Productivity of annual cropping and agroforestry systems on a shallow Alfisol in semi-arid India

An experiment was conducted at ICRISAT Center, Patancheru, India from June 1984 to April 1988 on a shallow Alfisol to determine whether the productivity of annual crop systems can be improved by adding perennial species such as Leucaena leucocephala managed as hedgerows. Except in the first year, crop yields were suppressed by Leucaena due to competition for moisture. The severity of competition was high in years of low rainfall and on long-duration crops such as castor and pigeonpea. Based on total biomass, sole Leucaena was most productive; even on the basis of land productivity requiring both Leucaena fodder and annual crops, alley cropping had little or no advantage over block planting of both components. Application of hedge prunings as green manure or mulch on top of 60 kg N and 30 kg P ₂ O ₅ha⁻¹ to annual crops did not show any benefit during the experimental period, characterized by below average rainfall. Indications are that (i) alley cropping was beneficial in terms of soil and water conservation with less runoff and soil loss with 3 m alleys than with 5.4 m alleys, and (ii) root pruning or deep ploughing might be effective in reducing moisture competition.     

Structure and Function of <Emphasis Type="Italic">Populus deltoides</Emphasis> Agroforestry Systems in Eastern India: 1. <Emphasis Type="Italic">Dry matter dynamics</Emphasis>

Agroforestry systems based on poplar (Populus deltoides) are becoming popular in eastern and northern parts of India. Therefore studies on the structure and function of the systems are important. The investigations included allometric equations for above- and belowground tree components, crop and plantation floor biomass and litter fall estimation at Pusa, Bihar, India. Biomass, floor litter mass, litter fall and net primary productivity (NPP) of plantations increased with an increase in age of trees whereas, crop biomass for any specific crop interplanted with poplar decreased with the age of the plantation. The total plantation biomass increased from 12.08 to 90.59 Mg ha⁻¹ and NPP varied from 5.69 to 27.9 Mg ha⁻¹ year⁻¹. The biomass accumulation ratio ranged from 2.1 to 3.2. Total annual litter fall was in between 1.95 and 10.00 Mg ha⁻¹ year⁻¹, of which 92–94% was contributed by leaf litter. Compartmental models were developed for dry matter distribution in agroforestry systems involving young (3-year-old) and mature (9-year-old) poplar trees interplanted with various crops, the crops being grown in two rotations maize (Zea mays) – wheat (Triticum aestivum) – turmeric (Curcuma domestica) and pigeonpea (Cajanus cajan) – turmeric. This study substantiates the potential of Populus deltoides G₃ under agroforestry combinations.     

A study of the potential of hedgerow intercropping in semi-arid India using a two-way systematic design

The potential of hedgerow intecrropping with Leucaena leucocephala was explored on vertic Inceptisols over 4 years at ICRISAT Center, Patancheru, India. The study was conducted using a systematic layout involving different alley widths ranging from 1.35 to 4.95 m and with varying distances between hedge and crops. The alleys were cropped with alternate rows of sorghum and pigeonpea. Hedges composed double Leucaena hedgerows 60 cm apart were periodically harvested for fodder. Sole crops of all components and a sorghum/pigeonpea intercrop were included in all four replications of the study.Starting in the second year, Leucaena was progressively more competitive to annual crops, causing substantial yield reduction. Competition (primarily for moisture) was most severe in narrow alleys and was greatest on pigeonpea.The growth of Leucaena was not sufficient to compensate for reduced crop yields. Land equivalent ratios (LERs) calculated on the basis of grain yield of crops and Leucaena fodder yields showed that hedgerow intercropping (HI) was advantageous over sole crops only during the first two years using wide alleys, but disadvantegeous in the last two years. LERs calculated on the basis of total dry matter indicated only a small advantage for HI (13–17 percent) over sole crops in wider (>4 m) alleys. Average returns per year from HI exceeded those of the most productive annual crop system (sorghum/pigeonpea intercropping) by 8 percent in 4.05 m alleys, and by 16 percent in 4.95 m alleys. Fodder production during the dry season was 40 percent of the annual total in these alley widths. Thus hedgerow intercropping at 4–5 m alley width is not very attractive for farmers in semi-arid India, which has 600–700 mm of annual rainfall. There is a need to examine the potential of HI in wider alleys. The merits and limitations of the systematic design are discussed.     

