宽叶缬草短缩茎的结构及石细胞的分布与发育

对宽叶缬草短缩茎的结构及其石细胞分布和发育进行系统观察.结果表明,宽叶缬草短缩茎具有茎尖、腋芽、叶迹和根迹维管束等组织和器官,是完整的营养繁殖体.茎尖明显分为顶端分生组织、伸长区和成熟区,成熟区为初生结构,所占比例最大,成熟区的上、下部结构组成成分上有所不同.短缩茎的薄壁细胞中含有大量白色体、造油体,少数有橙黄色橙皮苷针簇状结晶,是缬草素等次生代谢产物的贮藏器官.短缩茎内茎节的髓部有堆状或带状分布的石细胞,由石细胞原基细胞快速侵入式生长,细胞壁木质化增厚,周围薄壁细胞随之继续增厚而形成.

Structure of the shortening stem of Valeriana officinalis L.

By systematical observation to the shortening stem structure and developing process of the sclereids in stem of Valeriana officinalis L.var.latifolia Miq., this paper showed that the shortening stem was a vegetative propagule which consisted of shoot apex, axillary buds, vascular bundles of leaf trace and root trace,etc. The shoot tip revealed three regions called apical meristem, elongation zone, and maturation zone. Expect orange hesperidin crystal, there were many leucoplasts, oil bodies accumulated in the parenchyma cells in the shortening stem. Clusters of sclereids chambers were distributed widely in the pith of the shortening stem node, which was initiated from rapid apical intrusive growth of sclereid initial cell, the deposition of a secondary lignifed cell wall in it and the parenchyma cells around it as well.