古尔班通古特沙漠生物结皮中藓类植物形态解剖特征

对组成古尔班通古特沙漠生物土壤结皮的4种1变种藓类植物刺叶墙藓(Tortula desertorum Broth.)、真藓(Bryum argenteum Hedw.)、绿色流苏藓〔Crossidium squamiferum(Viv.)Jur.〕、泛生墙藓(Tortula muralis Hedw.)、泛生墙藓无芒变种(Tortula muralis var. aestiva Brid. ex Hedw.)的茎、叶形态解剖结构进行了观察。结果表明:除刺叶墙藓叶片上部为2层细胞组成外,其他4种叶片均由1层细胞组成。所有藓类植物叶片都具有明显或粗壮的中肋,起到了支持叶片和导水的作用。茎皮部外层细胞壁不同程度加厚。生物结皮中的藓类植物表现出叶片背卷或内卷、具疣状突起、透明毛尖(泛生墙藓无芒变种除外)以及边缘细胞壁加厚等特征,这些特征是对干旱荒漠环境长期适应的结果。研究结果为进一步揭示荒漠藓类植物形态特征与环境之间的关系提供理论依据。     

新疆薄罗藓科(Leskeaceae)植物分类的初步研究

通过对产自新疆的薄罗藓科5属8种的分类学研究,包括多毛藓属Lescuraea Bruch & Schimp. in B.S.G. 的弯叶多毛藓L. incurvata (Hedw.) Lawton,密根多毛藓 L. radicosa (Mitt.) Moeck.,石生多毛藓L. saxicola (Schimp. in B.S.G.) Mol. in Lor.;薄罗藓属Leskea Hedw. 的薄罗藓L. polycarpa Hedw. ;细罗藓属Leskeella (Brid.) Loeske 的细罗藓L. nervosa (Brid.)Loeske ;细枝藓属Lindbergia (C. Müll) Broth. 的中华细枝藓L. sinensis (C. Müll) Broth. ,以及假细罗藓属 Pseudoleskeella Kindb. 的假细罗藓P. catenulata (Brid. ex Schrad.) Kindb.和瓦叶假细罗藓P. tectorum (Brid. )Kindb.。其中,薄罗藓属Leskea Hedw.为新疆新记录属;薄罗藓Leskea polycarpa Hedw. 和密根多毛藓Lescuraea radicosa (Mitt.) Moeck. 是新疆新记录种。根据标本形态特征,编制了新疆薄罗藓科植物分属、分种检索表,简要讨论了其生境和地理分布特点。     

河北省薄罗藓科植物分类的初步研究

报道了河北省薄罗藓科植物6属共12种（含1变种），其中中国新记录种1种：北方假细罗藓Pseudoleskeella papillosa(Lindb.)Kindb.；河北省新记录属2属：薄罗藓属Leskea Hedw.和褶藓属Okamuraea Broth.；河北新记录种7种：薄罗藓Leskea polycarpi Hedw.，细枝藓Lindbergia brachyptera(Mitt.)Kindb.，细枝藓疣齿变种L. brachyptera var.austinii (Sull.)Grout.,阔叶细枝藓L. brevifolia(Gao)Gao，瓦叶假细罗藓Pseudoleskeella tectorum(Brid.)Kindb.,褶藓O.hakoniensis(Mitt.)Broth.和短枝褶藓O.brachydictyon(Card.）Nog..本文根据标本编制了河北省薄罗藓科植物分属、种检索表，简要讨论了其生境行理分布特点。旨在为河北省乃至中国苔藓植物的系统研究及苔藓植物资源的开发利用提供可借鉴的资料。研究标本保存在河北师范大学植物标本室（HBNU)。     

黄土高原生物结皮形成过程中土壤胞外酶活性及其化学计量变化特征

生物土壤结皮在增强土壤抵抗水蚀风蚀能力、改善土壤养分等方面发挥着重要作用,土壤胞外酶活性可作为土壤生化反应强度的微生物指标,对理解荒漠生态系统微生物参与的养分循环过程具有重要意义.选取黄土高原神木县六道沟小流域5个生物结皮发育阶段(裸沙地、全藻结皮、藻-藓混生结皮、藓-藻混生结皮、全藓结皮)的土壤作为研究对象,研究生物...     

