Influence of defoliation regime on herbage production and characteristics of intake by dairy cows as affected by grazing intensity

Three contrasting defoliation regimes for dairy cows—four cuts annually, severe rotational grazing and lax rotational grazing integrated with cutting—were compared in terms of herbage production, selection and intake per cow. Lax and severe grazing were compared simultaneously and grazing intensity was characterized by means of changes in herbage mass and sward height during grazing.
Herbage growth and yield were similar under cutting, severe grazing and lax grazing/cutting (120 t organic matter (OM) ha⁻¹). Herbage quality and leaf production were highest with severe grazing, which also had a less marked seasonal pattern of growth. The herbage intake per cow was 111 kg OM d⁻¹ with severe grazing and proportionately (0-20) higher at lax grazing/ cutting. 050 of the herbage yield was harvested by cutting at lax grazing/cutting. Selectivity was described with high certainly by the nutrient content prior to defoliation and by the degree of defoliation. Models to quantify this are presented.
Grazing intensity could be described by the size and degree of utilization of the fouled area, and herbage intake was quantified by means of herbage height before and after grazing. Within the range of 80-240 mm sward height prior to grazing, height measurements led to realistic and more accurate estimates than those obtained by measuring organic matter disappearance.
Herbage utilization was depressed significantly by increasing maturity of herbage due to lower nutritive value, but in particular due to reduced green leaf content and increased reproductive development. If sward height did not exceed 250 mm at any time, good utilization by grazing could be obtained.     

Interaction between water-soluble carbohydrate reserves and defoliation severity on the regrowth of perennial ryegrass (Lolium perenne L.)-dominant swards

A field-study was undertaken in Hamilton, New Zealand to determine if there was an interaction between water-soluble carbohydrate (WSC) reserve content and defoliation severity on the regrowth of perennial ryegrass-dominant swards during winter. Perennial ryegrass plants with either low or high WSC content were obtained by varying the defoliation frequency. At the third defoliation at the one-leaf stage and at the first defoliation at the three-leaf stage (harvest H₁), swards were mown with a rotary lawnmower to residual stubble heights of 20, 40 or 60 mm. All swards were then allowed to regrow to the three-leaf stage before again defoliating to their treatment residual stubble heights (H₂). Frequently defoliated plants contained proportionately between 0·37 and 0·48 less WSC in the stubble after defoliation, depending on the severity of defoliation. There was no interaction between WSC content and defoliation severity for herbage regrowth between harvests H₁ and H₂. Herbage regrowth was lower from swards containing low WSC plants compared with high WSC plants (2279 vs. 2007 kg DM ha⁻¹). Furthermore, swards defoliated to 20 or 40 mm had greater herbage regrowth compared with those defoliated to 60 mm (2266, 2249 and 1914 kg DM ha⁻¹ for swards defoliated to residual stubble heights of 20, 40 and 60 mm, respectively). Regrowth of perennial ryegrass was positively correlated with post-defoliation stubble WSC content within defoliation severity treatment, implying that WSC contributed to the defoliation frequency-derived difference in herbage yield. However, the effect of defoliation severity on herbage regrowth was not associated with post-defoliation stubble WSC content.     

Tiller size/density compensation in perennial ryegrass miniature swards subject to differing defoliation heights and a proposed productivity index

