云南松林分系统的耗散结构特性研究

云南松林分系统分布于北纬23°~29°、东经98°30′~106°、海拔高度为1 200~2 800 m的区域,面积约500万hm2,这一地理区域是由多种树种组成林分系统的生态区域。在诠释此云南松林分系统的生态环境及生长过程的前提下,引入了“耗散结构的进化观念”对本区域云南松林分系统的结构状态和演变动态进行了研究,导算出在8种地位级上发生的131条由“初始林分”走向“过熟林分”的“坐标序列”,表明其结构具有“对称分布”的特征,揭示了云南松林分系统与其生态阈限环境相互平衡的特性。并论述了云南松林分系统在由初始林分走向过熟林分的过程中都要与外界进行能量和物质转化,而由其产生的净生长量和死亡量是一种促成林分系统进化的理念,以此表明可用内部时间T(D-):3、4、5……32所刻画的131条“株数递减率/蓄积量递增率”过程,来反映云南松林分系统的演变动态的客观实际。     

论思茅松林分系统的耗散结构特性

思茅松林分系统在地球上形成确定的地理分布区域, 其包含幼林、中林、成熟林、过熟林等许多林分,在各地位级上形成“正态分布”, 表达了林分系统与外界环境的和谐状态。外界环境是非均匀分布的“能量”和“物质”、即多样性的“立地环境”。因此思茅松的不同地位级、不同密度、不同年龄的许多林分, 与环境进行能量、物质交换的规律都表现为由幼林走向过熟林的多种“坐标序列”, 每一条“坐标序列”产生的净生长量, 都大于耗散掉的死亡量。不同“立地环境”、不同初始密度的林分, 由初始郁闭走向过熟林分的演变过程,都可以用y=A D±B的数学模型来描述, 它揭示出林分系统的“周期行为”; 进入过熟林分以后, 林分结构变为多个“世代”组成的“异龄林分结构”, 而各个“世代”发生的“株数递减/蓄积量递增”过程都不同。“世代更迭”的行为不遵守“时间倍增”规律, 而是更为复杂的自组织行为, 因此称为“非周期行为”。“周期行为”趋向复杂的“非周期行为”是一条演化通路, 表达出思茅松林分系统在时、空中的“物质流”。这种“物质流”必定是与外界环境的“能量流”进行能量、物质交换所产生出来的。     

云南松林分系统的存在与演化

云南松林分系统在时、空中的随机分布，遵循“概率对称性”的描述。而林分产生的株数递减率和蓄积量递增率则遵循“非对称性转移概率”的描述。这就是云南松林分系统存在与演化的两种概率现象。由“对称性概率”描述的林分系统结构状态中产生“非对称性转移概率”，所造成的各种林分结构仍然遵循“对称性概率”的描述，由此可见林分系统在时、空中的演变，可以归结为“对称性概率”与“非对称性转移概率”的交替变换，因此能用内部时间刻划的“立木度／蓄积度”序列及其对应的“株数递减率／蓄积量递增率”序列，描述各种林分结构的连续演变过程。     

云南松林分系统的形成与演变

优良木成长壮大不断分化,被压木衰退消亡是云南松林分系统演变的两个方面。由优良木组成的上层林冠和被压木组成的下层林冠,是云南松林分系统中的两个子系统。在饱和林分系统内,占据上层林冠的优良木子系统每年分化为被压木的数量,与处于下层林冠的被压木子系统的死亡量相协调,以保持饱和林分系统结构,保持优良木株数占30—40％,蓄积量占60—70％。在饱和林分系统内间伐Ⅲ、Ⅳ、Ⅴ级木,则使优良木的生产量上升,同时使被压木的死亡量下降,直至林分回归于饱和林分系统结构。优良木子系统随全林分系统的演变,因林分系统密度而异,是生境、年龄、密度相互制约所表现的数量集合,都按照幂函数曲线轨迹演变,其曲线系数“A”随密度下降而上升,“B”值则随密度下降而减小。依照云南松林分系统的演变规律,对预测优良木保留量和确定间伐强度有实用价值。     

云南松林分系统的存在与演化

云南松林分系统在时,空中的随机分布,遵循“概率对称性”的描述。而林分产生的株数递减率和蓄积量递增率则遵循“非对称性转移概率”的描述。这就是云南松林分系统存在与演化的两种概率现象。由“对称性概率”描述的林分系统结构状态中产生“非对和性转移概率”,所造成的各种林分结构仍然遵循“对称性概率”的描述,由此可见林分系统在时,空中的演变,可以归结为“对称性概率”和“非对称性转移概率”的交替交变,因此能用内部时     

