Sward characteristics important for intake in six Lolium perenne varieties

An experiment was conducted to determine the genetic variation among diploid perennial ryegrass (Lolium perenne L.) varieties for sward structural characteristics considered to be important for intake by cattle. Assessments were made between June and September in 2000 and 2001. Six varieties (Abergold, Respect, Agri, Herbie, Barezane and Barnhem) were subjected to a cutting experiment where swards were cut after 3 to 4 weeks of regrowth during the growing season. The variables, measured in three 2‐week periods, were herbage mass of dry matter (DM), sward surface height (SSH), bulk density, proportion of green leaf, tiller density, tiller weight, extended tiller height, length of sheath and length of leaf blade. Significant differences among varieties were found in both years for herbage mass of DM, SSH, bulk density, proportion of green leaf, tiller density, tiller weight and length of sheath. The results show that there is significant genetic variation among diploid perennial ryegrass varieties for sward characteristics important for intake during grazing.     

The effect of defoliation practice in Western Australia on tiller development of annual ryegrass (Lolium rigidum) and Italian ryegrass (Lolium multiflorum) and its association with forage quality

The effect of defoliation on the vegetative, early reproductive and inflorescence stages of tiller development, changes in the dry‐matter yield of leaf, stem and inflorescence and the associated changes in forage quality was determined on plants of annual ryegrass (Lolium rigidum Gaud.) and Italian ryegrass (L. multiflorum Lam.). The field study comprised seventy‐two plots of 1 m × 2 m, sown with one annual ryegrass and seven Italian ryegrass cultivars with a range of heading dates from early to late; defoliation commenced 6 weeks after germination. During the vegetative stage of growth, plots were defoliated when the tillers had three fully expanded leaves (three‐leaf stage). During the early reproductive stage of growth, to simulate a cut for silage, plots were defoliated 6–7 weeks after 0·10 of the tillers displayed nodal development. The subsequent regrowth was defoliated every 3 weeks. Assessments of changes in tiller density, yield and quality were made in the growth cycle that followed three contrasting cutting treatments during the winter–spring period (from 10 July). In treatment 1, this growth cycle (following closing‐up before a subsequent conservation cut) commenced on 7 August following two defoliations each taken when the tillers were at the three‐leaf stage. In treatment 2, the growth cycle commenced on 16 October following: for early‐maturing cultivars, two cuts at the three‐leaf stage, a cut for silage and an additional regrowth cut; for medium‐maturing cultivars three cuts at the three‐leaf stage and a cut for silage; and late‐maturing cultivars, five cuts at the three‐leaf stage. In treatment 3, defoliation up to 16 October was as for treatment 2, but the growth cycle studied started on 27 November following two additional regrowth cuts for early‐ and medium‐maturing cultivars and cut for silage for the late‐maturing cultivars. Tiller development for all cultivars was classified into three stages; vegetative, early reproductive and inflorescence. In treatment 1, in vitro dry‐matter digestibility (IVDMD) and crude protein (CP) content were negatively associated with maturation of tillers. IVDMD ranged from 0·85 to 0·60 and CP ranged from 200 to less than 100 g kg^–1 dry matter (DM) during the vegetative and inflorescence stages respectively. This large reduction in forage quality was due to an increase in the proportion of stem, inflorescence and dead material, combined with a reduction in the IVDMD and CP content of the stem. A high level of forage quality was retained for longer with later‐maturing cultivars, and/or when vegetative tillers were initiated from the defoliation of early reproductive tillers (treatments 2 and 3). However, 15 weeks after the closing‐up date in treatment 1, defoliation significantly reduced the density of inflorescences with means (±pooled s.e_m.) of 1560, 1178 and 299 ± 108 tillers m^–2, and DM yield of inflorescence with means of 3·0, 0·6 and 0·1 ± 0·15 t ha^–1 for treatments 1, 2 and 3 respectively. This study supports the recommendation that annual and Italian ryegrass cultivars should be classified according to maturity date based on the onset of inflorescence emergence, and that the judicious defoliation of early reproductive tillers can be used to promote the initiation of new vegetative tillers which in turn will retain forage quality for longer.     