The objective of this study was to investigate the effects of cutting height on tiller population density, tiller weight and herbage harvested in perennial ryegrass swards. One hundred mini-swards of Lolium perenne were established from seed on 5 May 1993 in 150 mm × 150 mm plastic pots in a greenhouse. The experimental period lasted 6 months (May to November). On 4 June all pots were cut to 50 mm and randomly allocated to five treatments (20, 40, 80, 120 and 160 mm sward surface height) with twenty replicates (four per tray). All pots were cut twice weekly to the specified surface height from 11 June, and recording began on 14 June. Every 4 weeks from 9 July to 5 November the twenty pots contained in one tray were withdrawn for destructive measurements of tiller population density and the weights of tiller components. Tiller population density increased with reduction in defoliation height, except for the 20-mm treatment where tiller density was initially restricted. The slope of the size/density compensation (SDC) line was close to ?5/2 over the range of defoliation heights 40–120 mm, but was less than ?1·0 between 120 and 160 mm. These slopes are consistent with a recent theory, which proposes that variations from a slope of ?3/2 will be linked to defoliation-induced changes in canopy leaf area and to change in tiller leaf area:volume ratio, R. At defoliation heights below 120 mm, SDC slope was increased by reduction in canopy leaf area. Above 120 mm, increase in R forced a relatively rapid tiller population decline, resulting in a reduced SDC slope and decrease in canopy leaf area. Whereas traditional measures of leafiness such as leaf:stem ratio or leaf:non-leaf ratio decreased with increasing height of defoliation, the leaf area:volume ratio, R, increased with defoliation height. Distance from an arbitrarily positioned ?3/2 SDC line was correlated with sward productivity.     

Temperate grasslands and global atmospheric change: a review

Recent reviews confirm and extend previous observations that elevated carbon dioxide (CO₂) concentrations stimulate photosynthesis, leading to increased plant productivity. Elevated CO₂ concentrations tend to reduce the sensitivity of grassland ecosystems to low levels of precipitation but induce progressive nitrogen (N) limitations on plant growth which can be alleviated by supplying a significant external input of N in the form of mineral fertilizer or through the increased use of N-fixing legumes. Other nutrients, such as phosphorus, can act as the main limiting factor restricting the growth response in legumes to atmospheric CO₂ concentration. The botanical composition of temperate grasslands is affected by the rise in atmospheric CO₂ concentration, possibly through a decline in the relative abundance of grasses. Elevated CO₂ concentration will also alter the feeding value of herbage to grazers both in terms of fine-scale (for example, crude protein concentration and C:N ratio) and coarse-scale (legumes vs. grasses and C₃- vs. C₄-species) changes. The management guidelines of grasslands will need to be adapted to global atmospheric and climatic changes and to increased variability in climate.     

The effect of height and frequency of mowing on the yield and composition of perennial ryegrass—white clover swards in the autumn to spring period

Ninety-six plots (3 × 2 m) of well-established perennial rye grass/white clover pasture were mown to heights of 2·7 (Low) or 3·96 (High) cm (rising plate meter) at 14-, 28-, 84- or 112-d intervals in autumn-winter. A 7-, 14- and 28-d mowing interval was superimposed in spring on each autumn–winter mowing interval treatment with the low and high mowing heights altered to 2·92 and 4·80 cm, respectively.
With the low cutting height, accumulated herbage DM was more than doubled (1806 ± 79 kg DM ha^-1) compared to a 'high' (754 ± 49 kg DM ha^-1) cutting height in autumn–winter and this was due to increased harvesting efficiency rather than growth as estimated by leaf extension. Although defoliation interval had no effect on DM yield, the grass component increased and clover decreased. The composition effect carried over into spring. On average, 3·5 tillers were produced over winter for each ryegrass tiller present in autumn and tiller densities were higher in spring. Tillers produced over autumn–winter contributed more than 60% of ryegrass growth by early spring.
In early spring (16–30 September), the low cutting height increased herbage DM yield, in mid-spring (1–14 October) it reduced DM yields particularly in combination with short defoliation intervals, while in late spring (14 October to 11 November) cutting height had no effect on DM yields.
Over the entire spring period there was a very marked effect of defoliation interval on DM yields.     

Defoliation and productivity of a phalaris-subterranean clover sward, and the influence of grazing experience on sheep intake

Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha⁻¹) and high (22 sheep ha⁻¹) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.
There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward < lightly grazed sward < heavily grazed sward < repeatedly cut sward.
When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.     