思茅松林分演变过程周期性的研究

优良木（Ⅰ、Ⅱ、Ⅲ级木）的成长和分化,被压木（Ⅳ、Ⅴ级木）的衰退和死亡,是思茅松林分演变过程中相互联系的两方面。林分平均直径的变化与“内部时间（Ｔ＝Ｄ）”有关系。由“内部时间（Ｔ＝Ｄ）”刻划的“立木度／蓄积度”序列,与由“内部时间（Ｔ＝Ｄ）”刻划的“株数递减率／蓄积量递增率”序列的相互关联关系,反映出林分结构状态的演变。在８、１６、３２这几个临界时刻,“立木度／蓄积度”的数值都趋近１０,这是思茅松林分演变的３个周期,对应着周期性的转移概率序列。用在判断林分生长、预测林分演变过程、确定林分间伐强度等方面都有使用价值。     

云南松林分与小蠹虫的竞争状态研究

云南松林遭小蠹虫危害以后，常由于林分抵御功能的不同而形成区域性的蠹害林分系统，即在“虫群区域”内存在着“衰退”和“发展”两种林分演变过程，若按“演变属性”来分析，则看到蓄积量下降、蓄积量停滞、蓄积量上升等３种林分演变过程是蠹害林分系统的普遍性特征，前２种属于“衰退状态”，后１种属于“发展状态”。对３个蠹害林分系统的“演变属性”用二项分布Ｐ（衰退概率）＋Ｑ（发展概率）＝１来描述，那么二项式（Ｐ＋Ｑ）＾３方程中的系数１、３、３、１成为系统中不同“演变属性”的概率论判断。由人为因素、虫群因素、环境因素引起的林分“衰退”和“发展”当然是随机的，但从分析中看到人的正、反两面行为都会引起林分与虫群相互作用的变化。     

论林分系统的存在与演变

天然森林中包含着多种类型的林分类型,各个林分系统具有自然历史确定的地理分布范围,外界的“能量流”和“物质流”都呈非均匀分布状态,因此造成林分系统“结构状态”和“演变动态”的多样性。林分系统中的各个林分与外界进行能量、物质转换的规律相似,林分内部的“优胜劣汰”规律也相似。因此可用“内部时间T(D)”：3、4、5……32所刻画的“立木度／蓄积度”序列和“株数递减率／蓄积量递增率”序列的数学模型,来描述林分从幼林到过熟林的演变过程,这就是林分系统的“周期行为”。进入过熟林分以后,林分平均直径“D”的变化无规律性,但过熟林分仍然通过“新生林木更替衰老林木”而向前演变发展,这种“非周期行为”表达了林分系统“从存在到演化”的永恒性。     

云南松林分与小蠹虫的竞争状态研究

云南松林遭小蠹虫危害以后,常由于林分抵御功能的不同而形成区域性的蠢害林分系统,即在"虫群区域"内存在着"衰退"和"发展"两种林分演变过程,若按"演变属性"来分析,则看到蓄积量下降、蓄积量停滞、蓄积量上升等3种林分演变过程是蠹害林分系统的普遍性特征,前2种属于"衰退状态",后1种属于"发展状态"。对3个蠢害林分系统的"演变属性"用二项分布P(衰退概率)+Q(发展概率)=1来描述,那么二项式(P+Q)3方程中的系数1、3、3、1成为系统中不同"演变属性"的概率论判断。由人为因素、虫群因素、环境因素引起的林分"衰退"和"发展"当然是随机的,但从分析中看到人的正、反两面行为都会引起林分与虫群相互作用的变化。     

思茅松林分演变过程周期性的研究

优良木（Ⅰ、Ⅱ、Ⅲ级木）的成长和分化,被压木（Ⅳ、Ⅴ级木）的衰退和死亡,是思茅 松林分演变过程中相互联系的两方面。林分平均直径的变化与“内部时间（Ｔ＝Ｄ）”有关系。     

Browsing Damage on Pine (Pinus sylvestris and P. contorta) by a migrating moose (Alces alces) Population in Winter: Relation to Habitat Composition and Road Barriers

In Scandinavia, moose (Alces alces L.) sometimes cause severe browsing damage to economically-important pine. Moose-vehicle accidents have spurred construction of fences along roads, and these may interfere with moose migration between summer and winter ranges, or the road alone may be a barrier. If this happens and moose build up along roads, landowners may suffer economically. Therefore, this study investigated whether roads, fences or other factors influence the use of young pine stands by moose. Eighty stands along roads in northern Sweden were evaluated in which individually-browsed branches were counted on 9972 pines. Moose browsing was not significantly related to birch (Betula pendula Roth, B. pubescens Ehrh.) density, nor did it differ between pines (Pinus contorta Douglas or P. sylvestris L.). However, increased pine density, site productivity and proximity to a highway were associated with increased browsing. Further large-scale studies are needed to understand moose habitat selection and the effects of roads.     