Influence of cultivar and cutting date on the fatty acid composition of perennial ryegrass (Lolium perenne L.)

Two experiments were conducted to evaluate the effects of cultivar and season on the fatty acid (FA) composition of the lipids of perennial ryegrass (Lolium perenne L.). Eight diploid cultivars were cut at the same target yield of approximately 2000 kg dry matter ha^?1 between mid‐June and mid‐September. Two cultivars (Barlet and Magella) were harvested during four 2‐week periods and six cultivars (AberGold, Respect, Agri, Herbie, Barezane and Barnhem; cultivars 1–6) during three periods. The concentrations of individual FA were determined by gas chromatography. Barlet had higher concentrations of linolenic acid (C18:3) than Magella, but lower concentrations of linoleic acid (C18:2). Cultivars 1–6 were more variable in their leaf blade and stem proportions than Barlet and Magella. Despite this, there was no difference between cultivars 1–6 in the FA composition of the herbage. On average 0·74 of the FA consisted of C18:3. Higher concentrations of total FA were found in mid‐summer than in early summer. This was related to a high leaf blade proportion in the herbage, indicating that the proportion of leaf and stem of the herbage probably had more effect on lipid concentrations than the season per se in this period. However, in late August and mid‐September, the total FA concentration declined whereas the leaf blade proportion increased. Therefore, in this period environmental factors appeared to have a modifying effect. As consistent differences in the concentration of C18:3 were found among cultivars Barlet and Magella throughout the season, these studies demonstrate opportunities to change the composition of ruminant products through the choice of cultivars of perennial ryegrass.     

Is there a tiller morphology ideotype for yield differences in perennial ryegrass (Lolium perenne L.)?

The objectives of this study were to measure the variability in ‘realized’ tiller morphology and identify whether there are emergent generalizations about a tiller ideotype for productivity or survival. Morphological traits during the vegetative growth stage were measured for 2 years in the field for eight perennial ryegrass cultivars: six diploid and two tetraploid cultivars under low and high nitrogen fertilizer (50 and 225 kg N ha^?1 year^?1 respectively). Traits measured were lamina width, length and area; pseudo‐stem length and diameter; an index of tiller shape; tiller dry weight; tiller density; and herbage mass. Almost all the traits differed significantly between cultivars and significant correlations were found between the 2 years. Principal component analysis identified that tiller morphology and dry‐matter yield were independent. No trait associations indicating alternative cultivar‐specific tiller morphologies were detected but cultivars differed in tiller size. The patterns of change between the 2 years indicated a possible morphological trajectory as swards age. The genetic compensatory relationship between tiller size and density had a 1:1 slope, indicative of constant yield. This implies that larger tillers would be more leafy, and higher leafiness of tetraploid over diploid cultivars was confirmed by the leaf:non‐leaf ratio and tiller shape index.     

Effect of cutting date and frequency on yield and quality of kudzu in the southern United States

Kudzu [Pueraria lobata (Willd.) Ohwi.], a vigorous, perennial warm‐season legume, grows widely throughout the south‐eastern United States, predominantly as a volunteer species. It is tolerant of drought and acidic, infertile soils and may have potential as a low‐input forage for livestock. A field experiment was undertaken to determine the effects of cutting date and frequency on yield and nutritive value of kudzu (20‐year‐old stand with no fertilizer or lime inputs) in central Georgia. The cutting treatments included an uncut control, and plots cut once (September), twice (July and September), and three times (June, July, and September) during the 1994 growing season. Dry matter (DM) production and forage quality were determined for total herbage, leaf and stem tissues from quadrat samples taken on all plots in June, July, September, and November (after a killing frost) in 1994, and in June, 1995. Total herbage and leaf DM production was highest for the three‐harvest system and lowest in the no‐cut control plots in 1994, but these results were reversed in the harvest made in 1995. Total herbage production and crude protein concentrations were similar in kudzu cut in the initial summer harvest (either June, July, or September) during the first year. Whole plant and stem in vitro dry matter digestibility (IVDMD) in these samples declined throughout the growing season, while leaf IVDMD was unchanged until the first frost. Kudzu has the potential to be a low‐input forage for livestock, particularly as supplemental drought feed, or as a protein bank for summer or autumn grazing.     