Frequency of defoliation of native and naturalized species of the Flooding Pampas (Argentina)

This study describes the effect of herbage mass of species on defoliation patterns in a multispecies temperate–subhumid natural grassland of the Flooding Pampas at low, medium and high sward herbage mass [761  ±  8·7, 989  ±  137 and 1591  ±  44·0 kg green dry matter (DM) ha^–1 respectively] maintained by continuous grazing of cattle at variable stocking rates. Four native and naturalized warm-season species were studied: the grasses Cynodon dactylon , Leersia hexandra and Paspalum dilatatum and the legume Lotus tenuis . Sward state of the four species was described in terms of sward surface height, herbage biomass and the number of individual grass tillers and legume stems per unit area. The defoliation pattern was monitored twice-weekly during an experimental period of 3 weeks on permanently marked individual plants. Proportions of individual plants defoliated and the ranking of species selected differed among sward treatments and species. The results showed that the density of the species was a major factor determining the pattern of species defoliation and suggest that, in continuously grazed mixtures, the pattern of defoliation might be more dependent on the abundance of a species in the plant community than on species preferences. The results show the importance of considering this structural characteristic of swards in understanding the process of species selection by large herbivores in multispecies vegetation.     

Response of herbage regrowth and water-soluble carbohydrate concentration of ryegrass species to defoliation practices when grown in a Mediterranean environment

Three experiments were conducted to determine the association between leaf number per tiller at defoliation, water‐soluble carbohydrate (WSC) concentration and herbage mass of juvenile ryegrass plants when grown in a Mediterranean environment. Seedlings of ryegrass were grown in nursery pots arranged side‐by‐side and located outside in the open‐air to simulate a mini‐sward in Experiments 1 and 2, and a mixture of annual ryegrass and subterranean clover (Trifolium subterraneum L.) was grown in a small plot field study in Experiment 3. Swards were defoliated mechanically with the onset of defoliation commencing within 28 d of germination. Frequency of defoliation ranged from one to nine leaves per tiller, whilst defoliation height ranged from 30 mm of pseudostem height that removed all leaf laminae in Experiment 1, to 50 mm of pseudostem height with some leaf laminae remaining post‐defoliation in Experiments 2 and 3. A positive relationship between herbage mass of ryegrass, WSC concentration and leaf number per tiller at defoliation was demonstrated in all experiments. In Experiment 1, the herbage mass of leaf, pseudostem and roots of tillers defoliated at one leaf per tiller was reduced to 0·10, 0·09 and 0·06 of those tillers defoliated less frequently at six leaves per tiller. However, the reduction in herbage mass from frequent defoliation was less severe in Experiment 2 and coincided with a 0·20 reduction in WSC concentration of pseudostem compared with 0·80 measured during Experiment 1. In Experiment 3, the highest harvested herbage mass of ryegrass occurred when defoliation was nine leaves per tiller. Although the harvested herbage from this sward contained senescent herbage, the in vitro dry‐matter digestibility of the harvested herbage did not differ significantly compared with the remaining treatments that had been defoliated more frequently. Leaf numbers of newly germinated ryegrass tillers in a Mediterranean environment were positively associated with WSC concentration of pseudostem and herbage mass. A minimum period of two to three leaf appearances was required to restore WSC concentrations to levels measured prior to defoliation thereby avoiding a significant reduction in herbage mass. However, maximum herbage mass of a mixed sward containing ryegrass and subterranean clover was achieved when defoliation was delayed to nine leaves per tiller.     

Interactions between N supply and N uptake by perennial ryegrass, 15N recovery and soil pH for four acid Scottish soils

The effects of NH₄⁺−N, NO₃⁻-N or urea-N addition on N uptake by perennial ryegrass ( Lolium perenne ), ¹⁵N recovery and pH of four limed and unlimed soils were determined in a pot experiment over 10 weeks. The best form of N in terms of herbage N uptake and fertilizer recovery differed between the soils. Recovery of applied ¹⁵N in herbage was greatest for the soil with the lowest pH and highest organic matter content, and overall recovery in the soil–plant system was influenced more by soil type than by N form. There was always an apparent mineralization of soil N when perennial ryegrass was present. Soil pH changes ranged from +0·11 to −0·58 units for the unlimed soils and from +0·03 to −1·06 units for the limed soils. The use of NO₃⁻-N rather than NH₄⁺-N or urea-N avoided further acidification. Lime increased herbage N only from the soils with the lowest pH values. Although the acidifying effect of NH₄⁺-fertilizer was alleviated, liming may increase nitrification and possibly N loss via denitrification and/or leaching in the field.
The major influence of soil type on herbage N uptake, pH response to the treatments and fertilizer recoveries implies that liming and fertilizer N management decisions should consider soil characteristics, such as organic matter, clay contents and pH.     

