Winter cover crop: the effects of grass–clover mixture proportion and biomass management on maize and the apparent residual N in the soil

The advantages and disadvantages of varying mixture proportion of crimson clover (Trifolium incarnatum L.) and Italian ryegrass (Lolium multiflorum Lam.), used as winter cover crops, and cover crop biomass management before maize sowing (Zea mays L.) were studied in a series of field experiments in Eastern Slovenia. Pure stands and mixtures of cover crops on the main plots were split into different cover crop biomass management subplots: whole cover crop biomass ploughed down before maize sowing, aboveground cover crop biomass removed before ploughing and sowing, or aboveground cover crop biomass removed before sowing directly into chemically killed residues.Cover crop and cover crop biomass management affected the N content of the whole aboveground and of grain maize yields, and the differences between actual and critical N concentrations in the whole aboveground maize yield. The whole aboveground and grain maize dry matter yields, and the apparent remaining N in the soil after maize harvesting, showed significant interaction responses to cover crop × management, indicating positive and negative effects. Crimson clover in pure stand provided high, and pure Italian ryegrass provided low maize dry matter yields and N content in the yields in all the observed methods of biomass management. However, within individual management, mixtures containing high proportions of crimson clover sustained maize yields and N contents similar to those produced by pure crimson clover. Considering the expected ecological advantages of the mixtures, the results thereby support their use.     

Cover crops and interrow tillage for weed control in short season maize (Zea mays)

Weed competition can cause substantial maize (Zea mays L.) yield reductions. Interseeding maize with cover crops or a combination of interrow cultivation and interseeded cover crops are possible alternative methods of weed control. This study was conducted to examine the potential of interrow cultivation plus cover crops to reduce weed density in maize without reducing the grain yield. Field experiments were conducted in 1993 and 1994 at two sites in Québec to determine the effects of planting 12 cover crops with maize on weed control. Fall rye (Secale cereal L.), hairy vetch (Vicia villosa Roth), a mixture of red clover (Trifolium pratense L.) and ryegrass (Lolium multiflorum Lam), a mixture of white clover (Trifolium repens L.) and ryegrass, subterranean clover (Trifolium subterraneum L.), yellow sweet clover (Meliotus officinalis Lam), black medic (Medicago lupulina L.), Persian clover (Trifolium resupinatum L.), strawberry clover (Trifolium fragiferum L.), crimson clover (Trifolium incarnatum L.), alfalfa (Medicago sativa L.), and berseem clover (Trifolium alexandrinum L.) were seeded at two planting dates, 10 and 20 days after maize emergence. Interrow cultivation was carried out weekly until forage seeding, with a final cultivation being conducted just prior to cover crop seeding. Cover crop planting date did not affect maize yields or the ability of interrow tillage plus cover crops to suppress the development of weed populations. Maize yield was less affected by the interseeded cover crops under conditions of adequate rainfall. Corn planted in fields heavily infested with weeds resulted in substantial yield reductions even when rainfall was adequate. Except for 1993 at l'Assomption interrow tillage plus cover crop treatments had consistently lower weed biomass when compared to the weedy control. Most of the weed control was due to the interrow cultivation performed prior to seeding of the cover crops. The lowest weed density occurred in the herbicide treated plots. The ability of interrow tillage plus cover crops to suppress the development of weeds was affected by the level of weed infestation, the growing conditions and location. The cover crops provide additional weed control but the interrrow tillage or some herbicide application may still be necessary.     

Legume cover crops as living mulches for winter wheat: Components of biomass and the control of weeds

