Modelling of planted legume fallows in Western Kenya using WaNuLCAS. (I) Model calibration and validation

Poor soil fertility is the biggest obstacle to agricultural productivity in Sub-Saharan Africa. Improved fallows can help to raise agricultural productivity in these systems of low financial capital, however, experimental testing of their potential application domain and design is costly and time consuming. Models can evaluate alternative systems relatively quickly and at relatively low cost, but must first be validated to assess satisfactory simulation of the target systems. Specific climatic, edaphic, crop and fallow growth data was used from five sites in Western Kenya to calibrate and validate simulations of maize and improved fallow growth using the Water, Nutrient and Light Capture in Agroforestry Systems (WaNuLCAS) model. The model predicted continuous maize yields across the sites with an R ² of 0.72, an EF (model efficiency) of 0.66 and a CD (coefficient of determination) of 2.73, although the default pedotransfer functions (PTF) for volumetric soil water content used in the model had to be substituted for a tropical soils specific PTF before this was achieved. Predicted maize yield was consistently related to fallow biomass (i.e. higher fallow biomass correlated with higher subsequent maize yields) at two sites and the model predicted maize yields following fallow growth from this subset of the data with an R ² of 0.42. This relationship of fallow biomass to subsequent maize yield was not observed across the whole data set due to incomplete fallow litterfall data, factors not included in the model and associated poor model prediction of recycled tree biomass. After site and tree calibration, the model can thus be applied to assess fallow management strategies for sites limited by water and nitrogen.     

Short fallows of Tithonia diversifolia and Crotalaria grahamiana for soil fertility improvement in western Kenya

Managed short-duration fallows may have the potential to replace longer fallows in regions where population density no longer permits slow natural fallow successions. The purpose of fallows is not only to improve subsequent crop performance but also to restore soil fertility and organic matter content for the long term. We therefore evaluated the soil organic matter and nutrient flows and fractions in a short fallow experiment managed in the western Kenya highlands, and also compared the experimental area with a 9–12-yr-oldadjacent natural bush fallow. The factorial agroforestry field experiment with four land-use and two P fertilizer treatments on a Kandiudalfic Eutrudox showed that 31-wk managed fallows with Tithonia diversifolia(Hemsley) A. Gray and Crotalaria grahamiana Wight &Arn. improved soil fertility and organic matter content above those of a natural weed fallow and continuous maize (Zea mays L.). Post-fallow maize yields were also improved, although cumulative three-season increases in yield were small (0–1.2 Mg ha⁻¹) when the yield foregone during the fallow season was accounted for. Improvements in yield and soil quality could be traced to quantity or quality of biomass recycled by the managed fallows. The non-woody recycled biomass produced by the continuous maize, weed fallow, and tithonia treatments was near 2Mg ha⁻¹, whereas crotalaria produced three times more recyclable biomass and associated N and P. Increases in topsoil N due to the fallows may have been attributable in part to deep acquisition and recycling of N by the fallows. Particulate macro-organic matter produced by the fallows contained sufficient N(30–50 kg ha⁻¹) to contribute substantially to maize production. Organic Paccumulation (29 kg ha⁻¹) similarly may play a significant role in crop nutrition upon subsequent mineralization. The effect of the P fertilizer application on soil properties and maize yield was constant for all land-use systems (i.e., no land-use system × P fertilizer interactions occurred). There was an indication that tithonia may have stimulated infestation of Striga hermonthica (Del.) Benth., and care must be taken to evaluate the full effects of managed fallows over several seasons. This revised version was published online in June 2006 with corrections to the Cover Date.     