十种藓类植物茎的比较解剖学观察

利用石蜡切片的方法 ,对十种藓类植物 ,钝叶蓑藓 (Macromitriumjaponicum Doz.et.Molk .)、疏齿墙藓〔 Tortulanorvegica(Web .)Wahl.exLindb .〕、岸生连轴藓〔 Schistidiumrivularis(Brid .)Podp .〕、树形疣灯藓〔Trachycystisussuriensis(MaacketRegel )T .Kop .〕、齿边缩叶藓〔 Ptychomitriumdentatum(Mitt.)Jaeg .〕、山羽藓〔 Abietinellaabietina(Hedw .)Fleisch〕、东亚万年藓 ( Climaciumjaponicum Lindb .)鼠尾藓〔 Myurocladamaximowiczii(Borszcz)SteeretSchof.〕、卷叶凤尾藓 (FissidensdubiusP .Beauv .)及二形凤尾藓 (FissidensgeminiflorusDoz .etMolk .)的茎横切面的解剖观察结果表明 :茎的形状、中轴细胞的有无、形状及所占的比例、外皮部厚壁细胞的层数等特征因种类不同而有明显差异 ,可以作为分类依据之一。外皮部厚壁细胞的层数可能与藓类植物的生境有关。     

黄土高原藓结皮覆盖土壤导水性能和水流特征

生物结皮具有特殊的水文物理性质,为探究其对土壤水分渗透性和水流特征的影响,以黄土高原风沙土和黄绵土上3种典型地表覆盖类型(裸地、藓结皮、藓结皮-草本植物混合)为对象,采用环刀法和染色示踪法对其导水性质与水流特征进行探究。结果表明:藓结皮对2种土壤类型0~5 cm土层土壤理化性质影响较大,与裸地相比土壤容重降低了9.85%~10.00%,土壤黏粒含量增加了1.01~1.29倍,表层有机质含量提高了2.73~3.02倍;藓结皮使0~5 cm土层土壤饱和导水率降低了61.32%~88.89%,而在5~10 cm土层饱和导水率则有明显上升。另外,由于草本植物的影响,藓结皮-草本植物0~5 cm土层与藓结皮土壤相比土壤饱和导水率提高了1.32~6.43倍;黄绵土藓结皮与藓结皮-草本植物的染色面积比均高于裸地,且水分下渗深度增加了10 cm,而风沙土藓结皮与风沙土裸地的染色面积比差异不明显。综上所述,藓结皮和藓结皮-草本植物的存在改变了表层土壤水分渗透性以及水流运动特征和水分下渗深度,影响着黄土高原土壤水分保持和生态恢复。     

生物结皮富集营养元素和重金属元素的空间分异特性

通过测定库布齐沙漠腹地达拉特旗火力发电厂附近沙地中不同年代人工结皮和下层土壤中重金属和营养元素的含量,旨在研究生物结皮富集重金属和营养元素的能力,富集结果及空间差异性。通过单因素方差分析(ANOVA)和比较,得出以下结论:在水平格局,结皮中的重金属和营养元素含量表现为:Cr>Pb>Cu>Cd和C>K>N>P的顺序关系,相同发育年代结皮中的重金属和营养元素含量均表现为:藓结皮>藻结皮>物理结皮;不同发育年代结皮中各种重金属和营养元素含量均表现为随着发育时间的增加而增加;在垂直格局,各类型结皮中重金属和营养元素含量均表现为随着土壤垂直深度增加而降低。     

土壤水分对毛乌素沙地油蒿群落演替的影响

采用空间代替时间的方法,对毛乌素沙地三种不同演替阶段的油蒿(Artemisia ordosica)群落(半固定沙地油蒿群落、具生物结皮的油蒿群落、油蒿+本氏针茅群落)的群落特征及生物量、叶片气体交换过程、不同土层(0-10cm、10-20cm、20-40cm、40-60cm、60-80cm)土壤含水量、根系垂直分布等进行测定.结果表明:油蒿群落演替是油蒿与草本植物竞争、生物结皮发育、土壤发育以及土壤水分垂直变化等生物非生物因素综合作用的结果.随着油蒿群落演替的发展,群落物种数、草本层盖度和生物量、群落总盖度逐渐增加,而油蒿个体数、油蒿盖度、油蒿叶生物量以及生物土壤结皮盖度则先增加后减少,油蒿根系有向下分布的趋势,0-10cm土层逐渐以草本植物根系占优势.土壤水分是毛乌素沙地油蒿群落演替的主要驱动力,通过影响油蒿叶片气孔导度和净光合速率,以及根系垂直分布影响油蒿在群落中的竞争力.不同演替阶段油蒿群落土壤水分低值区不同,半固定沙地油蒿群落0-10cm和20-40cm土壤含水量最低,具生物结皮的油蒿群落10-20cm土壤含水量最低,而油蒿+本氏针茅群落60-80cm土壤含水量最低.     

古尔班通古特沙漠生物结皮影响下土壤水分的日变化

对新疆古尔班通古特沙漠生物结皮影响下土壤水分的日变化特征进行了定量研究,结果表明:①无论是结皮覆盖区还是无结皮覆盖区,其土壤含水量随深度的变化均表现出明显的层次性,其中以距地表 20 cm处的土壤含水量最高.②结皮覆盖区距地表5 cm和10 cm处的土壤含水量显著高于无结皮覆盖区(LSD检验, p ＜0.05);而45 cm和60 cm处的土壤含水量则表现为结皮覆盖区极显著低于无结皮区(LSD检验,p＜0.05),说明生物结皮具有较强的保持土壤表层水分的能力.③结皮覆盖区与无结皮覆盖区土壤水分的日变化特征明显不同于距地表5 cm处,而其他各土层的变化趋势则较为一致.无结皮覆盖区距地表5 cm处土壤水分的日变化呈先上升后下降的趋势,而结皮覆盖区的变化则恰恰相反.     