To gain information about the possible use of legume cover crops as an alternative and sustainable weed-control strategy for winter wheat (Triticum aestivum L.), an experiment was conducted at two sites in the Swiss Midlands in 2001/2002. Under organic farming conditions winter wheat was direct-drilled into living mulches established with four different legume genotypes or into control plots without cover crops. Compared to NAT (control plots without cover crops but with a naturally establishing weed community), white clover (Trifolium repens L.), subclover (Trifolium subterraneum L.), and birdsfoot trefoil (Lotus corniculatus L.) reduced the density of monocotyledonous, dicotyledonous, spring-germinating, and annual weeds by the time of wheat anthesis. Strong-spined medick (Medicago truncatula Gaertner) was less efficient in this regard. While the grain yield was reduced by 60% or more for all legumes when compared to NOWEED (control plots kept weed-free), a significant negative correlation between the dry matter of the cover crop and weeds as well as between the cover crop and the winter wheat was observed by the time of wheat anthesis. The effect of manuring (60 m³ ha⁻¹ liquid farmyard manure) was marginal for weeds and cover crops but the additional nutrients significantly increased total winter wheat dry matter and grain yields. The suppression achieved by some legumes clearly demonstrates their potential for the control of weeds in such cropping systems. However, before living legume cover crops can be considered a viable alternative for integrated weed management under organic farming conditions, management strategies need to be identified which maximise the positive effect in terms of weed control at the same time as they minimise the negative impact on growth and yield of winter wheat.     

Assessing innovative sowing patterns for integrated weed management with a 3D crop:weed competition model

Weed dynamics models are needed to design innovative weed management strategies. Here, we developed a 3D individual-based model called FlorSys predicting growth and development of annual weeds and crops as a function of daily weather and cropping practices: (1) crop emergence is driven by temperature, and emerged plants are placed onto the 3D field map, depending on sowing pattern, density, and emergence rate; plants are described as cylinders with their leaf area distributed according to height; (2) weed emergence is predicted by an existing submodel, emerged weed seedlings are placed randomly; (3) plant phenology depends on temperature; (4) a previously developed submodel predicts available light in each voxel of the canopy; after emergence, plant growth is driven by temperature; when shaded, biomass accumulation results from the difference between photosynthesis and respiration; shading causes etiolation; (5) frost reduces biomass and destroys plants, (6) at plant maturity, the newly produced seeds are added to the soil seed bank. The model was used to test different sowing scenarios in an oilseed rape/winter wheat/winter barley rotation with sixteen weed annuals, showing that (1) crop yield loss was negatively correlated to weed biomass averaged over the cropping season; (2) weed biomass was decreased by scenarios allowing early and homogenous crop canopy closure (e.g. reduced interrows, increased sowing density, associated or undersown crops), increased summer fatal weed seed germination (e.g. delayed sowing) or, to a lesser degree, cleaner fields at cash crop sowing (e.g. sowing a temporary cover crop for “catching” nitrogen); (3) the scenario effect depended on weed species (e.g. climbing species were little affected by increased crop competition), and the result thus varied with the initial weed community (e.g. communities dominated by small weed species were hindered by the faster emergence of broadcast-sown crops whereas taller species profited by the more frequent gap canopies); (4) the effect on weed biomass of sowing scenarios applied to one year was still visible up to ten years later, and the beneficial effect during the test year could be followed by detrimental effects later (e.g. the changed tillage dates accompanying catch crops reduced weed emergence in the immediately following cash crop but increased seed survival and thus infestation of the subsequent crops). This simulation showed FlorSys to predict realistic potential crop yields, and the simulated impact of crop scenarios was consistent with literature reports.     

Effect of spring fertilization on ecosystem services of organic wheat and clover relay intercrops

Nitrogen (N) deficiency and weed infestation are main factors limiting yield and yield stability in organic wheat. Organic fertilizers may be used to improve crop performance but off-farm input costs tend to limit profitability. Instead, forage legumes may be inserted into the crop rotation to improve the N balance and to control weed infestation. In opposition to simultaneous cropping, relay intercropping of legumes in organic winter wheat limits resource competition for the legume cover crop, without decreasing the performance of the associated wheat.The aim of this study is to evaluate the effect of spring organic fertilization on the performance of intercropped legumes and wheat, and on services provided by the legume cover.Two species of forage legumes (Trifolium pratense L. and Trifolium repens L.) were undersown in winter wheat (Triticum aestivum L. cv Lona) in five organic fields during two consecutive crop seasons. Organic fertilizer was composed of feather meal and applied on wheat at legume sowing. The cover crop was maintained after the wheat harvest and destroyed just before sowing maize.Spring organic nitrogen fertilization increased wheat biomass (+35%), nitrogen (+49%), grain yield (+40%) and protein content (+7%) whatever the intercropping treatment. At wheat harvest, red clover biomass was significantly higher than white clover one (1.4 vs. 0.7 t ha⁻¹). Nitrogen fertilization decreased forage legume above-ground biomass at wheat harvest, at approximately 0.5 t ha⁻¹ whatever the specie. No significant difference in forage legume biomass production was observed at cover killing. Nitrogen accumulation in legume above-ground tissues was significantly higher for white clover than for red clover. Both red and white clover species significantly decreased weed infestation at this date. Nitrogen fertilization significantly increased weed biomass whatever the intercropping treatment and decreased nitrogen accumulation in both clover species (−12%).We demonstrated that nitrogen fertilization increased yield of wheat intercropped with forage legume while the performance of legumes was decreased. Legume growth was modified by spring fertilization whatever the species.     