Nitrogen Dynamics in Cropping Systems in Southern Malawi Containing Gliricidia sepium, Pigeonpea and Maize

This study tested the hypothesis that incorporation of green leaf manure (GLM) from leguminous trees into agroforestry systems may provide a substitute for inorganic N fertilisers to enhance crop growth and yield. Temporal and spatial changes in soil nitrogen availability and use were monitored for various cropping systems in southern Malawi. These included Gliricidia sepium (Jacq.) Walp. trees intercropped with maize (Zea mays L.), with and without pigeonpea (Cajanus cajan L.), sole maize, sole pigeonpea, sole gliricidia and a maize + pigeonpea intercrop. Soil mineral N was determined before and during the 1997/1998, 1998/1999 and 1999/2000 cropping seasons. Total soil mineral N content (NO₃⁻ + NH₄⁺) was greatest in the agroforestry systems (p<0.01). Pre-season soil mineral N content in the 0–20 cm horizon was greater in treatments containing trees (≤85 kg N ha⁻¹) than in those without (<60 kg ha⁻¹; p<0.01); however, soil mineral N content declined rapidly during the cropping season. Uptake of N was substantially greater in the agroforestry systems (200–270 kg N ha⁻¹) than in the maize + pigeonpea and sole maize treatments (40–95 kg N ha⁻¹; p<001). Accumulation of N by maize was greater in the agroforestry systems than in sole maize and maize + pigeonpea (p<0.01); grain accounted for 55% of N uptake by maize in the agroforestry systems, compared to 41–47% in sole maize and maize + pigeonpea. The agroforestry systems enhanced soil fertility because mineralisation of the applied GLM increased pre-season soil mineral N content. However, this could not be fully utilised as soil N declined rapidly at a time when maize was too small to act as a major sink for N. Methods for reducing losses of mineral N released from GLM are therefore required to enhance N availability during the later stages of the season when crop requirements are greatest. Soil mineral N levels and maize yields were similar in the gliricidia + maize and gliricidia + maize + pigeonpea treatments, implying that addition of pigeonpea to the tree-based system provided no additional improvement in soil fertility.     

Growth and resource utilization of perennial pigeonpea (Cajanus cajan (L) Millsp.) at the tree-crop interface

Perennial pigeonpea (Cajanus cajan (L) Millsp.) was grown as a multipurpose tree species in strips or blocks with annual crops (sorghum, sunflower and chickpea) on a shallow Vertisol from 1987 to 1989. The interaction between the perennial pigeonpea and annual crops was measured at the tree-crop interface (TCI) by comparing the plants at the interface (I) and in the middle of the block planting (N).Perennial pigeonpea I plants had significantly more branches and bigger stems than N plants at the onset of the following rainy season. The greater number of flowers and grains of the I pigeonpea plants was partly due to a better lateral light level and partly due to a better access to water. On the other hand the negative effect on annual crops at the TCI extended to 1.5 m during the rainy season and to 2.5 m during the post-rainy season. Significant reduction in the growth of annual crops occurred at 30–40 days after sowing and was associated with the shading by the taller pigeonpea. Measurements of root profile of pigeonpea at the interface indicated that competition for moisture was the major cause for yield reduction of chickpea during the post-rainy season but an allelopathetic effect may also be involved. The results are compared with other TCI studies especially with Leucaena leucocephala in the semi-arid crops and the possible mechanisms for moisture interaction at the TCI are discussed.Submitted as ICRISAT Journal Article No. 1176 for Agroforestry Systems.     

Tree and crop productivity in gliricidia/maize/pigeonpea cropping systems in southern Malawi

