Bottom-up control of a northern Arizona ponderosa pine forest fire regime in a fragmented landscape

Fire regimes often vary at fine spatial scales in response to factors such as topography or fuels while climate usually synchronizes fires across broader scales. We investigated the relative influence of top-down and bottom-up controls on fire occurrence in ponderosa pine (Pinus ponderosa) forests in a highly fragmented landscape at Mount Dellenbaugh, in northwestern Arizona. Our study area of 4,000?ha was characterized by patches of ponderosa pine forest in drainages that were separated by a matrix of pinyon?Cjuniper woodlands, sagebrush shrublands, and perennial grasslands. We reconstructed fire histories from 135 fire-scarred trees in sixteen 25-ha sample sites placed in patches of mature ponderosa forest. We found that, among patches of ponderosa forest, fires were similar in terms of frequency but highly asynchronous in terms of individual years. Climate synchronized fire but only across broader spatial scales. Fires occurring at broader scales were associated with dry years that were preceded by several wet years. The remarkable level of asynchrony at finer scales suggests that bottom-up factors, such as site productivity and fuel continuity, were important in regulating fire at Mount Dellenbaugh. Understanding where bottom-up controls were historically influential is important for prioritizing areas that may best respond to fuel treatment under a warming climate.     

Long-term forest dynamic after land abandonment in a fire prone Mediterranean landscape (central Corsica,France)

Two hundred years of landscape changes were studied on a 3,760 ha area of central Corsica (France) representing a typical Mediterranean environment. Different historical sources, including an accurate land-cover map from 1774 and statistics on land cover from 1848 and 1913, were used. Three additional maps (1960, 1975 and 1990) were drawn, and a complete fire history from 1957 to 1997 was created. Forests expanded slowly by a border effect. Forest expansion was more rapid in unburnt sites (0.59% per year) than in burnt sites (0.23% per year), mostly because the initial amount of forests was greater. Because of the border effect, the combination of past landscape pattern and short distance colonization abilities of forest species may have allowed the shrublands to persist in some places after land abandonment. This persistence may explain the pattern of fire in the landscape, since shrubland burn more readily than forests.     

Arizona pine (Pinus arizonica) stand dynamics: local and regional factors in a fire-prone madrean gallery forest of Southeast Arizona,USA

In southwestern North America, large-scale climate patterns appear to exert control on moisture availability, fire occurrence, and tree demography, raising the compelling possibility of regional synchronization of forest dynamics. Such regional signals may be obscured, however, by local, site-specific factors, such as disturbance history and land use. Contiguous sites with similar physical environments, lower and middle Rhyolite Canyon, Arizona, USA, shared nearly the same fire history from 1660-1801, but then diverged. For the next 50 years, fires continued to occur frequently in lower Rhyolite, but, probably as result of flood-induced debris deposition, largely ceased in middle Rhyolite. We related stand dynamics of Arizona pine (Pinus arizonica) to fire history and drought severity and compared the dynamics in the two sites before and after the divergence in fire frequency. Fires occurred during unusually dry years, and possibly following unusually moist years. Arizona pine exhibited three age structure peaks: two (1810–1830 and 1870–1900) shared by the two sites and one (1610–1640) only in middle Rhyolite. The latter two peaks occurred during periods of unusually low fire frequency, suggesting that fire-induced mortality shapes age structure. Evidence was mixed for the role of favorable moisture availability in age structure. As expected, moisture availability had a prominent positive effect on radial growth, but the effect of fire was largely neutral. The two sites differed only moderately in stand dynamics during the period of divergence, exhibiting subtle age structure contrasts and, in middle Rhyolite only, reduced growth during a 50-year fire hiatus followed by fire-induced release. These results suggest that, despite local differences in disturbance history, forest responses to regional fire and climate processes can persist.     

Effects of site,landscape features,and fire regime on vegetation patterns in presettlement southern Wisconsin

The presettlement tree cover (1831–33) of 3 townships in a southern Wisconsin landscape was analyzed using original survey records. Four forest types were identified: closed forest, open forest, savanna, and prairie. Comparisons of vegetation types and landscape pattern were made between the east and west sides of the Pecatonica River, which bisects the landscape and could have acted as a natural fire barrier. West of the river, presettlement tree species richness and diversity were lower and trees were smaller in diameter and less dense than to the east. The major vegetation types to the west were prairie (42% of landscape) and savanna (40%), both fire-susceptible types. Prairie was more common on gentle slopes than on other landforms. To the east, the landscape was 70% forested (closed plus open forest). Here, prairie was more frequent on steep dry sites. These vegetation differences, including the contrasting landscape placement of prairie, are attributed to distinct site characteristics and to disturbance (fire) regimes, with the west likely having more frequent fires. In terms of the four vegetation types, the east landscape was more homogeneous, being dominated by closed forest (50%). West of the Pecatonica River, the landscape was more heterogeneous because of the high proportion of both prairie and savanna; however, in terms of flammability of vegetation, the west was essentially homogeneous (82% prairie plus savanna).     

Spatial patterns of nineteenth century fire severity persist after fire exclusion and a twenty-first century wildfire in a mixed conifer forest landscape,Southern Cascades,USA

Landscape Ecology - Spatial patterns of fire severity are influenced by fire-vegetation patch dynamics and topography. Since the late nineteenth century, fire exclusion has increased fuels and...     