Clover as a cover crop for weed suppression in an intercropping design: II. Competitive ability of several clover species

Undersown cover crop species introduced for weed management purposes should ideally combine adequate weed suppression with only marginal negative competitive effects on the main crop. The aim of this research was to identify the growth characteristics of clover species that determine weed suppressive ability and competitive ability against the main crop. In addition, the variation in these characteristics among clover species was determined, to identify whether species selection is an important component of the optimization of living mulch systems.In 2001, a field experiment was conducted in which weed suppression and competitive ability against leek of eight clover types was determined. Based on the results, three contrasting clover types were selected and their weed suppressive ability was further investigated in a field experiment conducted in 2002. Results of both experiments were related to growth characteristics that were simultaneously determined and described in a previous paper.Correlation analysis showed that the competitive effect of clover on transplanted leek was strongly correlated with maximum canopy height, indicating that yield reduction was mainly caused by competition for light. Subterranean clover, being the shortest species, gave inadequate weed suppression, whereas all other species were found to provide at least reasonable weed suppression. Subterranean clover reduced individual leek plant dry weight more strongly (60%) than expected based on its canopy height. Indications were found that this clover species was hindering the uptake of nitrogen by leek. Clover species with superior weed suppression, like Persian clover, red clover, alsike clover, berseem clover and crimson clover also gave the strongest negative effect on dry matter accumulation of leek (reductions between 70% and 90%). White clover was found to give the best compromise between adequate weed suppression and mild yield reduction, though also with this species reduction in leek plant dry weight was still 60%. It was concluded that for optimization of systems that introduce clover species as a weed suppressing cover crop species selection is important element, though additional management to restrict the competitive effect on the main crop remains a requirement.     

The influence of organic and conventional fertilisation and crop protection practices,preceding crop,harvest year and weather conditions on yield and quality of potato (Solanum tuberosum) in a long-term management trial

The effects of organic versus conventional crop management practices (fertilisation, crop protection) and preceding crop on potato tuber yield (total, marketable, tuber size grade distribution) and quality (proportion of diseased, green and damaged tubers, tuber macro-nutrient concentrations) parameters were investigated over six years (2004–2009) as part of a long-term factorial field trial in North East England. Inter-year variability (the effects of weather and preceding crop) was observed to have a profound effect on yields and quality parameters, and this variability was greater in organic fertility systems. Total and marketable yields were significantly reduced by the use of both organic crop protection and fertility management. However, the yield gap between organic and conventional fertilisation regimes was greater and more variable than that between crop protection practices. This appears to be attributable mainly to lower and less predictable nitrogen supply in organically fertilised crops. Increased incidence of late blight in organic crop protection systems only occurred when conventional fertilisation was applied. In organically fertilised crops yield was significantly higher following grass/red clover leys than winter wheat, but there was no pre-crop effect in conventionally fertilised crops. The results highlight that nitrogen supply from organic fertilisers rather than inefficient pest and disease control may be the major limiting factor for yields in organic potato production systems.     