Influence of cutting regime and fertilizer application on the botanical composition,yield and nutritive value of herbage of wet grasslands in Central Europe

The changes in dry matter (DM) yield, botanical composition and nutritive value of herbage to ruminants of two wet grasslands, Arrhenatherum elatius grassland (Experiment 1) and a Molinia caerulea fen meadow (Experiment 2), in which a range of cutting and fertilizer treatments were imposed in 1999, were assessed after 4–7 years of treatment imposition. Both experiments had a split‐plot design with four replicates. In Experiment 1 the three main‐plot cutting treatments were two cuts with a delayed first cut, three cuts and four cuts during the growing season of each year. In Experiment 2 the cutting treatments were two cuts with a traditional harvest time, two cuts with a delayed first cut and three cuts. The four sub‐plot fertilizer treatments were an unfertilized control, application of a phosphorus and potassium (PK) fertilizer, application of a nitrogen (N) and PK fertilizer to the first cut only (N₁PK) and application of PK plus N applied to each of two, three or four cuts (N_c PK). Application of fertilizer influenced yield and botanical composition of herbage more than the cutting treatments while the opposite occurred for nutritive value of the herbage. Application of fertilizer increased the proportion of tall grasses in Experiment 1 and forbs in Experiment 2. The proportion of Equisetum palustre, present only in Experiment 1, was reduced from 0·33 to less than 0·01 by increased cutting frequency together with the NPK fertilizer treatments. In Experiment 1 diversity of vascular plants was negatively affected only by the four‐cuts treatment while on both wet grasslands other cutting and fertilizer application treatments had no effect. Changes in DM yield of herbage caused by the cutting and fertilizer application treatments were similar for both vegetation types with DM yield increased significantly by fertilizer application but only slightly or not reduced by increasing the cutting frequency. Nutritive value of herbage was positively correlated with cutting frequency and was most influenced at the first cut.     

EFFECTS OF DIFFERENT GRAZING ROTATIONS ON PASTURE PRODUCTION

Different grazing treatments applied to pastures in which perennial ryegrass was dominant had little influence on DM production during the main reproductive growth period in late spring and early summer. Lax and infrequent grazing at this time did, however, reduce tiller density. During the dry summer period lax infrequent grazing increased production by 20%, while in the vegetative growth period in autumn those pastures which were alternately lax- and hard-grazed outyielded those which were hard grazed by 63%. In all treatments, initial recovery growth resulted almost exclusively from the production of leaf tissue. The rate of leaf growth declined three to five weeks after grazing, depending on treatment and season, and subsequent yield increase was dominated by sheath and stem growth.     

Relationships between morphological and chemical characteristics of perennial ryegrass varieties and intake by sheep under continuous stocking management

Plots of five intermediate‐heading varieties of perennial ryegrass (Lolium perenne L.) [AberDove, Belramo and Glen (diploid); Twins (tetraploid); and AberExcel (tetraploid hybrid)] were continuously stocked with sheep to maintain a target sward surface height of 40–50 mm. Daily dry matter (DM) intake was significantly different (F‐value = 0·032) between the varieties, with the tetraploid hybrid AberExcel having the highest values for daily DM intake and intake rate during eating. Amongst the diploid varieties, intake rate tended to be higher for sheep grazing Glen. The varieties comprised a wide range in potential growth habit, from the relatively prostrate, highly tillered Glen to the more‐erect AberExcel and there were differences between them in the vertical distribution of leaves within the sward canopy. The leaves of AberExcel weighed 3·6 mg DM cm^?2 leaf area in contrast to the other varieties (4·3–5·3 mg DM cm^?2 leaf area) resulting in a high leaf area index (LAI) in relation to the green leaf mass. Intake rate was not significantly correlated with extended tiller and sheath tube lengths, partition of herbage mass, number of tillers per square metre or LAI. However, canonical variates analysis showed that there were significant differences between the varieties for the morphological and chemical factors examined. Other factors also need to be explored to explain these differences in ingestive behaviour in order to identify plant traits that are correlated with herbage intake rate. These are needed for varieties destined for grazing use, both during the breeding programme and their subsequent evaluation.     