Differences in abundance,species richness,and body size of ground beetles (Coleoptera: Carabidae) between beech (Fagus sylvatica L.) forests on Podzol and Cambisol

Ten forest sites with a tree layer of almost pure beech in southern-central Scania, south Sweden, were studied. They were all located in the interior of large forest stands. Five of the stands were developed on mor Podzols with scanty or almost no ground layer vegetation. The other five sites were located on mull Cambisols with a rich ground layer of many vascular plant species and a shrub layer of varying density. In total, 42 carabid beetle species and 9260 individuals were captured with pitfall traps (15 per site, diameter 87 mm) during 12 weeks in April, July and September 2007. Composition of the carabid fauna differed considerably between the two types of sites. In the Cambisol sites 39 carabid species were found, compared to only 21 species in the Cambisol sites. Mean carabid species richness per site differed significantly between Cambisol and Podzol sites (n = 20.4 and 13.2, respectively). The difference was wholly due to a lower number of small carabid species (mean body length <12 mm) in the Podzol sites. Of four variables (pH-KCl, Clay, Organic matter, and Sum of plant cover percentages below tree canopy) the sum of plant cover variable usually accounted for more (>50%) of the carabid beetle variability compared to the three other variables. Carabus glabratus and Pterostichus oblongopunctatus were almost exclusively found on Podzol. Carabus coriaceus, Pterostichus melanarius and in particular Carabus nemoralis were more abundant on Cambisol, whereas Carabus hortensis and Pterostichus niger were almost ubiquitous and attained the highest total numbers. There was no significant difference between Podzol and Cambisol sites in the total number of carabid individuals captured. C. coriaceus, C. hortensis, and Carabus violaceus, species that were abundant enough in all sites for a comparison, had significantly larger mean body sizes in the Podzol than in the Cambisol sites, a finding valid for both sexes. There were more males than females of C. coriaceus, C. violaceus, and C. nemoralis in the catches, whereas sex distribution of C. hortensis was equal. Possible explanations for the findings are differences in microclimate and food web characteristics related to differences between Podzols and Cambisols in understorey vegetation and soil or litter properties.     

Spatial distribution of soil carbon and nitrogen storage and forest productivity in a watershed planted to Japanese cedar (Cryptomeria japonica D. Don)

Digital terrain modeling was used to evaluate landscape-level spatial variation of soil C and N storage and site productivity in Japanese cedar (Cryptomeria japonica D. Don) stands. Soil C and N storage were measured in samples from surface soils (0–25 cm depth) of 29 Japanese cedar stands in the 205-ha Myougodani watershed, Toyama Prefecture. The site index (C. japonica tree height at age 40 years) was used as a measure of forest productivity. Seven terrain attributes (elevation, slope gradient, aspect, profile curvature, plan curvature, openness, and wetness index) were calculated from a digital elevation model. Soil C and N storage were negatively correlated with slope gradient and positively correlated with openness. Variation in the site index was closely related to the wetness index. The prediction models using terrain attributes as explanatory variables explained 50% of the variability in soil C storage, 53% of the variability in soil N storage, and 75% of the variability in site index. This result demonstrated that this technique is useful for estimating the spatial distribution of soil properties and productivity in forest landscapes. On the other hand, there was no correlation between site index and soil C and N storage. Use of the prediction models in a geographic information system revealed that the spatial distribution of forest productivity differed considerably from those of soil C and N storage.     

Productivity of alley farming with leucaena (Leucaena leucocephala Lam. de Wit) and Napier grass (Pennisetum purpureum K. Schum) in coastal lowland Kenya

Inadequate supply of fodder is a serious constraint to the potentially-promising small-holder-dairy production system in coastal Kenya. Alley farming could be an approach to addressing this problem. A study of forage production based on Napier grass and leucaena in an alley cropping system was conducted on an infertile sandy soil in lowland coastal Kenya. The effects of leucaena hedgerows,Clitoria ternatea (L.) intercropping, addition of slurry (110 t ha^–1 yr^–1) and two harvesting managements (severe and lenient) on the yield of Napier grass fodder, were assessed. The study was initiated in 1989 and three years results are reported.Napier grass fodder yields did not decline in the presence of leucaena heggerows in most harvests. Generally, the Napier rows adjacent to the hedgerows gave greater yields than in either the centre or sole Napier rows. Interplanting clitoria between Napier rows had a beneficial effect on Napier yields in the later harvests. Compared with sole Napier, the hedgerow treatment receiving slurry increased yield ha^–1 of Napier by 50%, of total forage by 80%, and of nitrogen (measured over two harvests) by 200%.During the first two years of cropping most treatments did not affect the OM and nitrogen content of the soil but by the end of the third cropping year, slurry application increased soil OM by 21% and slurry and clitoria treatments each increased soil nitrogen by 33%.The study demonstrated the increases in the supply and quality of forages for dairy cattle that can be achieved by planting complementary forages, legume and grass species in an intercropping system like alley farming and the benefits to soil composition by the return of slurry.     

