Spatial distribution patterns of Symplocos congeners in a subtropical evergreen broad-leaf forest of southern China

Studies on spatial distribution of congeneric species can supplement our understanding of species ecological processes. We analyzed population structure, spatial distribution, intra-and interspecific associations among six Symplocos tree species on a large(50 ha) plot in a subtropical, evergreen broad-leaf forest in southern China using spatial point pattern analysis methods. Our results suggested that the six Symplocos tree species were all clustered at small scales. The aggregation intensity of S. wikstroemiifolia with low abundance but relatively numerous large-diameter trees was much higher than that of the other five Symplocos species that occurred at high abundance but included few large-diameter trees. Spatial associations among the six congeners showed that 12 of 30 pairs were associated positively at small scales, and 13 of 30 pairs were unrelated. For species among different size classes, 79 of 120 pairs were not correlated, and 17 of 120 pairs were associated positively. These results showed insufficient evidence for interspecific competition and congeneric Symplocos species commonly coexist within subtropical plant communities. The spatial patterns of Symplocos species and their correlations changed with size(DBH) class and were simultaneously affected by spatial scales; the intensity of their aggregation and association decreased with increasing area of the sample plot.     

哀牢山植物多样性垂直分布格局

以设于哀牢山各植物群的样地调查材料为依据,分析了哀牢山东、西坡不同海拔高度地带性植物群落乔木层林木的高、径垂直分布特征,植物群落生物多样性指数的垂直分布态势。结果表明:该山地植物群落乔木层和灌木层的植物物种多样性沿海拔梯度呈单峰型分布格局,以中山湿性常绿阔叶林乔木层林木的最大和平均树高、最大胸径值和物种多样性指数最大;而灌木层植物物种多样性指数最大值出现在中山湿性常绿阔叶林带下的季风常绿阔叶林和半湿性常绿阔叶林带内;而草本层植物的物种多样性指数则沿海拔梯度上升呈整体减小的趋势;累加其植物群落乔、灌、草3层植物的物种多样性指数,哀牢山东、西坡植物群落的Shannon-Weiner指数和物种丰富度最大值出现在海拔高度2000 m左右。在相同海拔高度处,西坡植物群落的植物物种多样性高于东坡,季风常绿阔叶林的物种多样性大于半湿性常绿阔叶林,而山顶苔藓矮林的植物物种多样性最小。     

亚热带常绿阔叶林结构、树种多样性和土壤特性的研究(英文)

采用样地调查法,对日本冲绳岛北部的亚热带常绿阔叶林的林分结构,树种组成以及土壤特性等进行了调查分析。结果表明,该亚热带林具有林冠低矮、小径木多等特点。林分的平均林冠高度仅10m,平均立木密度达5400株·hm2(≥3.0cmDBH),其中胸径小于10cm的立木占64%。林分胸高断面积为54.4m2·hm-2。30个调查样地共出现树种数(≥3.0cmDBH)达80种,分属31个科。根据树种的重要值,Castanopsis sieboldii和Schima wallichii分别为优势树种和亚优势树种。树种多样性指数(H’)平均达3.63,均衡度指数(J’)为0.71,丰富度指数(S’)为4.72。这些多样性指数都随着优势树种重要值的增加而显著降低。土壤养分测定结果表明,该森林的土壤肥力低,异质性大,并可能存在Al毒害。由相关分析知,林分密度和优势树高均与土壤pH值呈显著正相关关系;多样性指数则与土壤交换性K ,Ca2 以及Ca2 /Al3 比呈显著的正相关,而与土壤有机碳,全氮和磷呈显著的负相关。这些结果说明土壤性质是影响冲绳亚热带常绿阔叶林林分结构和树种组成多样性的重要因素。图3表4参38。     

Scaling-up from tree to stand transpiration for a warm-temperate multi-specific broadleaved forest with a wide variation in stem diameter

Previous studies have demonstrated a clear relationship between diameter at breast height (DBH) and tree transpiration (Q _T) in multi-specific broadleaved forests. However, these studies were conducted with a limited range of tree sizes and species, and thus many multi-specific broadleaved forests fall outside these conditions. Therefore, this study examined the relationship between DBH and Q _T in a warm-temperate multi-specific broadleaved forest (n = 12 species) with a wide range of tree sizes (5.0–70.0 cm DBH) using the Granier-type heat dissipation method. The results showed that, although sap flow density varied between individual trees and species, there was a significant relationship between log Q _T and log DBH (r ² = 0.66, P < 0.001) because of the strong dependence of sapwood area on DBH. This study confirmed the applicability of the relationship for the stand transpiration (E _C) estimates even in a multi-specific broadleaved forest with a wide variation in DBH. Our results also revealed that selecting the sample trees in descending order of DBH effectively reduced potential errors in E _C estimates for a specific sample size, as larger trees contribute more to E _C. This information should be useful for future studies investigating the transpiration of multi-specific broadleaved forests, reducing errors during the scaling-up procedure.     