The use of a rising plate meter to evaluate lucerne (Medicago sativa L.) height as an important agronomic trait enabling yield estimation

A rising plate meter (RPM) is an instrument used for the measurement of compressed sward height and estimation of yield on grasslands. The aim of this study was to investigate the use of an RPM to evaluate lucerne height as an agronomic trait enabling yield estimation. Compressed height (CH), stem length, plant and stem density and yield measurements were recorded from the late‐vegetative stage to flowering in the first cut and from the late‐bud stage to flowering in the second and third cuts, during three successive years. For lucerne maximal stem‐length prediction, CH could be used successfully (with R ² over 0·90) in developmental stages before flowering and for stem lengths up to 80 cm, irrespective of stand density or order of cut. The dry‐matter yield of lucerne was better predicted using CH (R ² = 0·72) than maximal stem length (MSL) (R ² = 0·53). The recommended number of observations was 7–9 per treatment in the first cut and twice this in subsequent cuts because of higher variability in stand structure. The use of RPM to measure CH of lucerne is not recommended when MSL exceeds 80 cm, or in the case of lodging. RPM may also be suitable for similar forage crops with erect or semi‐erect stems, subject to consideration of disc weight and diameter. Heavy discs are unsuitable for lucerne height prediction but could be more suitable for yield prediction.     

The productivity of oats and berseem clover intercrops. II. Effects of cutting date and density of oats on annual forage yield

Annual forage yields of intercrops of berseem clover (Trifolium alexandrinum L.) and oats (Avena sativa L.), as affected by timing of initial harvest and sowing rate of oats, were investigated. Berseem clover was intercropped with oats at 0, 30, 60, 90 and 240 plants m^?2 of oats in 1999 and 2000 in Alberta, Canada. Cutting date treatments involved initial harvest at 10‐d intervals between 35 and 88 d after planting (DAP), and one or two subsequent harvests of regrowth. Total intercrop dry‐matter (DM) yield averaged 12·9–13·3 t ha^?1 with proportions of 0·21–0·43 of berseem clover in the forage. Treatments with an initial cut at the silage stage of oats (76 DAP in 1999, 88 DAP in 2000, at about soft‐dough stage of oats) had greater yields of oats, lower yields of berseem clover, and lower proportions of berseem clover in total annual yields than treatments with an earlier initial cut. Total crude protein (CP) yield was greater with an initial cut at 65–66 DAP than with a silage‐stage initial cut. With an initial cut at 35 DAP (before stem elongation of oats) or after 65 DAP (after heading of oats), yield potential of oats was sometimes reduced compared with silage‐stage treatments, but this was balanced by a greater yield of berseem clover. The impact of harvest timing on total yield decreased as the proportion of berseem clover in the intercrops increased. With decreasing density of oats, DM yield of first‐cut intercrops and total DM yield of oats decreased, while regrowth and total DM yields of berseem clover increased. Intercrops with oats at 60 plants m^?2 had equal or greater total DM and CP yields than intercrops with 240 plants m^?2 of oats. For intercrops with oats at 60 plants m^?2, with initial cuts at 65–66, 75–76, or 88 DAP, yields of regrowths were 0·30–0·35, 0·16–0·26 and 0·09 of the total yield respectively. Oats–berseem clover intercrops showed potential to manipulate the pattern of annual forage yield and to provide flexibility of harvest without reducing annual yields.     