Model evaluation of cover crops,application to eleven species for banana cropping systems

Cover crops are increasingly used for weed management. But selecting the most suitable species of cover crop to be associated with a main crop requires long-term trials. We present a model-based method that uses a reduced number of parameters to help select cover crops in the context of banana cover-cropping systems. We developed the SIMBA-CC model to focus on radiation interception. The model was calibrated for 11 cover crop species by measuring their growth in 4 m² plots with three levels of shade (0, 50, and 75%). The SIMBA-CC model served to predict the long term growth potential of the 11 cover crop species in function of the radiation under the banana crop canopy. The model was validated using three species in association with banana plants. We defined three indicators based on outputs of the model to assess the ability of each of the 11 species (i) to compete with weeds and (ii) to be maintained in the long-term under the canopy of the main crop, and (iii) to evaluate competition with the main crop for nitrogen resource. This ex ante evaluation revealed the most promising species to be intercropped with banana. Finally, the SIMBA-CC model was used to define the light interception traits of a virtual cover crop that satisfy the three indicators in the case of intercropping with banana. We showed that to satisfy the three criteria, cover crops with low values of optimal photosynthetically active radiation (PARopti) should have moderate maximal biomass productivity, while crops with higher PARopti values should have a higher maximal productivity. The use of functional traits and modeling appears effective to disentangle the relations between intrinsic traits of cover crops and effect traits that affect the performances of the intercropping system.     

Dynamisches Entscheidungsmodell zur Lenkung von Unkrautkontrollmaßnahmen in Wintergetreide mit Hilfe digitaler Bildverarbeitung

Dynamic decision model for weed control methods in cereals by means of digital image analysis
Field experiments were conducted to study weed-crop competition in winter wheat, -barley and -rye. Weed cover and crop cover was determined over time by digital image analysis of slides taken in the field. The relation of percentage weed coverage and crop coverage was most suitable to quantify the competition of weed population on the crop. In addition the model "UNPROG" was established to predict yield losses caused by weed competition and to estimate the profitability of chemical or mechanical weed control. The model was tested on 22 farms in Nordrhein-Westfalia in autumn 1991 and spring 1992. In 65 out of 72 field plots (92 %) of winter cereals "UNPROG" gave correct prediction with respect to necessity of weed control.     

Effects of organic farming on weed flora composition in a long term perspective

In 1987, the Ekhaga Experimental Farm in Sweden was established on a site that previously had been subjected to conventional farming, and has been managed since then as an organic farm. To study the effects of organic farming on weed population development and crop yields, two different crop rotations were designed, one adapted for animals (six fields) and one without animals (six fields). Each field contained a fixed 1 m² reference plot in which all the weed observations were done each year. During the period 1988–2002, number of weed plants in spring and weed biomass at harvest were recorded in the reference plots. No differences in these two parameters were observed between the crop rotations. Number of weed plants in spring did not differ between annual crops and did not increase over the 15-year period. Neither did weed biomass at harvest nor weed species diversity change over the 15 years. The two crop rotations kept weed pressure at the same level as under the previous conventional farming practice. General field observations suggest that invasion of Cirsium arvense (L.) Scop. is occurring along the field borders. Competitive ability of the crop showed to be important in weed regulation. Peas, a weak competitor, had significantly higher weed biomass at harvest compared with oats and winter wheat. Weather conditions during the period from April to September caused weed occurrence and development through the season to vary between years. To improve weed management in organic farming, advisors and farmers should recognise the importance of individual field and farm analyses to design location-specific, farm-adapted crop rotations.     

Shortening generation times for winter cereals by vernalizing seedlings from young embryos at 10 degree Celsius

Methods allowing quick generation cycles are available for various crop species but are limited to spring genotypes. However, winter types predominate in many crops including wheat and barley. Different from spring ones, winter genotypes need to be treated in low temperature for extended periods to accelerate flowering. Combined with known factors that reduce generation cycles for spring wheat and barley, the generation cycle effects of vernalizing seedlings from young embryos of winter types at 10°C of several crop species were studied. Here, we described how to obtain up to seven generations of winter barley, six generations for winter wheat or five generations for both winter oat and triticale genotypes per annum. This procedure should find wide applications in breeding and other biological studies.     