This study examined the hypothesis that incorporation of Gliricidia sepium (Jacq.) Walp.) (gliricidia), a fast-growing, nitrogen-fixing tree, into agroforestry systems in southern Malawi may be used to increase the input of organic fertilizer and reduce the need for expensive inorganic fertilizers. The productivity of maize (Zea mays L.), pigeonpea (Cajanus cajan L.) and gliricidia grown as sole stands or in mixed cropping systems was examined at Makoka Research Station (latitude 15° 30′ S, longitude 35° 15′ E) and a nearby farm site at Nazombe between 1996 and 2000. Treatments included gliricidia intercropped with maize, with or without pigeonpea, and sole stands of gliricidia, maize and pigeonpea. Trees in the agroforestry systems were pruned before and during the cropping season to provide green leaf manure. Maize yields and biomass production by each component were determined and fractional light interception was measured during the reproductive stage of maize. Substantial quantities of green leaf manure (2.4 to 9.0 Mg ha⁻¹ year⁻¹) were produced from the second or third year after tree establishment. Green leaf manure and fuelwood production were greatest when gliricidia was grown as unpruned sole woodlots (c. 8.0 and 22 Mg ha⁻¹ year⁻¹ respectively). Improvements in maize yield in the tree-based systems also became significant in the third year, when c. 3.0 Mg ha⁻¹ of grain was obtained. Tree-based cropping systems were most productive and exhibited greater fractional light interception (c. 0.6 to 0.7) than cropping systems without trees (0.1 to 0.4). No beneficial influence of pigeonpea on maize performance was apparent either in the presence or absence of gliricidia at either site in most seasons. However, as unpruned gliricidia provided the greatest interception of incident solar radiation (>0.9), coppicing may be required to reduce shading when gliricidia is grown together with maize. As pigeonpea production was unaffected by the presence of gliricidia, agroforestry systems containing gliricidia might be used to replace traditional maize + pigeonpea systems in southern Malawi. This revised version was published online in June 2006 with corrections to the Cover Date.     

Management of pigeon pea in short fallows for crop-livestock production systems in the Guinea savanna zone of northern Ghana

Crop and livestock production in the Guinea savanna zone of northern Ghana has been declining over the past years as a result of increasing pressure on land. To sustain soil productivity, pigeon pea(Cajanus cajan), a leguminous perennial crop was evaluated for its potential as a short duration fallow crop for fodder and grain, and maize (Zea mays)production. It involved comparing a natural fallow (i.e., control) and four improved fallows of pigeon pea pruned annually at 30 cm, 60 cm and 90 cm from the ground, and unpruned pigeon pea over a two-year period. After this time, the land was cleared manually and planted to maize. The highest mean annual biomass of pigeon pea over the two-year period of 6.1 t ha⁻¹ dry matter (DM) was obtained by pruning at 60 cm. The highest leaf litter production and pigeon pea seed yield was obtained from the no pruning treatment. The mean maize grain yield from the improved fallow (3.02 t ha⁻¹) in the first year after clearing was significantly (P < 0.05) greater than that of the natural fallow (1.54 t ha⁻¹). Considering the biomass of pigeon pea from pruning, pigeon pea seed yield and maize grain yield after the pigeon pea, pruning pigeon pea at 60 cm is the most promising regime for crop-livestock production systems. This revised version was published online in June 2006 with corrections to the Cover Date.     

Can agricultural management emulate natural ecosystems in recharge control in south eastern Australia?

Water balance information at locations across southern Australia is analysed to identify mechanistic causes for higher water tables under agricultural systems compared to natural vegetation. Contrasting patterns of water use indicate pronounced physiological activity in summer by natural ecosystems ensuring the persistence of the dominant perennial species. A strategy of tempered water use during periods of rainfall excess in winter/spring enables seasonal carryover of soil water to be withdrawn by deep roots for summer functioning. Characteristic patterns of water use by agricultural systems, comprised mainly of determinant annual species, are truncated in time but feature elevated peak rates coinciding with periods of maximum soil water availability during the cooler months. Low seasonal vapour pressure deficit explains the observed benefits of enhanced biomass production due to high water use efficiency. The associated limited scope for soil water uptake carries the penalty of increased frequency of drainage and the subsequent resource degradation currently associated with farmland. Inclusion of a perennial lucerne phase in rotation with crops confers environmental benefit through summer uptake from a rooting depth double that of crops. An effective buffer is thus provided to restrict drainage under an ensuing cropping phase in a manner that more closely emulates natural systems. Simulations of phase farming point to a halving in long term drainage. They suggest adoption over a range of arable land units could cause local retreat of water tables by 0.3 m y⁻¹ initially, leading to a new equilibrium level which would still remain elevated with respect to that under natural vegetation. This revised version was published online in June 2006 with corrections to the Cover Date.     