Effect of heading date of perennial ryegrass cultivars on tillering and tiller development in spring and summer

Three cultivars (two diploid and one tetraploid) in each of three maturity groups (early, intermediate and late) of perennial ryegrass were sown in 10 m² plots, replicated four times, in Northern Ireland in June 1997 in a study of the effect of heading date on tiller development (including initiation to flower) and turnover of tillers produced at specific times in spring in 1998 and 1999. The plots were harvested seven times in each year. Annual dry‐matter production was similar for all groups in each year. In spring and early summer of both years, tiller density of the diploid cultivars was 1·5 times greater than that of the tetraploid cultivars and the mean tiller density over all swards in June was about 0·40 times greater than that in April. Maximum proportions of reproductive tillers in the early, intermediate and late maturity groups, determined from apical dissections, were found in early April, mid‐May and early June, respectively. Although a high proportion of tillers, which were present when annual observations commenced in spring, was decapitated at the first harvest in the early group, the previous population density was maintained by rapid production of new tillers during May, including those from suppressed tiller buds during reproduction. It is concluded that the relationship between heading date and rate of tiller turnover (including flowering) at specified times in spring is important in sward management throughout the early part of the growing season and should be taken into account in tiller‐based grass growth models.     

Nitrogen and Italian ryegrass. 3. Growth up to 14 weeks: yields, proportions, digestibilities and nitrogen contents of crop fractions, and tiller populations

The regrowth of Italian ryegrass following a cut in late April was studied at weekly intervals up to 14 weeks. There were four levels of applied N: 28, 84, 140 and 196 kg ha^-1. The proportion of green leaf blade declined from 80% of DM yield after 2 or 3 weeks of regrowth to 5% after 13–14 weeks. About three-quarters of the increase in DM yield of total herbage from week 3 to week 11 was attributable to increased yield of ‘stem’. The level of applied N had little effect on the proportion of crop fractions except during the period immediately before and after the begiiming of ear emergence when the higher rates of N increased the proportion of ‘stem’ and reduced that of green leaf. The most digestible crop fractions were green leaf, inflorescence and immature ‘stem’. D-value of ‘stem’ declined by about 20 units from week 1 to week 14, while D-value of green and dead leaf declined at about half this rate and there was little or no decline in D-value of inflorescence. The N content of green leaf was consistently higher than that of the other fractions. The N content of ‘stem’declined markedly to around 0.5% by week 12. The N content of dead leaf and inflorescence was intermediate between that of green leaf and mature ‘stem’. The N content of all fractions was increased by applying more N. The tiller population increased during the flrst 3 weeks of regrowth and declined during the next 4. Applying more N increased both tiller population and weight per tiller.     

Yield profile of tall fescue (Festuca arundinacea) in comparison with meadow fescue (F. pratensis) in Finland

Yield profile characteristics of tall fescue (Festuca arundinacea Schreb.), cv. Retu, were compared with those of meadow fescue (Festuca pratensis Huds.). The study was conducted in Finland and was based on official variety trial data recorded between 1980 and 1998 at 17 trial sites between latitudes 60° and 66°N. The crops were managed according to silage‐cutting regimes. The pattern of yield formation of the tall fescue cv. Retu differed significantly from that of meadow fescue cultivars, both within a growing season as well as in sward age. Tall fescue cv. Retu established slowly, and the dry‐matter (DM) yield from the first cut, made in the first year of harvest, was significantly lower than that for meadow fescue. In the second and third years, the DM yield from the first cut did not differ between tall fescue and meadow fescue. Tall fescue produced significantly higher DM yield as regrowth (second and third cuts) than meadow fescue. The sward age significantly affected total DM production. In first‐year swards, there were no significant differences in total DM yield between tall fescue and meadow fescue but, in the second‐ and third‐year swards, tall fescue produced significantly higher DM yields than meadow fescue. The difference in yield profile between tall fescue and meadow fescue was similar in all the environments included in the study. DM yield for the first cut (kg DM ha^–1) for tall fescue cv. Retu, in comparison with meadow fescue cultivars, was 2495 vs. 3099 (P < 0·001), 3735 vs. 3741 (NS, P=0·94) and 3553 vs. 3468 (NS, P=0·30) in the first‐, second‐ and third‐year swards respectively. The respective DM regrowth yields (second plus third cut) were: 6059 vs. 5416 (P < 0·001); 5445 vs. 4221 (P < 0·001); and 5580 vs. 4113 (P < 0·001) in first‐, second‐ and third‐year swards. Total DM yields per season for tall fescue vs. meadow fescue were (kg DM ha^–1) 8554 vs. 8515 (NS, P=0·69), 9180 vs. 7962 (P < 0·001) and 9133 vs. 7581 (P < 0·001) in first‐, second‐ and third‐year swards respectively. Over the three‐year sward rotation period, which is common in Finland, tall fescue produced on average 12% higher DM yield than meadow fescue. Both tall fescue and meadow fescue suffered little winter damage in Finnish conditions; the differences between cultivars of the two species were small. The results indicated that tall fescue cv. Retu is a productive and persistent forage species suited to Finnish growing conditions.     