东莞清溪山地常绿阔叶林立木结构与多样性

文章研究了广东省东莞清溪山地常绿阔叶林群落组成和立木结构,并用丰富度指数、Shannon-Wiener多样性指数、均匀度指数及Simpson指数分析了群落物种多样性的变化。结果表明：林分内立木的树高和胸径变化总趋势均为小级别的个体占大多数,而且各主要树种的重要值差异不大,这些都表明该群落处于进展演替阶段;处于南亚热带的东莞清溪山地常绿阔叶林Shannon-Wiener多样性指数在0.673~2.539,物种均匀度为0.619~0.977,Simpson指数则为0.480~0.905,其物种多样性指数与陈和洞自然保护区常绿阔叶林的多样性指数相近。总的来说,东莞清溪山地常绿阔叶林的物种多样性指数、均匀度和Simpson指数都比较高。并通过对物种多样性指数的分析,探讨了如何才能维持较高的生物多样性。     

Growth strategy of trees related to successional status II. Leaf dynamics

The leaf dynamics of two early (Alnus nepalensis and Schima khasiana) and three late (Machilus kingii, Quercus dealbata and Q. griffithii) successional tree species of a subtropical humid, evergreen, montane forest at Upper Shillong (1900 m) in north-eastern India has been studied using demographic techniques. The open-grown early successional species show greater leaf production and higher turnover rate than the open-grown late successional species. The latter, under forest situations, have still slower turnover rates but with leaves of several age groups. It is suggested that the early successional species are attuned to environments of temporary resource availability while the late successional species are adapted to the relatively predictable but highly competitive environments of closed forests.     

Density dependence on tree survival in an old-growth temperate forest in northeastern China

? Density dependence is a major mechanism for shaping plant communities. However, its role in regulating diverse, mixed natural tree communities is less certain.

? In this study we investigated density-dependent effects in a large-scale (25 ha) old-growth temperate forest in northeastern China. Spatial patterns of neighborhood distribution in the plot were analyzed using various methods for inferring competition, including (1) pair correlation function to determine spatial patterns of pre-mortality and post-mortality and (2) neighborhood analysis of individuals to examine the extent to which tree survival is correlated with other covariates.

? Results showed that, for common species, 3 of 5 canopy species and 3 of 8 midstory and understory species were random in mortality. Negative density-dependent mortality was not found when trees reach 1 cm in DBH. There was no significant correlation for canopy species between tree survival and conspecific abundance, but largely positive correlations for midstory and understory species. In contrast, tree survival was found to negatively correlate with conspecific basal area for most species, indicating strong intraspecific competition. No strong interspecific density dependence was found in the forest.

     

栎属树种生长模型研究进展

栎属树种是亚热带常绿阔叶林和温带落叶阔叶林的主要建群树种之一,分布范围极为广泛,在国内外被广泛应用于城市园林绿化、水源涵养林、水土保持林,也是重要的珍贵用材树种,同时其果实、栓皮等具有重要的工业和药用价值.研究栎林的生长过程,对其经营抚育决策具有重要的参考意义.文中对近年来国内外关于栎属树种的生长过程,特别是栎属树种生长模型的研究进行了综述,包括栎属树种全林分生长模型、单木生长模型、径阶分布模型等;阐述了栎属树种生长模型的研究现状及发展趋势,以期为栎林的经营抚育决策提供参考.     

Evaluation of leaf display of evergreen broadleaved tree species and deciduous tree species in warm temperate conifer plantations

We investigated the sapling leaf display in the shade among trees of various leaf lifespans co-occurring under the canopy of a warm-temperate conifer plantation. We measured leaf-area ratio (aLAR) and morphological traits of saplings of evergreen broadleaved tree species and a deciduous tree species. Although we found large interspecific and intraspecific differences in aLAR even among saplings of similar size in the homogeneous light environment, we did not find a consistent trend in aLAR with leaf lifespan among the species. While deciduous trees annually produced a large leaf area, some evergreen broadleaved trees retained their leaves across years and had aLAR values as high as those of deciduous trees. Among leaf-level, shoot-level, and individual-level morphological traits, aLAR was positively correlated with current-year shoots mass per aboveground biomass in deciduous trees, and with the area of old leaves per aboveground mass in evergreen broadleaved trees. Thus, tree-to-tree variation in the degrees of annual shoot production and the accumulation of old leaves were responsible for the interspecific and intraspecific variations in aLAR.     