Gas exchange responses of <Emphasis Type="Italic">Eucalyptus</Emphasis>, <Emphasis Type="Italic">C. africana</Emphasis> and <Emphasis Type="Italic">G. robusta</Emphasis> to varying soil moisture content in semi-arid (Thika) Kenya

A dramatic decline in forest cover in eastern Africa along with a growing population means that timber and poles for building and fuelwood are in short supply. To overcome this shortage, the region is increasingly turning to eucalyptus. But eucalyptus raises environmental concerns of its own. Fears that it will deplete water supply, affect wildlife and reduce associated crop yields have caused many countries in the region to discourage farmers from planting this exotic. This paper is part of a series of investigations on the growth and water use efficiency of faster growing eucalyptus hybrids, which was introduced from South Africa to Kenya. The hypothesis is that the new hybrids are more efficient in using water and more suitable for the semi-arid tropics than existing eucalyptus and two popular agroforestry species. Gas exchange characteristics of juvenile Eucalyptus grandis (W. Hill ex Maiden), two eucalyptus hybrids (E. grandis × Eucalyptus camaldulensis Dehnh.), Grevillea robusta (A. Cunn) and Cordia africana (Lam) was studied under field and pot conditions using an infrared gas analyzer was used to measure photosynthetic active radiation (PAR), net photosynthetic rate (A), stomatal conductance (g _s) and transpiration rate (E) at CO₂ concentrations of 360 μmol mol⁻¹ and ambient humidity and temperature. A, E and g _s varied between species, being highest in eucalyptus hybrid GC 15 (24.6 μmol m⁻² s⁻¹) compared to eucalyptus hybrid GC 584 (21.0 μmol m⁻² s⁻¹), E. grandis (19.2 μmol m⁻² s⁻¹), C. africana (17.7 μmol m⁻² s⁻¹) and G. robusta (11.1 μmol m⁻² s⁻¹). C. africana exhibited high E values (7.0 mmol m⁻² s⁻¹) at optimal soil moisture contents than G. robusta (3.9 mmol m⁻² s⁻¹) and eucalyptus (5.3 mmol m⁻² s⁻¹) in field experiment and G. robusta (3.2 mmol m⁻² s⁻¹) and eucalyptus (4.2 mmol m⁻² s⁻¹) in pot-grown trees. At very low soil moisture content, extremely small g _s values were recorded in GC 15 and E. grandis (8 mmol m⁻² s⁻¹) and G. robusta (14 mmol m⁻² s⁻¹) compared to GC 584 (46.9 mmol m⁻² s⁻¹) and C. africana (90.0 mmol m⁻² s⁻¹) indicating strong stomatal control by the species. Instantaneous water use efficiency ranged between 3 and 5 μmol mmol⁻¹ and generally decreased with decline in soil moisture in pot-grown trees but increased with declining soil moisture in field-grown trees.     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica II. Cacao and wood production,litter production and decomposition