Photosynthetic rate and carbon balance of grazed ryegrass pastures

The relationship between net canopy photosynthesis (Pnc) measured at 400 J m⁻² s-¹ and leaf area index (LAI) was determined on ryegrass-dominant swards over a range of her bage masses in five grazing experiments. In three experiments the swards were continuously stocked either by sheep or by cattle to maintain a number of herbage heights and hence LAI values. In two experiments the swards were intermittently grazed with 21 -d regrowth intervals. On the continuously stocked swards measurements were made over nine periods each of 7 to 21 d in length. On the intermittently grazed swards measurements were made over six periods while the herbage was being grazed down from a high to a low LAI and over three periods during the regrowth phase.
On all the swards where measurements were made while grazing was in progress, the relation of Pnc to LAI was linear. The photosynthetic rate was greater on the continuously grazed than on the intermittently grazed swards up to LAI 35 and in two experiments to LAI 5. The relationship of Pnc to LAI of swards in the regrowth phase was predominantly curvilinear and such that over much of the LAI range they had a greater photosynthetic potential than the same swards being grazed down from high to low mass. Daily rates of CO₂ uptake and growth in the field were calculated for one continuously stocked experiment. Both were linearly related to LAI in the range LAI 2–4·5. The agronomic implications of the results are discussed.     

Herbage growth rates on heterogeneous swards as influenced by sward-height classes

The contribution of four classes of sward height to daily herbage growth rates of a heterogeneous sward in eight periods throughout a grazing season was investigated in two continuous cattle-grazing systems differing in intensity (moderate stocking rate: MC; lenient stocking rate: LC). At the beginning and end of periods of 12 to 28 d, the compressed sward height (CSH) was measured in exclusion cages at eighteen fixed points per cage to derive daily growth rates for the four classes of sward height. Stratified calibrations were made to relate sward height to herbage mass for each treatment in each period. Quadratic regressions described the relationship between herbage growth rate and initial CSH for each treatment in each period. For scaling up to the scale of the plot, CSH was measured monthly at 100 points per plot. Daily herbage growth rates declined from more than 100 kg dry matter (DM) ha⁻¹ d⁻¹ on both treatments at the beginning of the grazing season to 20 kg DM ha⁻¹ d⁻¹ or less, especially on treatment LC. This was due to the larger area covered by tall herbage on treatment LC than on treatment MC. On treatment MC, daily herbage growth rate was predominantly derived from short sward areas of up to 12 cm in height while sward areas taller than 12 cm contributed most to daily growth rates on treatment LC in early summer. The method used is considered suitable for estimating daily herbage growth rates of different classes of sward height in extensively managed pastures and can easily be adapted to deal with more heterogeneous swards than used in this study.     

