The effects of shade on the plant communities of rides in plantation woodland and implications for butterfly conservation

During 1985-6 the vegetation in the rides (forest tracks) of eight English lowland woods, planted on ancient woodland sites was surveyed. Shade was measured in each transect by hemispherical photography. Dicotyledonous species richness and the majority of dicot plant species declined as shade increased. The results for individual species compare favourably with Ellenberg's (L) value in most cases, and a method is suggested for modifying Ellenberg's data in the light of new information. The woodlands contained species of both open and shaded habitats and communities were influenced by soil acidity. Because shade in woodland rides increases as the trees mature, theory states that there will almost certainly be a reduction in plant species diversity. Butterfly populations would also be adversely affected, partly because of the reduction in direct sunlight, and partly because their larval foodplants decline in both quantity and quality since they are generally associated with low to moderate shade.