Seasonal dynamics of mineral N pools and N-mineralization in soils under homegarden trees in South Andaman,India

Agroforestry trees are now well known to play a central role in the build up of nutrients pools and their transformations similar to that of forest ecosystem, however, information on the potential of homegarden trees accumulating and releasing nitrogen (mineralization) is lacking. The present study reports seasonal variations in pool sizes of mineral N (NH₄⁺-N and NO₃⁻-N), and net N-mineralization rate in relation to rainfall and temperature under coconut (Cocos nucifera L.), clove (Eugenia caryophyllata Thunb) and nutmeg (Myristica fragrans Houtt. Nees) trees in a coconut-spice trees plantation for two annual cycles in the equatorial humid climate of South Andaman Island of India. Concentration of NH₄⁺-N was the highest during wet season (May–October) and the lowest during post-wet season (November–January) under all the tree species. On the contrary, concentration of NO₃⁻-N was the lowest in the wet season and the highest during the post-wet season. However, concentrations of the mineral N were the highest under the nutmeg and the lowest under the coconut trees. Like the pool sizes, mean annual mineralization was the highest under the nutmeg (561 mg kg⁻¹ yr⁻¹) and the lowest under the coconut trees (393 mg kg⁻¹ yr⁻¹). Rate of mineralization was the highest during the post-wet season and the lowest during the dry season (February–April) under all the tree species. High rainfall during the wet season, however, reduced the rate of nitrification under all the tree species. The mean annual mineralization was logarithmically related with rainfall amount and mean monthly temperature.     

Intercropping field crops between rows of Leucaena leucocephala under rainfed conditions in northern India

A field study was conducted for six years (1981–1986) on sandy loam soil on intercropping hedgerows of Leucaena leucocephala (Lam) de Wit. with three field crops viz. maize (Zea mays L), black gram (Vigna mungo L) and cluster bean (Cyamopsis tetragonoloba L Taub.). In treatments 1 and 2 Leucaena hedges were planted as pure crops at close (25 cm × 75 cm) and wide (25 cm × 375 cm) spacings. In treatments 3, 4 and 5 the three field crops were intercropped between the hedgerows of Leucaena at the wide spacing, and in treatments 6, 7 and 8 the field crops were raised as pure crops. Leucaena was topped to 75 cm each time it attained a height of 175 cm. The pure crop of Leucaena at close spacing produced an average, over the six years, of 34 t ha⁻¹a⁻¹ of green fodder and 9.4 t ha⁻¹a⁻¹ of air dry fuelwood. The Leucaena at wide spacing produced 18.9 t ha⁻¹a⁻¹ of green fodder and 6.3 t ha⁻¹a⁻¹ of fuelwood. Intercropping with field crops decreased the yield of green fodder and fuelwood. The yield of all the field crops was less when raised as intercrops than as pure crops. Mean maximum net returns were obtained from intercrops of Leucaena and cluster bean (Rs 3540 ha⁻¹a⁻¹) which were significantly higher than the returns from pure crop of Leucaena at wide spacing but similar to the returns from pure crops of cluster bean. Leucaena with maize (Rs 3273 ha⁻¹a⁻¹) and black gram (Rs 3125 ha⁻¹a⁻¹) gave significantly higher net returns over pure crops of Leucaena at wide spacing, maize and black gram. ICRISAT = International Crops Research Institute for the Semi-Arid Tropics- Hyderabad, India. CIAT = Centro International de Agricultura Tropical - Cali - Columbia     

Seasonal variations in species diversity, dry matter and net primary productivity of herb layer of Quercus leucotrichophora-Pinus roxburghii mixed forest in Kumaun Himalaya, India

Plant biomass, species diversity and net primary productivity are presented for herb layer of banj oak (Quercus leucotrichophora A. Camus)-chir pine (Pinus roxburghii Sarg.) mixed forest in Kumaun, central Himalaya, India. The species diversity declined from a maximum (3.56) in September to a minimum (2.11) in December. The monthly live shoots biomass exhibited a single peak growth pattern with highest live shoot biomass of 185 g·m-2 in August. The seasonal pattern showed that the maximum above-ground production (131 g·m-2) occurred during the rainy season and the minimum (1 g·m-2) during winter season. The below-ground production was maximum during winter season (84 g·m-2) and minimum during summer season (34 g·m-2). The annual net shoot production was 171 g·m-2 and total below-ground production was 165 g·m-2. Of the total input 61% was channeled to above-ground parts and 39% to below-ground parts. Transfer of live shoots to dead shoots compartments and that of dead shoots to litter compartments was 61% and 66%, respectively. The total dry matter disappearance was 61% of the total input within annual cycle. The herb layer showed a net accumulation of organic matter, indicating the seral nature of the community.     