Spatial Pattern of Bird-Dispersed Seed Rain ofDaphniphyllum macropodum in an Evergreen Broad-Leaved Forest

Abstract

We investigated how the locations of conspecific fruiting trees, other fruiting species, and canopy gaps affected spatial patterns of bird-dispersed seed rain for a common ornithochorous tree: Daphniphyllum macropodum.In a mature evergreen broad-leaved forest in southwestern Japan, seed rain was monitored with seed traps in a 1.2 ha plot on a site with a generally closed canopy. Many of the bird-dispersed seeds of D. macropodumwere concentrated near the conspecific fruiting trees, while other seeds were scatter-dispersed. The spatial distribution of scatter-dispersed seeds was independent of the distance from conspecific fruiting trees. The scatter-dispersed seed rain of D. macropodumwas biased by abundant and widely distributed fruiting trees of Cleyerajaponica.Other rare fruiting species and canopy gaps had no clear effect on the seed rain of D. macropodum.Results show that the spatial pattern of bird-dispersed seed rain in this forest is affected by the location of fruiting trees of the same and different ornithochorous species rather than by canopy gaps.     

中国不同地带性森林乔木叶片热值特征及其影响因素

[Objective]To study the leaf calorific value of different forest types.[Method]This study focused on four common forest types widely distributed across China, including boreal coniferous forest, warm temperate deciduous broad-leaved forest, subtropical evergreen broad-leaved forest, and tropical monsoon forest. The leaf calorific values of 175 dominant (or common) tree species were measured. The leaf calorific value characteristics of the trees found in the different forest types were analyzed, and the primary factors affecting leaf calorific value were investigated, including leaf morphological traits, nutrient elements, climate, and soil traits. [Result]The results showed that in these forest types, the leaf calorific values ranged from 14.84 to 21.98 KJ·g^-1, with an overall mean of 19.06 KJ·g^-1. The presence of organisms appeared to affect the leaf calorific value, which differed among forest types as follows: coniferous trees > broadleaf trees or evergreen trees > deciduous trees. [Conclusion]The latitudinal pattern of tree leaf calorific value, ordered from north to south, is as follows: warm temperate deciduous broad-leaved forest > subtropical evergreen broad-leaved forest > tropical mountain rainforest. The mean leaf calorific value of boreal coniferous forest was slightly lower than those of warm temperate deciduous broad-leaved forest and subtropical evergreen broad-leaved forest. The leaf calorific values were significantly correlated with leaf carbon content (R²= 0.89, P<0.001). A multiple regression equation was established to describe the relationships among leaf calorific value, leaf carbon content, leaf nitrogen content, and leaf thickness.     

长三角城市乡土树种的天然林木群落分析与应用

长三角城市群园林绿化程度很高,但一些城市仍缺少乡土树种特色,危及生态系统稳定。长三角地跨北、中两个亚热带,有三类林木群落组成,即含有常绿阔叶树种的落叶阔叶林、落叶常绿阔叶混交林、常绿阔叶林。长三角范围乡土树种丰富,城市绿化以乡土树种为主,应大力推广应用。     

川西山地岷江冷杉风景林林分空间结构分析

应用传统林分结构因子配合混交度、角尺度、大小比数和开敞度4个林分空间结构参数,分析了川西山地退化风景林的空间结构特征.结果表明:林分结构较为简单,林分乔木层由7个树种组成,树种密度分布差异较大,树种结构相对单一;林分垂直结构特征明显,可分为3个林层,呈现出典型的复层林结构特征;林分平均混交度为0.46,以中强度混交单元为主,所占比例达61％;平均角尺度为0.483,属于随机分布状态;林木的胸径大小比数为0.286,有接近30％的林木处于优势状态;林分平均敞开度为0.309,林木生长空间基本充足.岷江冷杉种群优势度明显,群落处于相对稳定状态,其他5个树种由于株数较少,且径级分布不连续,自然更新较困难,经过一定时间的演化,将死亡,因此必须切实采取保护措施,保证其顺利生长,以保持物种多样性和群落的稳定性.     