Red clover for silage: management impacts on herbage yield,nutritive value,ensilability and persistence,and relativity to perennial ryegrass

This 6‐year experiment quantified the impacts of management factors on red clover yield, persistence, nutritive value and ensilability, and compared these with perennial ryegrass receiving inorganic N fertilizer. Within a randomized complete block design, field plots were used to evaluate a 2 (cultivar, Merviot and Ruttinova) × 2 (alone and with perennial ryegrass) × 2 (0 and 50 kg fertilizer N ha^?1 in mid‐March) × 2 (harvest schedule) combination of the factors relating to red clover, and a 2 (harvest schedule) × 4 (0, 50, 100 and 150 kg N ha^?1 for each cut) combination of the factors relating to perennial ryegrass. The early and late harvest schedules both involved four cuts per year, but commenced a fortnight apart. Red clover treatments averaged 14 906 kg dry matter (DM) ha^?1 per year, whereas perennial ryegrass receiving 600 kg inorganic N fertilizer per year averaged 14 803 kg DM ha^?1 per year. There was no yield decline evident across years despite a decline in the proportion of red clover. The early harvest schedule and sowing ryegrass with red clover increased the herbage yield and digestibility. March application of fertilizer N to red clover treatments reduced the annual yield. Early harvest schedule increased and both fertilizer N and sowing with ryegrass decreased the proportion of red clover. Sowing with ryegrass improved the indices of ensilability, but reduced the crude protein content. Both red clover cultivars had similar performance characteristics. A selected red clover‐based treatment, considered to exhibit superior overall production characteristics, outyielded N‐fertilized perennial ryegrass in mid‐season. However, it had poorer digestibility and ensilability indices.     

Effects of perennial ryegrass (Lolium perenne L.) cultivars on herbage production, nutritional quality and herbage intake of grazing dairy cows

Four perennial ryegrass (Lolium perenne L.) cultivars were compared for differences in herbage production, nutritive value and herbage intake of dry matter (DM) during the summers of 2002 and 2003. Two paddocks were sown with pure stands of four cultivars in a randomized block design with three replicates. Each plot was subdivided into fourteen subplots (22 m × 6 m) which were grazed by one cow during 24 h. Twelve lactating dairy cows were assigned to one cultivar for a period of 2 weeks in a 4 × 4 Latin square experimental design; the experiment lasted 8 weeks in each year. Sward structure (sward surface height, DM yield, green leaf mass, bulk density and tiller density) and morphological characteristics were measured. The ash, neutral‐detergent fibre, acid‐detergent lignin, crude protein and water‐soluble carbohydrate concentrations, and in vitro digestibility of the herbage were measured. The sward was also examined for infestation by crown rust (Puccinia coronata f. sp. lolii). Herbage intake of dairy cows was estimated using the n‐alkane technique. Cultivar differences for all sward structural characteristics were found except for bulk density and tiller density in 2003. Cultivars differed for proportions of pseudostem, stem (in 2003 only) and dead material. The chemical composition of the herbage was different among cultivars, with the water‐soluble carbohydrate concentration showing large variation (>0·35). Cultivars differed in susceptibility to crown rust. Herbage intake differed among cultivars in 2002 (>2 kg DM) but not in 2003. Herbage intake was positively associated with sward height, DM yield and green leaf mass. Canopy morphology did not affect herbage intake. Crown rust affected herbage intake negatively. It was concluded that options for breeders to select for higher intake were limited. High‐yielding cultivars and cultivars highly resistant to crown rust were positively related with a high herbage intake.     