Structure and function of traditional agroforestry systems in the western Himalaya. I. Biomass and productivity

Species composition, biomass, and productivity patterns of three types of traditional agroforestry systems, namely, agrisilvicultural, agrihorticultural, and agrihortisilvicultural, commonly practiced in the western Himalaya, were studied. Species composition in the systems varied depending upon the size of the land holdings, and the basic requirements of the farmer. Among three systems agrihortisilvicultural was highly diverse in vegetation, with as many as 13 tree and 5 agricultural crops mixed together. This system showed the highest productivity up to 25.8 t ha⁻¹ yr⁻¹, out of which 68 percent was contributed by the trees and the remainder by the annuals. Agrisilvicultural system having predominantly annuals had the lowest productivity of 20.4 t ha⁻¹ yr⁻¹ with only 27 percent contribution by the trees. Total aboveground biomass in agrihortisilvicultural or agrihorticultural system was around 48 t ha⁻¹ and it was about 2-fold higher than agrisilvicultural system. In fodder trees, a significant percentage of annual production, up to 48 percent, was allocated in current twigs, while in horticultural trees a major portion, up to 63 percent was partioned towards fruits. Implications of the results have been discussed in context with the growing problems of fuel, fodder, and packaging material in the western Himalaya.     

Productivity and nutrient cycling in young Acacia senegal farming systems on Vertisol in the Blue Nile region, Sudan

Acacia senegal, the gum arabic-producing tree, is the most important component of traditional dryland agroforestry systems in the␣Sudan. The spatial arrangement of trees and the type of agricultural crop used influence the interaction between trees and crops. Tree and crop growth, gum and crop yields and nutrient cycling were investigated over a period of 4 years. Trees were grown at 5 × 5 m and 10 × 10 m spacing alone or in mixtures with sorghum or sesame. No statistically significant differences in sorghum or sesame yields between the intercropping and control treatments were observed (mean values were 1.54 and 1.54 t ha⁻¹ for sorghum grain and 0.36 and 0.42 t ha⁻¹ for sesame seed in the mixed and mono-crop plots, respectively). At an early stage of agroforestry system management, A. senegal had no detrimental effect on crop yield; however, the pattern of resource capture by trees and crops may change as the system matures. A significant positive relationship existed between the second gum picking and the total gum yield. The second gum picking seems to be a decisive factor in gum production and could be used as an indicator for the prediction of the total gum yield. Soil organic carbon, N, P and K contents were not increased by agroforestry as compared to the initial levels. Soil OC was not increased by agroforestry as compared to sole cropping. There was no evidence that P increased in the topsoil as the agroforestry plantations aged. At a stocking density of 400 trees ha⁻¹ (5 × 5 m spacing), A. senegal accumulated in its biomass a total of 18.0, 1.21, 7.8 and 972 kg ha⁻¹ of N, P, K and OC, respectively. Agroforestry contributed ca. 217 and 1500 kg ha⁻¹ of K and OC, respectively, to the top 25-cm of soil during the first four years of intercropping.     

Growth and yield of maize and timber trees in smallholder agroforestry systems in Claveria,northern Mindanao,Philippines

On-farm experiments were conducted in the Philippines to study over a 4-year period the growth of two timber trees, gmelina (Gmelina arborea R. Br.) and bagras (Eucalyptus deglupta Blume), and their impact on the grain yield of intercropped maize. The experiment consisted of maize monocropping plots (control) and maize intercropped between trees planted in block (2 × 2.5 m), and hedgerow arrangement (1 × 10 m). Three maize crops were planted in the block plots before canopy closure, and seven maize crops were planted in the hedgerow and monocropping plots. Maize grain yield in the hedgerow and in the block arrangement with gmelina were respectively 37% (16.58 tons ha⁻¹) and 68% (8.3 tons ha⁻¹) lower than in monocropping (26.21 tons ha⁻¹). In the plots with bagras, maize grain yield in hedgerow and in block arrangement were respectively 19% (24.8 tons ha⁻¹) and 66% (10.4 tons ha⁻¹) lower than in monocropping (30.6 tons ha⁻¹). For both tree species, the diameter at breast height (dbh) was greater in hedgerow than in block arrangement, with the difference being more pronounced with age. It was estimated that gmelina planted in hedgerows would produce 6–8 m³ ha⁻¹ of merchantable volume more than if planted in block. The study verifies the hypothesis that intercropping between widely-spaced trees rows (planted at 10 m or more) is more profitable and feasible to smallholders than either maize monocropping or woodlots, and concludes with recommendations on how to further improve the productivity of tree-intercropping systems.     