A plate meter inadequately estimated herbage mass in a semi-arid grassland

A resting plate meter was tested in the laboratory and on a field site to determine its effectiveness for estimating peak herbage mass on semi-arid grasslands. In laboratory tests, data from the plate meter closely predicted the herbage mass of four perennial bunch grasses (thirteen of sixteen tests with r ² ≥ 0·90) but the closeness of the prediction varied with the pressure of the plate. Field tests took place on a southern Arizona, USA semi-arid grassland in 2005, 2006, 2007 and 2008. Twenty teams of two or three people sampled the site; five teams in each year measured eight to thirteen plate heights and clipped the vegetation beneath. Consistent with a need for yearly calibration, a significantly different ( P  <   0·05) linear relationship was detected between 3 of the 4 years which was associated with differences in average herbage mass of dry matter (DM) (1525 kg ha⁻¹ for 2005, 2093 kg ha⁻¹ for 2006, 1338 kg ha⁻¹ for 2007 and 1370 kg ha⁻¹ for 2008). Plate height poorly predicted herbage mass within years ( r ²  = 0·21, 0·51, 0·49 and 0·41 respectively) with plate heights explaining half or less of the variability in field herbage mass and having a mean prediction error of 466 kg DM ha⁻¹. The plate meter technique had limited potential for estimating peak herbage mass in semi-arid grasslands.     

Effects of concentrate supplementation and herbage allowance on the performance of grazing suckling lambs

An experiment was conducted to examine the effect of concentrate supplementation on the performance of suckling lambs while grazing at variable levels of herbage allowance. Twenty-four ewes suckling twin lambs were allocated 55 d after lambing between four grazing treatments (two levels of herbage allowance × two levels of supplementation) in a 2×2 factorial design. The sward consisted of Tall Fescue and was grazed rotationally.
Daily herbage organic matter (OM) allowances were 57.0 (H) and 38.5 (L) g OM kg LW⁻¹ above 1·5 cm cutting height. Lambs were either supplemented (S) ad libitum with high-moisture whole maize grain or unsupplemented (U).
Supplement intake during the 60-d grazing period was 16 kg DM and 17.5 kg DM respectively for LS and HS lambs. The effect of supplementation on lamb growth rate differed significantly between allowances: at the low allowance level, supplementation increased growth rate (287 g d⁻¹ vs 226 g d⁻¹), whereas it had no effect at the high level (277 g d⁻¹ vs 276 g d⁻¹). Growth rate of unsupplemented lambs was significantly reduced at the low level of allowance compared to the high level (226 g d⁻¹ vs 276 g d⁻¹). The effect of sward height on time spent grazing by unsupplemented lambs is described during the defoliation of a plot.     

The effect of herbage allowance on intake and performance of ewes suckling twin lambs

The effect of herbage allowance on the herbage intake and performance of ewes and their twin lambs at pasture was investigated. Daily herbage allowances of 40,80.120 and 160 g organic matter (OM) kg^-1 ewe live weight, based on herbage mass measured to ground level, were offered during the first 12 weeks of lactation.
The sheep were grazed rotationally around four paddocks of a perennial ryegrass pasture for 7-day periods and herbage mass, extended tiller length and digestibility of the herbage consumed by the animals were estimated. Herbage intake by the ewes was estimated during weeks two to twelve and live weights were recorded weekly: during the last two weeks of the experiment grazing behaviour of one ewe on each treatment was recorded continuously.
Herbage intakes by the ewes (164, 1–81, 2–42 and 268 ± 0153 kg d^-1) and live weight gains of the lambs (202, 245, 274 and 300 ± 7–3 g d^-1) increased with increasing herbage allowances. Herbage intakes by the ewes and growth rates of their lambs increased up to a herbage allowance which was over five times the amount of herbage eaten by the ewes.
As the animals reduced herbage mass and sward height, biting rates by the ewes during grazing increased by 4 (± 0·08) bites min^-1 cm^-1 and masticating rates decreased. Mastications reached a maximum of 90 (± 3·5) min^-1 at a sward surface height of 9 cm.     

Dynamics of perennial ryegrass and white clover in sown swards in NW Greece

Seasonal dynamics of white clover and perennial ryegrass were examined in sown perennial ryegrass/white clover swards subject to a 2 × 2 factorial treatment combination of defoliation (rotational grazing by sheep and cutting) and nitrogen fertilizer application (0 and 40 kg N ha^–1 year^–1) in NW Greece. Sward surface height and percentage cover were measured before and after five defoliation periods in 1996 within permanent microplots (30 × 30 cm, divided into nine cells) in which white clover was either initially present or absent. Both white clover and perennial ryegrass achieved maximum height and cover in April–May. Defoliation treatment and whether white clover was present initially significantly affected height and cover of both species. Total plant cover was similar prior to all defoliation periods except in July, a time of drought. Cover of perennial ryegrass was greater where white clover was initially absent, but total plant cover was greater in microplots containing white clover and the extent of the differences varied during the year. In contrast, N fertilizer application had little effect on species cover, other than small reductions in white clover cover. When white clover was present in April, it was found in virtually every microplot cell until July, but if it was absent in April there was little colonization of the microplot.     