Dry-matter yields and crude protein and rumen-degradable protein concentrations of three Arachis pintoi ecotypes at different stages of regrowth in the humid tropics

Well‐established stands of three ecotypes of Arachis pintoi (CIAT 17434, 18744 and 18748) were harvested from replicated plots (three blocks, each containing three plots for each ecotype) during the two dominant seasons (dry and wet) of the low altitude, humid tropics of Costa Rica. Each plot was further divided into six subplots so that, within each season, samples corresponding to 4, 6, 8, 10, 12 or 14 weeks of regrowth could be collected. For each harvest, dry matter (DM) yield of the leaf, stem and whole plant, and the leaf:stem ratio, were recorded. Samples of the whole plant were analysed for crude protein (CP), rumen‐degradable (RDP) and rumen‐undegradable protein (RUP) concentrations. DM yield of the leaf, stem and whole plant increased with advancing period of regrowth but the effects of period of regrowth varied somewhat among ecotypes and across seasons. Generally, DM yield was greater during the wet than during the dry season. The greatest difference between ecotypes for stem and total DM yields was evident during the dry season. In general, DM contents were low in the whole plant, leaf and stem samples (<220 g kg^?1) and increased with increasing period of regrowth. Increases in leaf:stem ratio were most dramatic during the dry season with greater periods of regrowth, although the ratio was fairly constant during the wet season. Whole‐plant CP concentration was relatively high after short periods of regrowth (up to 279 g kg^?1 DM) but declined with longer periods of regrowth; the relative decline was much greater during the dry season. The RDP concentration was relatively constant during the wet season (mean 115 g kg^?1 DM), but declined with longer periods of regrowth during the dry season (range 194–111 g kg^?1 DM). In general, the concentrations of RDP, on a CP basis, were greater during the dry season and ranged from 590 to 700 g kg^?1 CP. Season, ecotype and period of regrowth all exerted an effect on RUP concentrations.     

Changes during growth in the nutritive value of components of berseem clover (Trifolium alexandrinum L.) under different cutting treatments in a Mediterranean region

The nutritive value of berseem clover (Trifolium alexandrinum L.) may be influenced by changes in the nutrient concentrations of morphological fractions as a consequence of cutting treatment. The aim of this study was to evaluate the effects of different cutting treatments on the nutritive value of herbage and leaf and stem components in two Mediterranean berseem genotypes during growth in order to develop management approaches for harvesting forage with a high nutritive value. Spring growth of genotypes of Egyptian (cv. Giza 10) and Italian (cv. Sacromonte) origins was harvested in each of 2 years beginning 196 days after sowing and thereafter every 6 days (twelve harvests in total) to measure dry matter (DM) yield and nutritive value. Cutting treatments were initiated at sixth internode elongation (A) and early flowering (B) and there was an uncut control treatment (C). In vitro digestibility of organic matter (IVOMD) and crude protein (CP) concentration were determined for leaf, stem and total forage of each cultivar at each harvest. For both cultivars, in the uncut treatment (C), DM yield increased linearly to a maximum of 14 800 kg ha^?1, on average, by 250 days after planting, the same time at which stem DM accumulation peaked, whereas the nutritive value, in terms of IVOMD and CP concentration, declined with age, coinciding with a reduction in leaf:stem ratio (LSR) from 1·00 to 0·30. Plant parts differed (P < 0·01) in nutritive value with stems being of lower nutritive value than leaves, in the two cutting treatments. There were strong positive correlations between LSR and herbage IVOMD and CP concentration, in both cutting treatments, indicating that, during growth, part of the changes in IVOMD and CP concentration of berseem clover plants was due primarily to the changes in the LSR. Defoliation induced a considerable reduction in DM yield, but an increase in the LSR and a small increase in nutritive value. Plants cut at the sixth internode elongation (A) showed a small proportional decrease (0·19) in total DM yield, but higher LSR values and similar or higher IVOMD and CP concentration than initiating cutting at early flower (B). Delaying defoliation to the early flowering stage (B) increased the proportion of stems and, therefore, decreased nutritive value. Therefore, harvest management in which cuts are applied at the stage of sixth internode elongation appears to be the most favourable for obtaining relatively high yields of forage with high nutritive value in berseem clover grown in Mediterranean regions.     