Growth characteristics of multipurpose tree species,crop productivity and soil properties in agroforestry systems under subtropical humid climate in India

Multipurpose tree species （MPTs） were studied in an agroforestry arboretum under subtropical humid climate in Northeast India. Out of 12 MPTs planted under agroforestry systems, Acacia auriculiformis in spacing of 2 m × 2 m （2500 stems·hm^-2） could have the potentiality to meet the timber/fuelwood requirement due to its high wood production of 635 m^3·hm^-2 with mean annual increment （MAI） of 2.54×10^-2 m^3.treel.a^-1 in a short rotation period of 10 years. Thus, A. auriculiformis is a short rotation forest tree species suitable to grow in subtropical humid climate. On the other hand, at 16 years of age, Eucalyptus hybrid and Michelia champaca in spacing of 3 m × 3 m （1111 stems.hm^2） produced appreciably high timber volume of 315 m^3.hm^-2 and 165 m^3.hm^-2 with MAI of 1.77×10^-2 m^3.tree^-1·a^-1 and 0.92×10.2 m^3.tree^-1.a^-1, respectively. At 16 years of age, Gmelina arborea produced a timber volume of 147 m^3.hm^-2 with MAI of 1.47×10^-2 m^3.tree^-1.a^-1 followed by Samania saman （140 m^3.hm^-2）, Albizziaprocera （113 m^3·hm^-2） and Tectona grandis （79 m3.hm^-2） with MAI of 1.40, 1.13 and 0.78 × 10^-2 m^3 .tree^-1a^-1, respectively in 4 m × 4 m spacing （625 stems.hm^-2）. Gliricidia maculata and Leucaena leucocephala could be used as live fences around the farm boundary to supply their N-rich leaves for mulch as well as manure to crops. In agroforestry arboretum, direct seeded upland rice （Oryza sativa - variety, AR-11）, groundnut （Arachis hypogaea - variety, JL-24） and sesamum （Sesamum indicum - variety, B-67） were grown during the initial period upto 8 years of tree establishment. Under other MPTs, there was a reduction in crop productivity as compared to open space. After 8 years of tree establishment, horti-silvi and silvi-pastoral systems were developed and pineapple （Ananas comosus - variety Queen）, turmeric （Curcuma longa -variety RCT -1） and cowpea （Vigna sinensis - variety Pusa Barsati） as forage crop were raised. The productivity of p     

Alley cropping in the moist savanna of West-Africa: II. Impact on soil productivity in a North-to-South transect in Togo

The moist savanna of West-Africa is characterized by a wide range of climates and soil types. The impact of the biophysical environment on hedgerow N uptake, wood production and maize grain yield was assessed for three years in three alley cropping trials with a selected number of hedgerow species in Glidji (Southern Togo), Amoutchou (Central Togo), and Sarakawa (Northern Togo). Senna siamea hedgerows accumulated significantly more N in the first pruning in Glidji (129−138 kg N ha−1) and Sarakawa (102−185 kg N ha−1) than in Amoutchou (17–26 kg N ha−1). This difference in N uptake was attributed to the infertile subsoil in Amoutchou, which was sandy up to 1 m and had a shallow groundwater-table. The amount of N accumulated in the Gliricidia sepium biomass varied between 38 kg N ha−1 in Glidji and 142 kg N ha−1 in Amoutchou. Averaged over all species and sites, 9 to 29% and 9 to 39% of the annual N accumulation in the hedgerow biomass is incorporated in the second, respectively third pruning. The Gliricidia trees produced between 12 and 26 ton fresh matter ha−1 of wood and the Senna trees between 4 and 38 ton fresh matter ha−1. Maize grain yield in Glidji was not affected by treatments (3196 kg ha−1, on average). In Amoutchou, the highest grain production was observed in the Gliricidia treatment (2774 kg ha−1 vs 1007 kg ha−1 in the control), while in Sarakawa, the Gliricidia (3786 kg ha−1) and Senna (3842 kg ha−1) plots produced a greater grain yield than the control plots (2123 kg ha−1). Maize yield increase in the alley cropping systems relative to the control plots was related to the soil total N content. Top and sub-soil characteristics were shown to be an important modifier of the functioning of alley cropping systems and should be taken into account when deciding on whether to use alley cropping and when selecting the hedgerow species. This revised version was published online in June 2006 with corrections to the Cover Date. This revised version was published online in June 2006 with corrections to the Cover Date.     