Effect of regrowth interval on the productivity of swards defoliated by cutting and grazing

Over three grazing seasons (1984-1986) a sward of perennial ryegrass, cv. Talbot, which received a total of 336 kg N ha^-1 each season, was cut or grazed with ewes at 3- or 4-week intervals on a rotational basis.
Sward productivity was higher under cutting than under grazing irrespective of the interval between defoliations. Under cutting, mean herbage organic matter (OM) yields over both intervals were 8·66, 9·62 and 8·17 t ha-¹ in 1984, 1985 and 1986 respectively while under grazing the corresponding yields were 7·65, 8·63 and 7·50 t ha^-1. The mean annual yield of herbage defoliated at 3-week intervals was 7·50, 8·64 and 7 ·20 t OM ha^-1 compared with 8·80, 9·60 and 8·46 t OM ha^-1 for swards defoliated at 4-week intervals in the three years respectively.
The nitrogen (N) content of both the available and the residual herbage was consistently higher under grazing than under cutting. Available herbage contained 31·3 and 27·7 g N kg OM^-1 and residual herbage 26·1 and 22·7 g N kg OM^-1 under grazing and cutting respectively.
The mean yield of N under cutting was 284 kg ha^-1 compared with 304 kg ha^-1 under grazing. Defoliation interval had no effect on N yield, the overall mean yield being 294 kg ha^-1 under both 3- and 4-week defoliation intervals. The effect of the treatments on tiller population was slight and inconclusive.
The process of grazing reduced yield probably as a result of damage to the sward through trampling; the positive effect of excretal N on yield was minimal on account of the short grazing periods.     

Seasonal patterns of grass production under cutting or continuous stocking managements

A comparison was made between the seasonal pattern of net herbage accumulation under cutting and the pattern of intake under continuous variable stocking on a Lolium perenne sward, which was well supplied with nitrogen fertilizer and irrigated to minimize moisture stress.
The seasonal pattern of production under monthly cutting showed characteristic asymmetry with a marked peak in production in late May/early June, a rapid decline in June followed by an increase in rate in August and the total amount harvested over the season was 9–28 t organic matter ha⁻¹.
By contrast, the seasonal patterns of production on swards grazed by ewes to maintain sward surface heights of 30, 50 and 70 mm all differed fundamentally from that observed under cutting. Production was more evenly distributed throughout the season under these grazing treatments which, it has been proposed, should optimize performance. Total amounts harvested by grazing were 10–84, 10.12 and 8.51 t OM ha⁻¹, respectively. The different patterns of production under cutting and grazing were considered to be associated with differences in the expression of reproductive development in the grass crop and the maintenance of a smaller leaf area per unit ground area under grazing.
The results of the study suggest that it is not appropriate to regard the seasonal pattern of net accumulation under a standard cutting regime as representing the potential production of grass under continuous stocking. Moreover, in practice, the more uniform seasonal pattern of production seen in swards maintained on the basis of a controlled sward surface height makes it easier to match the supply of herbage to the appetite of grazing animals as the season progresses.     