Successional process of Picea crassifolia forest after logging disturbance in semiarid mountains: A case study in the Qilian Mountains,northwestern China

Selective logging is the most widely employed method of commercial timber production in Asia, and its impact on forest structure, composition, and regeneration dynamics is considerable. However, the successional processes in forest communities after logging in semiarid mountains are poorly understood. To provide more information on these processes, we used data from tree rings, direct and indirect age determinations, and field measurements of stand structure to reconstruct the historical disturbance regime, stand development patterns, and successional processes in a natural Picea crassifolia forest community in the Qilian Mountains of northwestern China. The results showed that the density of P. crassifolia forest increased significantly after logging. The densities of second growth forests 30 and 70 years after logging disturbance had increased to 2874% and 294% of primary forest's density, respectively. Logging disturbance did not alter tree species composition of logged stands. However, the diversity of understory species changed significantly among the successional phases. Logging disturbance decreased the spatial heterogeneity of second growth forest. The spatial distributions of recruitment were affected by the location of the remaining trees. There was less recruitment near the remaining trees than near forest that had been cut. In addition, logging disturbance also induced a growth release for the trees on the sites sampled. Our results imply that the succession and regeneration of P. crassifolia forest may be improved if the remaining trees could be retained relative uniform distribution pattern, thinning or selective logging could be performed to height density, exotic shrubs could be removed or the shrubs cover could be reduced during the earlier successional stages.     

亚热带次生常绿阔叶林主要乔木各自然恢复１５年来的变化

在瓦屋山定位研究了中亚热带湿性常绍阔叶林次生群落１５a来各乔木种且成及其重要值、密度，大小结构、高度结构和材积生产力的动态变化，结果表明，自然恢复过程中次生常绿阔叶林群落乔木层物种组成变化明显，有１１种从群落乔木层消失，也有１０种出现，乔木层树种多样性（丰富度）基本保持动态平衡和发展；优势种地位保持不变，但次优势种地位得以加强；群落结构变化较大，突出表现为种群密度、胸径和树高分布格局的变化和立体空间的分布，群落由单优或双优种向双优或多优势种发展，由单一乔木屋向多层次立体结构发展，空间加厚，群落更复杂、稳定性增强。乔木种群平均高生长速度在２４－４８cm.a^-1 ,平均ＤＢＨ生长率在０.３４－１.１０cm.a^-1，各种群生长速度差异明显，按生长速度可划分为适应性强的速生树种如细梗呈茱萸叶五加，华木荷（平均ＤＢＨ生长在０．７０cm.a^-1以上），中等速度的小叶青冈、扁制栲（平均ＤＨＢ生长在０．４０cm.a^-1以上），慢生种如润楠、石栎（平均ＤＨＢ生长在０．４cm.a^-1以下）；种群材积生产力高低不一，优势种扁刺种群材积生产力并不是直增长，而是在后期出现下降；次优势和睦产长速度较快。１９９７年３固定样地材积量２１８－２２５m^3.hm^-2，优势种群材积生产力是群落材积生产力的高低的关键，在相同的环境条件下和各种群相互作用下，密度是各种群生长和材积生产力高低的决定因素之一。     

Sprouting characteristics of a subtropical evergreen broad-leaved forest following clear-cutting in Okinawa,Japan

This study explores the sprouting characteristics of an evergreen broad-leaved forest after clear-cutting based on a survey of 1,893 stumps of 62 tree species in Okinawa, Japan. The sprouting capabilities of the stumps varied among tree species. The stumps of 60 species could produce sprouts, while those of the other two species could not. In 10 of the sprouting species, the mean sprout-stem number was higher than 4.0 stems per stump; nine sprouting species showed low sprouting capabilities. Additionally, Castanopsis sieboldii and Schima wallichii, the dominant species, exhibited high sprouting capabilities. Further, the sprouting capabilities varied with stump diameter at breast height (DBH). Stumps with a larger DBH tended to have a higher mean number of sprout stems per stump, higher DBH, and higher tree height than the smaller stumps. The forest stands regenerated by sprouting might recover into a tree community similar to that before clear-cutting.     