Effects of cover crops on soil mineral nitrogen and on the yield and nitrogen content of maize

Current agricultural practice favours winter cover crops, which can not only optimize N management in field crop rotation; but also affect subsequent crops. Three field experiments were carried out in Eastern Slovenia to examine the effects of Italian ryegrass (Lolium multiflorum Lam.), winter rape (Brassica napus ssp.oleifera (Metzg.) Sinsk), subclover (Trifolium subterraneum L.), and crimson clover (Trifolium incarnatum L.) as winter cover crops on the mineral N (N_min) content of soil and on the yield and N content of subsequent maize (Zea mays L.), fertilized with 120 kg N ha⁻¹. Italian ryegrass and winter rape decreased soil N_min contents before winter and in spring more than both clovers. In contrast, clovers accumulated significantly higher amounts of N in organic matter and had lower C/N ratios than winter rape and especially Italian ryegrass. In comparison to the control (bare fallow without cover crop), clovers increased the whole above ground maize dry matter yield, maize grain yield and N contents in whole above ground plants and in grain. The yields and N contents of maize following winter rape were on the same level as the control, while yields and N contents of maize following Italian ryegrass were, in two of the experiments, at the same level as the control. The effects of Italian ryegrass on the maize as subsequent crop in the third experiment were markedly negative. Maize in the control treatment exploited N much more efficiently than in treatments with cover crops. Therefore, cover crop N management should be improved, especially with a view to optimizing the timing of net N mineralization in accordance with the N demands of the subsequent crop.     

Impact of Soil Tillage and Crop Rotation on Barley (Hordeum vulgare) and Weeds in a Semi-arid Environment

Experiments were conducted to evaluate the effect of mouldboard‐ or chisel‐ploughing and rotations on barley crops and associated weeds in a semi‐arid location. Two primary soil tillage operations and eight crop rotation‐tillage operation combinations were evaluated over two successive seasons. Drought conditions prevailed (<152 mm annual precipitation) and affected the measured parameters. Barley grown in mouldboard‐ploughed plots had higher biomass compared with chisel‐ploughed plots. Barley grain yield was greater in mouldboard‐ploughed plots in a fallow‐fallow‐barley rotation. Weed species densities varied between tillage systems and rotations. Density of Hordeum marinum, for example, was high in fallow‐barley‐fallow in chisel‐ploughed plots, and was high under more continuous fallow in mouldboard‐ploughed plots. Similar variations were also observed in weed fresh weights and in numbers of seed produced. The results describe the productivity of barley under extremely dry conditions, where an advantage for mouldboard ploughing was observed. The results also indicate the complexity of weed communities in their response towards different tillage‐rotation combinations.     

Multi-model uncertainty analysis in predicting grain N for crop rotations in Europe

Realistic estimation of grain nitrogen (N; N in grain yield) is crucial for assessing N management in crop rotations, but there is little information on the performance of commonly used crop models for simulating grain N. Therefore, the objectives of the study were to (1) test if continuous simulation (multi-year) performs better than single year simulation, (2) assess if calibration improves model performance at different calibration levels, and (3) investigate if a multi-model ensemble can substantially reduce uncertainty in reproducing grain N. For this purpose, 12 models were applied simulating different treatments (catch crops, CO₂ concentrations, irrigation, N application, residues and tillage) in four multi-year rotation experiments in Europe to assess modelling accuracy. Seven grain and seed crops in four rotation systems in Europe were included in the study, namely winter wheat, winter barley, spring barley, spring oat, winter rye, pea and winter oilseed rape. Our results indicate that the higher level of calibration significantly increased the quality of the simulation for grain N. In addition, models performed better in predicting grain N of winter wheat, winter barley and spring barley compared to spring oat, winter rye, pea and winter oilseed rape. For each crop, the use of the ensemble mean significantly reduced the mean absolute percentage error (MAPE) between simulations and observations to less than 15%, thus a multi–model ensemble can more precisely predict grain N than a random single model. Models correctly simulated the effects of enhanced N input on grain N of winter wheat and winter barley, whereas effects of tillage and irrigation were less well estimated. However, the use of continuous simulation did not improve the simulations as compared to single year simulation based on the multi-year performance, which suggests needs for further model improvements of crop rotation effects.     