Effects of Root Freezing on the Physiology and Growth of Picea glauca,Picea mariana and Pinus banksiana Seedlings Under Different Soil Moisture Regimes

The root systems of 2-yr-old Picea glauca, Picea mariana and Pinus banksiana seedlings were submitted to various frost temperatures during an artificial frost to induce different levels of root damage. Frost-damaged and control seedlings were placed in a greenhouse under high and low soil moisture regimes. Seedling growth and physiology were evaluated periodically. Seedling survival was reduced when root damage reached levels of 60-80%. Root systems of all three species showed partial to total recovery by the end of the experiment. In general, root freezing damage caused reductions in seedling growth, with these reductions becoming less significant over time. Root damage had little to no effect on black spruce and jack pine seedling physiology, while white spruce CO 2 uptake decreased with increasing root damage. Shoot nitrogen content of all three species decreased slightly with increasing root damage.     

Stand Structure in Pine,Spruce and Deciduous Forests: A Comparison Between Finnish and Russian Karelia

This study investigated the stand structure in pine, spruce and deciduous forests in the border district of Finland and Russia. A total of 46 mature forest stands was selected as pairs, the members of each pair being as similar as possible with respect to their forest site type, age, moisture and topography. The stands were then compared between the two countries by means of basal areas and number of stems. The proportions of dominating tree species were 2-12% lower, and correspondingly the proportions of secondary tree species higher, in Russian forests. The density of the forest stock was also higher in each forest type in Russia. The forests in the two countries differed most radically in terms of the abundance of dead trees. The amount was two to four times higher in Russian deciduous and spruce forests, and in pine forests the difference was 10-fold. The stand structures indicated that Russian coniferous stands, in particular, were more heterogeneous than intensively managed pine and spruce stands in Finland.     

Growth and Terminal Bud Formation in Picea abies Seedlings Grown with Alternating Diurnal Temperature and Different Light Qualities

The root collar diameter and the height:diameter ratio are of particular importance in container-grown seedlings where a high density in the containers may produce spindly seedlings. Temperature regimes and light quality are known to affect plant growth. The aim of this study was to identify responses in Picea abies (L.) Karst. seedlings grown with light providing different red:far-red ratios and under temperature regimes with alternating day (DT) and night temperature (NT) from negative (DT < NT) to positive (DT > NT) difference (DIF) between DT and NT. Experiments were conducted in controlled environment chambers and in a daylight phytotron. Only limited thermoperiodic responses appeared in P . abies seedlings with respect to seedling height and dry weight accumulation. The formation of terminal buds, however, was clearly delayed in seedlings grown at negative DIF. The results indicate a requirement for day extension light that is high in far-red, to prevent terminal bud formation under natural short-day conditions. An extended study should be conducted to clarify the minimum level of light intensity and the optimal light quality needed to prevent terminal bud formation under natural short-day conditions.     

CAD与建筑设计的关系

本文分析了CAD在设计中引起的正面和负面影响,并进行了系统的阐述,从而使设计者在应用中能够保持客观的态度。     

Natural Regeneration in Clear-cuts: Effects of Scarification,Slash Removal and Clear-cut Age

The effects of soil scarification (mounding), slash removal and clear-cut age on the natural regeneration in clear-cuts was evaluated using data from four sites in southern Sweden. The treatments were carried out during a good seed and establishment year for birch ( Betula pubescens Ehrh. and B. pendula Roth). Scarification had the strongest positive effect on the density of naturally regenerated seedlings, especially in birch, but also in pine ( Pinus sylvestris L.) and spruce [ Picea abies (L.) Karst]. Slash removal had a positive effect on birch density. No statistically significant effect of clear-cut age was found. In addition, no statistically significant interactions between clear-cut age and scarification or slash removal were found. The ingrowth of field vegetation was the fastest in areas that were not scarified, less rapid in areas scarified on old clear-cuts, and the slowest after scarification in fresh clear-cuts. In conclusion, it may be possible to control the density of birch during a good establishment year for birch. If birch is desired, the best combination of treatments is to remove the slash and scarify; otherwise, these treatments should be avoided.     