Effects of landscape patterns of fire severity on regenerating ponderosa pine forests (<Emphasis Type="Italic">Pinus ponderosa</Emphasis>) in New Mexico and Arizona,USA

Much of the current effort to restore southwestern ponderosa pine forests to historical conditions is predicated upon assumptions regarding the catastrophic effects of large fires that are now defining a new fire regime. To determine how spatial characteristics influence the process of ponderosa pine regeneration under this new regime, we mapped the spatial patterns of severity at areas that burned in 1960 (Saddle Mountain, AZ) and (La Mesa, NM) 1977 using pre- and post-fire aerial photography, and quantified characteristics of pine regeneration at sample plots in areas where all trees were killed by the fire event. We used generalized linear models to determine the relationship of ponderosa pine stem density to three spatial burn pattern metrics: (1) distance to nearest edge of lower severity; (2) neighborhood severity, measured at varying spatial scales, and (3) scaled seed dispersal kernel surfaces. Pine regeneration corresponded most closely with particular scales of measurement in both seed dispersal kernel and neighborhood severity. Spatial patterns of burning remained important to understanding regeneration even after consideration of subsequent disturbance and other environmental variables, with the exception of a few cases in which simpler models were equally well-supported by the data. Analysis of tree ages revealed slow progress in early post-fire years. Our observations suggest that populations spread in a moving front, as well as by remotely dispersed individuals. Based on our results, recent large fires cannot be summarily dismissed as catastrophic. We conclude that management should focus on the value and natural recovery of post-fire landscapes. Further, process centered restoration efforts could utilize our findings in formulating reference dynamics under a changing fire regime.     

Mixed-severity fire regime in a high-elevation forest of Grand Canyon,Arizona, USA

Fire regime characteristics of high-elevation forests on the North Rim of the Grand Canyon, Arizona, were reconstructed from fire scar analysis, remote sensing, tree age, and forest structure measurements, a first attempt at detailed reconstruction of the transition from surface to stand-replacing fire patterns in the Southwest. Tree densities and fire-/non-fire-initiated groups were highly mixed over the landscape, so distinct fire-created stands could not be delineated from satellite imagery or the oldest available aerial photos. Surface fires were common from 1700 to 1879 in the 4,400 ha site, especially on S and W aspects. Fire dates frequently coincided with fire dates measured at study sites at lower elevation, suggesting that pre-1880 fire sizes may have been very large. Large fires, those scarring 25% or more of the sample trees, were relatively infrequent, averaging 31 years between burns. Four of the five major regional fire years occurred in the 1700s, followed by a 94-year gap until 1879. Fires typically occurred in significantly dry years (Palmer Drought Stress Index), with severe drought in major regional fire years. Currently the forest is predominantly spruce-fir, mixed conifer, and aspen. In contrast, dendroecological reconstruction of past forest structure showed that the forest in 1880 was very open, corresponding closely with historical (1910) accounts of severe fires leaving partially denuded landscapes. Age structure and species composition were used to classify sampling points into fire-initiated and non-fire-initiated groups. Tree groups on nearly 60% of the plots were fire-initiated; the oldest such groups appeared to have originated after severe fires in 1782 or 1785. In 1880, all fire-initiated groups were less than 100 years old and nearly 25% of the groups were less than 20 years old. Non-fire-initiated groups were significantly older (oldest 262 years in 1880), dominated by ponderosa pine, Douglas-fir, or white fir, and occurred preferentially on S and W slopes. The mixed-severity fire regime, transitioning from lower-elevation surface fires to mixed surface and stand-replacing fire at higher elevations, appeared not to have been stable over the temporal and spatial scales of this study. Information about historical fire regime and forest structure is valuable for managers but the information is probably less specific and stable for high-elevation forests than for low-elevation ponderosa pine forests.This revised version was published online in May 2005 with corrections to the Cover Date.     

Historical patterns of fire severity and forest structure and composition in a landscape structured by frequent large fires: Pumice Plateau ecoregion,Oregon, USA

Context

Lack of quantitative observations of extent, frequency, and severity of large historical fires constrains awareness of departure of contemporary conditions from those that demonstrated resistance and resilience to frequent fire and recurring drought.

Objectives

Compare historical and contemporary fire and forest conditions for a dry forest landscape with few barriers to fire spread.

Methods

Quantify differences in (1) historical (1700–1918) and contemporary (1985–2015) fire extent, fire rotation, and stand-replacing fire and (2) historical (1914–1924) and contemporary (2012) forest structure and composition. Data include 85,750-ha tree-ring reconstruction of fire frequency and extent; >?375,000-ha timber inventory following >?78,900-ha fires in 1918; and remotely-sensed maps of contemporary fire effects and forest conditions.

Results

Historically, fires?>?20,000 ha occurred every 9.5 years; fire rotation was 14.9 years; seven fires?>?40,469 ha occurred during extreme drought (PDSI <?? 4.0); and stand-replacing fire occurred primarily in lodgepole (Pinus contorta var. murrayana). In contemporary fires, only 5% of the ecoregion burned in 30 years, and stand-replacing fire occurred primarily in ponderosa (Pinus ponderosa) and mixed-conifer. Historically, density of conifers?>?15 cm dbh exceeded 120 trees/ha on?<?5% of the area compared to 95% currently.

Conclusions

Frequent, large, low-severity fires historically maintained open-canopy ponderosa and mixed-conifer forests in which large fire- and drought-tolerant trees were prevalent. Stand-replacing patches in ponderosa and mixed-conifer were rare, even in fires >?40,469 ha (minimum size of contemporary “megafires”) during extreme drought. In this frequent-fire landscape, mixed-severity fire historically influenced lodgepole and adjacent forests. Lack of large, frequent, low-severity fires degrades contemporary forest ecosystems.