Effect of Sowing Technique on Growth of Undersown Crop and Yield of Spring Barley

Undersowing a main crop enables establishment of a catch crop in areas characterized by a short post-harvest period before the onset of winter. Techniques with lower costs than conventional undersowing by separate drilling are often regarded as unreliable. Undersowing by drilling after sowing spring barley ( Hordeum vulgare L.) was compared with broadcast sowing simultaneously with drilling barley. Various implements were coupled behind the combined drill in cases where seed was broadcast: a press-wheel attachment, a long-tined harrow and a cage roller. A fourth treatment did not include an implement coupled behind the drill. The undersown crop was sown as a seed mixture of 3 kg ha^–1 red clover ( Trifolium pratense L.) and 6 kg ha^–1 meadow fescue ( Festuca pratensis Hudson). The numbers of plants and weeds and the plant height were measured five times during the growing season. Above-ground biomass of the undersown species was determined at barley harvest and in late autumn. Grain yield of spring barley was recorded. Drilling resulted in the highest yield of undersown crop when an early summer drought occurred, but broadcasting in combination with use of seed covering equipment led to the least variation in biomass production over the 4 years the experiment was conducted. The relative proportion of meadow fescue in the crop was low in three years, and lower when broadcast than when drilled. Barley grain yield was highest when the seed was broadcast and seed covering equipment was used. Use of a cage roller increased weed biomass, but press-wheels and a long-tined harrow did not. Separate rolling after undersowing increased undersown crop yield in one year, but decreased grain yield in some cases.     

Response of Weed Communities to Legume Living Mulches in Winter Wheat

In order to obtain information about the impact of legume cover crops on the weed community in organic farming, winter wheat (Triticum aestivum L.) was directly drilled in rows 0.1875 and 0.3750 m apart in living mulches that consisted of Trifolium repens L. (TRFRE), T. subterraneum L. (TRFSU), Medicago truncatula Gaertner (MEDTR), and Lotus corniculatus L. (LOTCO). A control treatment without cover crops (NAT, the site‐specific weed community) was also established. The vegetation between the wide rows was either mulched or left undisturbed. The effect of liquid farmyard manure (60 m³ ha⁻¹) was also tested. TRFRE, TRFSU, and LOTCO effectively suppressed Poa annua L. and Matricaria recutita L. at site 1 and P. annua, Capsella bursa‐pastoris (L.) Med., and Stellaria media (L.) Vill. at site 2 when compared with NAT. MEDTR, which died during the winter, provided little weed suppression. Mulching significantly suppressed dicotyledonous weed species, but favoured Poa trivialis L. No manure effect was observed. Winter hardy legume cover crops contribute to weed suppression in winter wheat. However, careful evaluation of cover crop × weed × management interactions is necessary to understand the risk for the establishment of problematic weeds.     

Clover as a cover crop for weed suppression in an intercropping design: I. Characteristics of several clover species

Weeds often form a major problem in weakly competitive vegetable crops, particularly in low input systems. Undersown cover crops can be used to suppress weeds, but often put too high a competitive pressure on the main crop. Cover crop selection is one of the potential means that can be used to design or optimize these intercropping systems. The objective of the current research was to investigate the variability among a range of clover species in morphological and physiological traits that are considered relevant for interplant competition. To this purpose, field experiments with pure stands of eight clover species (2001) and a selection of three clover species (2002) were conducted, in which regular observations and periodic harvests were taken. Clear differences in the time in which full soil cover was obtained, total accumulated biomass, growth duration, height development and N-accumulation were observed. Persian clover (Trifolium resupinatum L.) and subterranean clover (T. subterraneum L.) were the two most contrasting species in this study, particularly differing in the period in which full soil cover was obtained. Persian clover's faster soil cover could not be attributed to a single trait, but resulted from a number of intrinsic characteristics, like light extinction coefficient, light use efficiency and specific leaf area that together determine the relative growth rate. The study also demonstrated the importance of differences in relative starting position, caused by, for instance, seed size, seeding rate and fraction establishment, for the analysis of early growth characteristics. Alsike clover (T. hybridum L.), berseem clover (T. alexandrinum L) and crimson clover (T. incarnatum L.) developed slower than Persian clover, but all produced a higher amount of accumulated dry matter, due to a longer growing period. Clear differences in height and height development between species were observed. These differences were not associated with dry matter accumulation, as the tallest (red clover; 80 cm) and the shortest species (subterranean clover; 12 cm) produced similar amounts of dry matter. A strong positive correlation between early soil cover development and N-accumulation was observed. The large variability among clover species indicates that species selection is a very important aspect of the development of cropping systems that include clover as a cover crop.     