Sward growth under cutting and continuous stocking managements: sward canopy structure, tiller density and leaf turnover

The change in structure of continuously grazed versus infrequently cut swards of perennial ryegrass ( Lolium perenne L), cv. S23, was investigated during their first full harvest year. Measurements were made from early May until late September. The intensity of stocking by sheep in the grazed sward was adjusted in an attempt to maintain a high level of radiation interception and the cut sward was harvested at approximately monthly intervals.
The herbage mass, lamina area index and radiation interception of the cut sward varied in a cyclic pattern between harvests but in the grazed sward these parameters showed considerably less variation, although they all increased early in the season and then declined later. The proportion of dead material above ground increased throughout the season in both sward types but was more marked in the grazed sward.
There were major differences between the grazed and cut swards in the number of tillers per unit ground area; the difference became more marked throughout the season and by September the tiller densities in the grazed and cut swards were 3·20⁴ m^-2 and 6·20³ m^-2 respectively. Divergence in tiller density was associated with differences in specific stem weight and leaf area per tiller.
Rates of appearance and death of leaves on tillers in the grazed sward were determined. During May, leaf appearance exceeded leaf death but this was reversed in June. During the rest of the season as a new leaf appeared on a tiller so the oldest leaf died.     

Spring and autumn animal treading effects on pre‐grazing herbage mass and tiller density on two contrasting pasture types in Ireland

A perennial ryegrass (Lolium perenne L.)‐dominated sward on a well‐drained soil (Experiment 1) and a creeping bent (Agrostis stolonifera L.)‐dominated sward on a poorly drained soil (Experiment 2) were subjected to four treading treatments: control (C, no damage), light damage (L), moderate damage (M) or severe damage (S) to quantify the effects on herbage dry‐matter (DM) production and tiller density. In Experiment 1, treading damage was imposed in spring. In Experiment 2, one‐third of the site was damaged in autumn, one‐third in spring and one‐third in both spring and autumn. Both sites were rotationally grazed after treading treatments. Pre‐grazing herbage mass was measured eight times in Experiment 1 and seven times in Experiment 2 on each plot, and tiller density was assessed four times in each experiment. In Experiment 1, pre‐grazing herbage mass was reduced by 30% in S plots at the first harvest after damage, but cumulative pre‐grazing herbage DM production was not different between treatments (12·7 t DM ha^?1). In Experiment 2, annual cumulative pre‐grazing herbage mass was reduced by between 14 and 49%, depending on intensity of treading damage event and season when damage occurred. Tiller density was not affected by treatment in either experiment. A perennial ryegrass‐dominated sward on a well‐drained soil was resilient to heavy treading damage. A creeping bent‐dominated sward on poorly drained soil requires a more careful grazing management approach to avoid major losses in cumulative pre‐grazing herbage mass production during wet weather grazing events.     

Influence of initial cutting date and cutting frequency on yield and quality of star, elephant and Guinea grasses

Two experiments were conducted to estimate the influence of initial cutting dates in March and April and of cutting frequencies on Guinea grass, elephant grass and star grass productivity. Increasing delay in date of first cut in the season resulted in a progressive increase in dry-matter yield. 72–81% of the tiller population at any cut were vegetative and this helped to maintain good aftermath yield for each date of first cut. Early April cuts gave the highest dry-matter yield and early March the least.
Harvesting frequencies affected the sward productivity such that the annual dry matter yield increased with increasing harvesting interval from 6800 kg ha^-1 for a 3-week interval to 13,000 kg ha^-1 for a 10-week interval. The proportion of green leaf in the dry matter dropped from 57·7% at 3-week intervals to 32·0% at 10-week intervals. Seasonal effects showed that potential yield was reduced by a short interval between harvests early in the season but not late in the season. Species differences in quality and in the harvesting interval that gave the maximum yields were noted. These results are discussed in relation to management of the sward throughout the growing season.