Biomass, diversity and composition of epiphytic macrolichens in primary and secondary forests in the subtropical Ailao Mountains, SW China

Epiphytic macrolichen litterfall was collected over 3 years from primary evergreen broad-leaved forests (PF), Populus bonatii secondary forests (PBSF), middle-aged oak secondary forests (MOSF) and old-aged oak secondary forests (OOSF) of the Ailao Mountains in SW China. To assess changes in the epiphyte communities of the subtropical forests, we compared the differences in biomass, species diversity and community structure of epiphytic lichens from the four forest types. A total of 51 species were recorded in this study. Species richness was highest in the PF, while α-diversity was highest in the MOSF. Lichen biomass differed markedly across the four forest types and was highest in the MOSF. The contribution of each dominant species to total biomass, except Nephromopsis ornata, was significantly different among forest types. Moreover, the percent contribution of foliose chlorolichens to litterfall tended to be higher in later-succession forest types, whereas the contributions of cyanolichens and fruticose chlorolichens were lower in these forest types. Compared to the PF, the lichen community structure in secondary forests differed significantly, implying that at least a few hundred years were needed for the restoration of these lichen communities. In particular, nineteen lichens were suggested as indicators, and eleven of them were present in the MOSF. Canonical correspondence analysis (CCA) indicated that the observed differences were mainly attributed to canopy openness and the size of the largest tree, which represented an environmental gradient from exposed to sheltered habitats. The diversity of host tree species was also important in determining the composition and distribution of macrolichens. Our findings support the idea that the maintenance of the forest landscape mosaic of heterogeneous forest types may be an important management practice to maintain or promote the epiphytic lichen community of the subtropical Ailao Mountains.     

19个地区种子植物区系数量分类的初步研究

运用最小距离聚类法及主成分分析法对全国 19个地区种子植物区系的种类组成和地理成分进行分析 ,结果表明 ,19个地区可划分为 5类 :第 1类是以泛热带植物区成分为主的亚热带常绿阔叶林区系类型 ,包括闽、赣、浙、皖、湘、鄂 6省的15个地区 ;第 2类是山东泰山 ,是以温带落叶阔叶林为主的植物区系 ;广西大瑶山属于第 3类 ,是古热带植物区成分为主的亚热带常绿阔叶林区系类型 ;元谋干热河谷属于第 4类 ,是热带稀树草原—亚热带常绿阔叶林组合的特殊类型 ;长白山归于为第 5类 ,是典型的温带性植物区系类型     

哀牢山湿性常绿阔叶林木质藤本植物的物种多样性及其与支柱木的关系

分别在哀牢山湿性常绿阔叶林沟谷和坡面调查了10个20m×50m的样地,研究哀牢山湿性常绿阔叶林木质藤本植物的物种多样性及其与支柱木的关系。结果表明:研究区共记录到DBH≥1cm的木质藤本植物402株(隶属于23种21属16科)和DBH≥10cm的林木1522株(隶属于47种30属15科);与其他亚热带森林比较,该森林中木质藤本植物物种较为丰富但多度较低;藤本植物的物种丰富度、密度和基面积在沟谷显著高于坡面,而林木的差异性不显著;木质藤本植物在支柱木上呈集群分布,并且不同种支柱木被藤本植物攀援的百分比间存在显著的差异(P0.001),说明藤本植物的攀援对支柱木具有选择性;云南越桔、薄叶马银花和景东冬青等树种因其树皮光滑而不易被藤本植物攀援,而腾冲栲、七裂槭、山矾和多花山矾等则易于被藤本植物攀援;大径级支柱木被藤本植物攀援的比率高于小径级支柱木;茎缠绕和钩刺攀援藤本的胸径与支柱木胸径极显著相关(P0.001),根攀援和卷须攀援藤本的胸径与支柱木胸径相关性不显著(P0.05)。     

The sustainability of selection cutting in a late successional broadleaved community forest in Bhutan

This study assessed the sustainability of selection cutting in a community forest (CF) in Bhutan. The harvesting approach differed from cable crane logging operations in an adjacent commercially managed forest by creating much smaller canopy openings. This had many implications for natural regeneration of preferred species. The study was conducted in a late successional broadleaved CF containing 32 genera of tree species dominated by Quercus and Castanopsis and managed for timber, firewood, non-wood forest products and forest grazing. The study was based on a comparison of two forest inventories to assess forest structure and regeneration, a study of stumps to quantify harvesting intensities, and a household survey to quantify livestock holdings and grazing patterns. The study examined different intensities of selection cutting in three blocks of the CF and found that higher intensities of selection cutting did not have a negative impact on: (a) natural regeneration of seedlings and saplings of preferred timber species; (b) the diameter distribution of all species and use categories except for Quercus; (c) the diversity of tree genera; and (d) the percentage of remaining trees with favorable bole shape and form. These results contrasted with findings in the adjacent commercially managed forest. Community management of broadleaved forests with selection cutting appeared to be sustainable and avoid some of the unresolved silvicultural problems associated with commercially managed forests in Bhutan. However more research is needed on the sustainable management of Quercus spp.