Mechanical wounding under field conditions: A potential tool to increase the allelopathic inhibitory effect of cover crops on weeds?

To increase the inhibitory effect of soil-incorporated cover crop residues on germination and early growth of weeds, the allelochemical content of the cover crop at the time of soil incorporation should be maximal. We investigated whether mechanical damaging in spring induced the production of allelochemicals in late-summer sown lucerne, winter rye and winter oilseed rape. Allelopathic activity and biomass of intact cover crop plants were determined throughout spring, and mechanically damaged plants were monitored till 14 days after wounding. Allelopathic activity was determined directly by lettuce seedling bioassays (lucerne and winter rye) or indirectly by glucosinolate quantification (winter oilseed rape). During spring, the allelopathic activity per unit biomass of intact plants showed a gradual decline for winter rye and lucerne, and a steep decline at the onset of flowering for winter oilseed rape. All cover crop species attained the highest allelopathic activity per unit area at the end of the sampling period as the increase in biomass surpassed the decline in allelopathic activity per unit biomass. Although mechanical wounding enhanced the allelopathic activity per unit biomass of all three species, the effect was only minor and often just sufficient to compensate for the loss in biomass resulting from wounding. This study therefore indicates that the best option for maximizing the inhibitory effect is to incorporate residues of intact cover crops as late as possible.     

Relay-intercropped forage legumes help to control weeds in organic grain production

In organic grain production, weeds are one of the major limiting factors along with crop nitrogen deficiency. Relay intercropping of forage legume cover crops in an established winter cereal crop might be a viable option but is still not well documented, especially under organic conditions.Four species of forage legumes (Medicago lupulina, Medicago sativa, Trifolium pratense and Trifolium repens) were undersown in six organic wheat fields. The density and aerial dry matter of wheat, relay-intercropped legumes and weeds were monitored during wheat-legume relay intercropping and after wheat harvest until late autumn, before the ploughing of cover crops.Our results showed a large diversity of aerial growth of weeds depending on soil, climate and wheat development. The dynamics of the legume cover crops were highly different between species and cropping periods (during relay intercropping and after wheat harvest). For instance, T. repens was two times less developed than the other species during relay intercropping while obtaining the highest aerial dry matter in late autumn. During the relay intercropping period, forage legume cover crops were only efficient in controlling weed density in comparison with wheat sole crop. The control of the aerial dry matter of weeds at the end of the relay intercropping period was better explained considering both legumes and wheat biomasses instead of legumes alone. In late autumn, 24 weeks after wheat harvest, weed biomass was largely reduced by the cover crops. Weed density and biomass reductions were correlated with cover crop biomass at wheat harvest and in late autumn. The presence of a cover crop also exhibited another positive effect by decreasing the density of spring-germinating annual weeds during the relay intercropping period.     

Zur Eignung nicht eingesäter Teilflächen in Getreideschlägen für die störungsfreie, bildanalytische Ermittlung von Unkrautschadschwellen

Suitability of not seeded test squares in crop fields for reliable ascertainment of weed threshold levels by quantitative image analysis
The possibility to measure weed ground cover using a computerized image analysis system has been reported in former investigations. In the present studies suitability of not seeded test squares in crop fields for reliable detection of weed ground cover for precise determination of weed threshold levels (beyond that crop yield losses are economically greater than costs for weed control) has been tested.
Test squares in which weed grew without competing crop as well as corresponding squares including both, growing crop and weeds, were examined for weed ground cover by computer assisted quantitative image analysis of color slides taken from the respective squares. In the squares, that includes crop and weeds, the former was removed immediately before taking the slides.
In both seasons from the beginning of weed growth in October to the latest feasible application time of herbicides in April weed growth as measured by ground cover in the not seeded test squares did not differ significantly from that grown under competition of winter barley.
Results indicate, that up to the growth stage of weed where herbicides if ever should be applied at the latest, crop-free test areas in the same field could serve for correct determination of weed growth by image analysis.