Soil water dynamics in cropping systems containing <Emphasis Type="Italic">Gliricidia sepium</Emphasis>, pigeonpea and maize in southern Malawi

The water dynamics of cropping systems containing mixtures of Gliricidia sepium (Jacq.) Walp trees with maize (Zea mays L.) and/or pigeonpea (Cajanus cajan L.) were examined during three consecutive cropping seasons. The trees were pruned before and during each cropping season, but were left unpruned after harvesting the maize; prunings were returned to the cropping area in all agroforestry systems to provide green leaf manure. The hypothesis was that regular severe pruning of the trees would minimise competition with crops for soil moisture and enhance their growth by providing additional nutrients. Neutron probe measurements were used to determine spatial and temporal changes in soil moisture content during the 1997/98, 1998/99 and 1999/00 cropping seasons for various cropping systems. These included gliricidia intercropped with maize, with and without pigeonpea, a maize + pigeonpea intercrop, sole maize, sole pigeonpea and sole gliricidia. Soil water content was measured to a depth of 150 cm in all treatments at 4–6 week intervals during the main cropping season and less frequently at other times. Competition for water was apparently not a critical factor in determining crop performance as rainfall exceeded potential evaporation during the cropping season in all years. The distribution of water in the soil profile was generally comparable in all cropping systems, implying there was no spatial complementarity in water abstraction by tree and crop roots. However, available soil water content at the beginning of the cropping season was generally lower in the tree-based systems, suggesting that the trees continued to deplete available soil water during the dry season. The results show that, under rainfall conditions typical of southern Malawi, the soil profile contains sufficient stored water during the dry season (ca. 75–125 mm) to support the growth of gliricidia and pigeonpea, and that gliricidia trees pruned before and during the cropping season did not deleteriously compete for water with associated crops. Water use efficiency also appeared to be higher in the tree-based systems than in the sole maize and maize + pigeonpea treatments, subject to the proviso that the calculations were based on changes in soil water content rather than absolute measurements of water uptake by the trees and crops.     

Growth and ecological impacts of traditional agroforestry tree species in Central Himalaya, India

A number of multipurpose tree species are conserved as scattered trees in settled farms on terraced slopes by the traditional farmers in Central Himalaya, India. Knowledge on growth rates and ecological impacts of these tree species is limited. Ten locally valued multipurpose tree species, viz., Albizzia lebbek, Alnus nepalensis, Boehmeria rugulosa, Celtis australis, Dalbergia sissoo, Ficus glomerata, Grewia optiva, Prunus cerasoides, Pyrus pashia and Sapium sebiferum, were established as mixed plantations at a degraded community forest land site and an abandoned agricultural land site in a village at 1200 m altitude in District Chamoli, India. At the abandoned agricultural land site, annual food crops were grown, along with planted trees, providing supplemental irrigation and organic manure following traditional farming practices. Survival, height, stem circumference, crown depth and width, number of branches, above-ground biomass and soil physico-chemical characteristics were monitored up to five years of plantation growth. Above-ground tree biomass accumulation at the abandoned agricultural land site was 3.9 t ha⁻¹ yr⁻¹ compared with 1.1 t ha⁻¹ yr⁻¹at the degraded forest land site. B. rugulosa, C. australis, F. glomerata, G. optiva, P. cerasoides and S. sebiferum showed more prominent differences in growth at the two sites compared with A. lebbek, A. nepalensis, D. sissoo and P. pashia. A. nepalensis and D. sissoo showed best growth performance at both the sites. A significant improvement in soil physico-chemical characteristics was observed after five years at both of the sites. Carbon sequestration in soil was higher than that in bole biomass. This revised version was published online in June 2006 with corrections to the Cover Date.     