Dry-matter yield and herbage quality of a perennial ryegrass/white clover sward in a rotational grazing and cutting system

The expected reduction in the use of fertilizer nitrogen (N) on grassland in the Netherlands has led to renewed interest in white clover. Therefore, the performance of a newly sown perennial ryegrass/white clover sward on clay soil was assessed during 4 consecutive years. The experiment consisted of all combinations of two defoliation systems, i.e. one or two silage cuts per year (S1, S2), spring N application rate, i.e. 0 or 50 kg ha⁻¹ year⁻¹ (N₀, N₅₀), and the management system, i.e. rotational grazing and cutting, or cutting only (RGC, CO). The overall mean white clover cover was 30%. All treatments affected white clover cover, which was 8% higher with S2 than with S1, 6% higher with N₀ than with N₅₀ and 12% higher with CO than with RGC. The overall mean annual dry-matter (DM) yield (13·1 t ha⁻¹ year⁻¹) was significantly affected only by the management system: in two relatively wetter years, the annual DM yield was 1·19 t ha⁻¹ higher with RGC than with CO, whereas there was no difference in two relatively drier years. Nitrogen application increased the DM yield in the first cut by 7·0 kg kg⁻¹ N applied, but had no significant effect on the annual DM yield. Herbage quality was not affected by the experimental treatments. The average in vitro organic matter digestibility was 0.801, and the average crude protein content was 193 g kg⁻¹ DM. With the expected reduction in the use of fertilizer N, perennial ryegrass/white clover swards should be seriously considered as an alternative option to perennial ryegrass swards on these clay soils.     

Seasonal changes in the crude protein concentration of mixed swards of white clover/perennial ryegrass grown without fertilizer N in an organic farming system in the United Kingdom

Changes in the crude protein (CP) concentration of white clover and perennial ryegrass herbage from a mixed sward were determined on six sampling dates from May to October in each of 2 years. The swards were grown without fertilizer N in an organic farming system and continuously grazed by dairy cows during the grazing season. The annual mean contents of white clover in the dry matter (DM) of the sward were 272·3 and 307·0 g kg⁻¹ in Years 1 and 2. The mean CP concentrations of the white clover and perennial ryegrass herbage were 251·6 and 151·9 g kg⁻¹ DM in Year 1 and 271·9 and 174·0 g kg⁻¹ DM in Year 2 respectively. The CP concentration of the white clover increased significantly during the grazing season from 220·0 to 284·1 g kg⁻¹ DM in Year 1 and from 269·0 to 315·5 g kg⁻¹ DM in Year 2. In the perennial ryegrass herbage the CP concentration increased from 112·2 to 172·6 g kg⁻¹ DM in Year 1 and from 142·7 to 239·5 g kg⁻¹ DM in Year 2. The rate of increase during the season in the CP concentration of the perennial ryegrass herbage was similar to the rate of increase recorded in the white clover herbage.     

The liveweight gain of Limousin × Friesian heifers grazing perennial ryegrass/white clover swards of different clover content and the effects of their grazing on sward botanical composition

An experiment was carried out in 1992 and 1993 to examine the effect of white clover content of perennial ryegrass/white clover swards on the performance of Limousin × Friesian heifers. Swards with low (L), medium (M) and high (H) white clover contents were established and managed by continuous variable stocking. A compressed sward height of 5·5 cm was maintained using a buffer fence to vary plot areas, with herbage surplus to grazing requirements cut, removed and yields measured. The mean white clover proportions for treatments L, M and H were 0·02, 0·19 and 0·18 in 1992 and 0·13, 0·16 and 0·31 in 1993 respectively. White clover contents of the swards reached a maximum in August and September, and differences between treatments diminished. There was no significant difference between treatments in the content of white clover in the swards in autumn 1993.
Liveweight gains of heifers increased asymptotically with increasing white clover content of the sward. Below a white clover herbage mass of 300 kg DM ha^–1, there was little effect on liveweight gain, which was 0·70 kg day^–1 over the grazing season. Between 400 and 450 kg DM ha^–1 white clover, liveweight gains were 0·85–0·90 kg day^–1. While clover content of the sward did not significantly affect utilized metabolizable energy output; the mean output over the grazing season in the two years from liveweight gain and herbage yield was 78 GJ ha^–1. It is suggested that, using this grazing system, white clover reached an equilibrium with a mean herbage mass of about 400 kg DM ha^–1 over the grazing season.