Risk Perception Among Non-industrial Private Forest Owners

Based on an enquiry, risk perception among non-industrial private forest owners is described in relation to climate change and forestry hazards. Of the respondents, 11% took action to remedy the effects of climate change. Out of a given set, hazards were ranked according to each respondent's experience of recent substantial financial loss to the estate and in relation to his or her willingness to make investments aimed at risk reduction. For each hazard, the respondent assessed the risk in four classes ranging from very high to negligible risk. Six hazards were considered most problematic in all three aspects: browsing damage, falling timber prices, damage by wind, spruce bark beetle, root rot and pine weevil. A majority of the respondents claimed to take action to reduce the risk associated with at least one hazard, while 35% did not know whether they did. Excluding climate change, the need for decision support was the largest in relation to damage by wind owing to a combination of perceived high risk and a high level of ignorance in relation to whether risk-reducing measures were taken.     

Interactions in decomposition and N mineralization between tropical legume residue components

In situ produced plant residues contain a mixture of different plant components of varying quality. To assess synergistic or antagonistic interactions occurring during the decomposition and mineralization of such mixtures, individual plant parts (stems, leaves, leaf litter and roots) or the mixture of stems, leaves and leaf litter of the agroforestry species pigeonpea (Cajanus cajan) or of crop residues of peanut (Arachis hypogaea) or of the weed hairy indigo (Indigofera hirsuta) were incubated in pots for 19 weeks. Periodically, remaining plant residues were sieved out (>2 mm), weighed and N content as well as soil mineral N determined. Above- and below-ground residues of peanut decomposed fastest and showed the largest N release in agreement with their high N concentration and low-acid detergent fibre (ADF) : N ratio. Hairy indigo was hypothesized to be of lower quality than pigeonpea because of its high-polyphenol content. However, it decomposed faster than pigeonpea, largely because of the higher N and lower lignin concentration of its components. Ranking of individual plant components for N mineralization resulted in the following pattern, leaves > leaf litter > roots > stems. In mixtures of the different plant components a similar species order in decomposition was obtained, e.g. peanut > hairy indigo > pigeonpea. The amount of N released from the mixture was dominated by stem material that comprised 46–61% of the mixture. The interactions in mixtures were relatively small for peanut (generally high-quality components) as well as for pigeonpea (low proportion of high-quality components, i.e. N rich leaf material). However, a positive interaction occurred during later stages of N mineralization in the mixture of hairy indigo as it had a significant proportion of N rich components and absence of highly reactive polyphenols. Thus, for plants with low to intermediate chemical quality attributes, manipulating plant composition (e.g. by varying harvest age, affecting stem and leaf proportions) will be important to obtain significant interactions during decomposition when its components are mixed.     

Pasture production and tree growth in a young pine plantation fertilized with inorganic fertilizers and milk sewage in northwestern Spain

The use of organic waste materials such as milk sewage as an organic fertilizer could have the dual advantages of organic-waste disposal and reduced dependence on inorganic fertilizers. The effects of fertilization with (1) conventional mineral fertilization, (2) milk sewage sludge at 40 kg N ha⁻¹ target rate and (3) no fertilization on pasture production and tree growth were examined in an experiment consisting of two pasture mixtures under a one-year-old Pinus radiata plantation with a density of 2500 trees ha⁻¹. The two pasture mixtures were: (1) Dactylis glomerata L. var. saborto (25 kg ha⁻¹) + Trifolium repens L. group Ladino (4 kg ha⁻¹) + Trifolium pratense L. var. Marino (1 kg ha⁻¹); (2) Lolium perenne L. var. Tove (25 kg ha⁻¹) + Trifolium repens L. group Ladino (4 kg ha⁻¹) + Trifolium pratense L. var. Marino (1 kg ha⁻¹). The experiment began in the spring of 1995 using a randomized block design with three replicates in Castro Riberas de Lea (Lugo, Galicia, north-western Spain). Plot size was 12 × 8 m², with a 1 m buffer strip between plots. Two-year data showed that fertilization with either material had a positive effect on pasture production, with no significant difference between the two fertilization treatments. Tree growth in the milk sewage sludge plot was significantly higher than in the control plots. Inorganic fertilization increased pasture production, but affected tree growth negatively. The results show that milk sewage sludge could be used as a fertilizer in silvo-pastoral systems. This revised version was published online in June 2006 with corrections to the Cover Date.