Biomass production, foliar and root characteristics and nutrient accumulation in young silver birch (Betula pendula Roth.) stand growing on abandoned agricultural land

The above-ground biomass and production, below-ground biomass, nutrient (NPK) accumulation, fine roots and foliar characteristics of a 8-year-old silver birch (Betula pendula) natural stand, growing on abandoned agricultural land in Estonia, were investigated. Total above-ground biomass and current annual production after eight growing seasons was 31.2 and 11.9 t DM ha⁻¹, respectively. The production of stems accounted for 62.4% and below-ground biomass accounted for 19.2% of the total biomass of the stand. Carbon sequestration in tree biomass reaches roughly 17.5 t C ha⁻¹ during the first 8 years. The biomass of the fine roots (d < 2 mm) was 1.7 ± 0.2 t DM ha⁻¹ and 76.2% of it was located in the 20 cm topsoil layer. The leaf area index (LAI) of the birch stand was estimated as 3.7 m² m⁻² and specific leaf area (SLA) 15.0 ± 0.1 m² kg⁻¹. The impact of the crown layer on SLA was significant as the leaves are markedly thicker in the upper part of the crown compared with the lower part. The short-root specific area (SRA) in the 30 cm topsoil was 182.9 ± 9.5 m² kg⁻¹, specific root length (SRL), root tissue density (RTD) and the number of short-root tips (>95% ectomycorrhizal) per dry mass unit of short roots were 145.3 ± 8.6 m g⁻¹, 58.6 ± 3.0 kg m⁻³ and 103.7 ± 5.5 tips mg⁻¹, respectively. In August the amount of nitrogen, phosphorus and potassium, accumulated in above ground biomass, was 192.6, 25.0 and 56.6 kg ha⁻¹, respectively. The annual flux of N and P retranslocation from the leaves to the other tree parts was 57.2 and 3.7 kg ha⁻¹ yr⁻¹ (55 and 27%), respectively, of which 29.1 kg ha⁻¹ N and 2.8 kg ha⁻¹ P were accumulated in the above-ground part of the stand.     

Alley cropping in the moist savanna of West-Africa: I. Restoration and maintenance of soil fertility on ’terre de barre‘ soils in Bénin Republic

The potential of alley cropping systems supplied with a limited amount of fertilizer to restore crop productivity on a degraded site and to maintain crop productivity on a recently cleared, non-degraded site on ‘terre de barre‘ soils in Southern Bénin was investigated from 1994 to 1996. Leucaena leucocephala, Senna siamea and Gliricidia sepium were used as hedgerow species. Maize yields of the no-tree control plots dropped from the initial (1990) 401 kg ha−1 and 2181 kg ha−1 on the degraded and non-degraded sites, respectively, to 109 kg ha−1 and 1346 kg ha−1 in 1996, even with application of a minimal amount of mineral fertilizer. The alley cropping systems produced on average (mean of three treatments and three years) 107% more grain than the initial 1990 values on the degraded site and 11% less grain than the initial 1990 values on the non-degraded site. Especially the Senna and to a lesser degree the Leucaena treatment yielded consistently more grain than the control. The Senna trees contained a larger amount of N and produced more wood during the first pruning on the degraded site (155 kg N ha−1 and 14.0 ton fresh wood ha−1) than on the non-degraded site (49 kg N ha−1 and 6.6 ton fresh wood ha−1) most likely because of differences in subsoil fertility, as indicated by the higher clay, exchangeable bases, and N content between 60 and 125 cm cm. N accumulation and wood production by the Leucaena and Gliricidia trees was similar in both sites (82 and 36 kg N ha−1 and 4.6 and 9.3 ton fresh wood ha−1, respectively). When a limited amount of fertilizer is available, Senna appears to be the best choice as hedgerow species on sites with a relatively fertile subsoil. For other soils, a N2-fixing species may be a better choice. This revised version was published online in June 2006 with corrections to the Cover Date. This revised version was published online in June 2006 with corrections to the Cover Date.