During 7 years (1979–1985) cacao harvests (beans and husks) have been recorded for the agroforestry systems ofTheobroma cacao underCordia alliodora andErythrina poeppigiana shade trees. The mean oven dry cacao yields were 626 and 712 kg.ha⁻¹.a⁻¹ cocoa beans underC. alliodora andE. poeppigiana respectively. Harvests have gradually increased over the years and the plantation has now reached maturity. Annual extraction of N, P, K, Ca and Mg in fruits, which is relatively small, was calculated on the basis of chemical analyses. The following average values were found (kg.ha⁻¹.a⁻¹): At the age of 8 years, theC. alliodora trees have reached 26.7 cm diameter (DBH) and 14.0 m in height. Mean annual growth (from age 5 to 7) is 14.6 m³.ha⁻¹.a⁻¹. Natural plant residue production has been measured for 4 years (Nov. 1981–Oct. 1985). UnderE. poeppigiana it has reached a value of 8.91 t.ha⁻¹.a⁻¹ and underC. alliodora 7.07 t.ha⁻¹.a⁻¹. The shade trees have contributed 57 and 47% respectively. Transference and decomposition rates are high and important in the nutrient cycles. The nutrient content of the litter was analysed and corresponding average yearly transfers were (kg.ha⁻¹.a⁻¹): For part I see Vol. 4, No. 3, 1986. Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesselschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica II. Cacao and wood production,litter production and decomposition

During 7 years (1979–1985) cacao harvests (beans and husks) have been recorded for the agroforestry systems ofTheobroma cacao underCordia alliodora andErythrina poeppigiana shade trees. The mean oven dry cacao yields were 626 and 712 kg.ha^–1.a^–1 cocoa beans underC. alliodora andE. poeppigiana respectively. Harvests have gradually increased over the years and the plantation has now reached maturity.Annual extraction of N, P, K, Ca and Mg in fruits, which is relatively small, was calculated on the basis of chemical analyses. The following average values were found (kg.ha^–1.a^–1): At the age of 8 years, theC. alliodora trees have reached 26.7 cm diameter (DBH) and 14.0 m in height. Mean annual growth (from age 5 to 7) is 14.6 m³.ha^–1.a^–1.Natural plant residue production has been measured for 4 years (Nov. 1981–Oct. 1985). UnderE. poeppigiana it has reached a value of 8.91 t.ha^–1.a^–1 and underC. alliodora 7.07 t.ha^–1.a^–1. The shade trees have contributed 57 and 47% respectively. Transference and decomposition rates are high and important in the nutrient cycles.The nutrient content of the litter was analysed and corresponding average yearly transfers were (kg.ha^–1.a^–1): For part I see Vol. 4, No. 3, 1986.Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesselschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica III. Cycles of organic matter and nutrients

Models for cycles for organic matter and nutrients element (N, P, K, Ca and Mg) are presented for the agroforestry systems of cacao (Theobroma cacao) withCordia alliodora orErythrina poeppigiana in Turrialba, Costa Rica. For the models, system reserves (soil, humus, vegetation divided into leaves, branches, stems, fine roots, fruits) and transference between compartments (production and decomposition of litter residues) inputs (fertilizer, rainfall) and outputs (harvests) of the system are considered. The implications of the models are discussed in detail. Aspects of net primary production in the systems studied are considered. N fixation is calculated on the basis of balances. Analysis of soil water showed high variations that coincided with rainfall patterns and pruning of theE. poeppigiana. For part I see Vol. 4, No. 3, 1986 For part II see this issue Agroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesellschaft für Technische Zusammenarbeit), Turrialba, Costa Rica     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) and poro (Erythrina poeppigiana) in Costa Rica III. Cycles of organic matter and nutrients

Models for cycles for organic matter and nutrients element (N, P, K, Ca and Mg) are presented for the agroforestry systems of cacao (Theobroma cacao) withCordia alliodora orErythrina poeppigiana in Turrialba, Costa Rica.For the models, system reserves (soil, humus, vegetation divided into leaves, branches, stems, fine roots, fruits) and transference between compartments (production and decomposition of litter residues) inputs (fertilizer, rainfall) and outputs (harvests) of the system are considered.The implications of the models are discussed in detail.Aspects of net primary production in the systems studied are considered.N fixation is calculated on the basis of balances. Analysis of soil water showed high variations that coincided with rainfall patterns and pruning of theE. poeppigiana.For part I see Vol. 4, No. 3, 1986 For part II see this issueAgroforestry Project, CATIE/GTZ (Tropical Agricultural Research and Training Center/Gesellschaft für Technische Zusammenarbeit), Turrialba, Costa Rica