毛乌素沙地生物结皮对土壤温度的影响

土壤温度对作物生长发育、土壤水盐运移、生物的数量与活性、土壤碳平衡等均有较大的影响。为了研究毛乌素沙地生物结皮区和裸沙区土壤温度的差异,于2010年9~11月在陕北毛乌素沙地东南缘神木县圪丑沟地区,选择晴朗天气进行定位观测试验。结果表明: ① 同一剖面深度,生物结皮样地土壤温度的日变化趋势同裸沙样地一致;且从6:00~21:00时每隔3 h,土壤垂直剖面的温度沿着“\” 型、 微“（”型、明显“（”型、“/” 型、微“）”型、“\”型进行周期性变化。② 生物结皮的覆盖极显著降低了地表的土壤温度（[WTBX]P[WTBZ]＜0.01）。③ 生物结皮样地和裸沙样地表层土壤温度变幅最大,随剖面深度的增加土壤温度变化趋缓。当气温和土壤含水量较低时,生物结皮对同一深度土壤温度变化幅度的影响更明显。从垂直剖面角度看,生物结皮降低了土壤温度及其变幅。④ 生物结皮提高了0~25 cm深度土壤温度对气温的敏感程度。表明土壤温度的变化是多因素作用的结果,前人仅根据生物结皮的颜色来判断其对土壤温度的影响是不够的。生物结皮对土壤温度造成影响的主要原因,可能是其改变了土壤表层结构及土壤水分状况。在今后相关研究中如水分蒸发、土壤化学性状的改变等,要特别注意生物结皮的温度效应对这些过程的影响,从而使分析更加客观深入。     

Carbon fixation and influencing factors of biological soil crusts in a revegetated area of the Tengger Desert,northern China

Biological soil crusts （BSCs） are an important type of land cover in arid desert landscapes and play an important role in the carbon source-sink exchange within a desert system. In this study, two typical BSCs, moss crusts and algae crusts, were selected from a revegetated sandy area of the Tengger Desert in northern China, and the experiment was carried out over a 3-year period from January 2010 to November 2012. We obtained the effec- tive active wetting time to maintain the physiological activity of BSCs basing on continuous field measurements and previous laboratory studies on BSCs photosynthesis and respiration rates. And then we developed a BSCs carbon fixation model that is driven by soil moisture. The results indicated that moss crusts and algae crusts had significant effects on soil moisture and temperature dynamics by decreasing rainfall infiltration. The mean carbon fixation rates of moss and algae crusts were 0.21 and 0.13 g C/（m2.d）, respectively. The annual carbon fixations of moss crusts and algae crusts were 64.9 and 38.6 g C/（m2.a）, respectively, and the carbon fixation of non-rainfall water reached 11.6 g C/（m2.a） （30.2% of the total） and 8.8 g C/（m2.a） （43.6% of the total）, respectively. Finally, the model was tested and verified with continuous field observations. The data of the modeled and measured CO2 fluxes matched notably well. In desert regions, the carbon fixation is higher with high-frequency rainfall even the total amount of seasonal rainfall was the same.     

Responses of microbial activities and soil physical-chemical properties to the successional process of biological soil crusts in the Gurbantunggut Desert,Xinjiang

Biological soil crusts(BSCs) are capable of modifying nutrient availability to favor the establishment of biogeochemical cycles. Microbial activities serve as critical roles for both carbon and nutrient transformation in BSCs. However, little is known about microbial activities and physical-chemical properties of BSCs in the Gurbantunggut Desert, Xinjiang, China. In the present research, a sampling line with 1-m wide and 20-m long was set up in each of five typical interdune areas selected randomly in the Gurbantunggut Desert. Within each sampling line, samples of bare sand sheet, algal crusts, lichen crusts and moss crusts were randomly collected at the depth of 0–2 cm. Variations of microalgal biomass, microbial biomass, enzyme activities and soil physical-chemical properties in different succession of BSCs were analyzed. The relationships between microalgal biomass, microbial biomass, enzymatic activities and soil physical-chemical properties were explored by stepwise regression. Our results indicate that microalgal biomass, microbial biomass and most of enzyme activities increased as the BSCs developed and their highest values occurred in lichen or moss crusts. Except for total K, the contents of most soil nutrients(organic C, total N, total P, available N, available P and available K) were the lowest in the bare sand sheet and significantly increased with the BSCs development, reaching their highest values in moss crusts. However, pH values significantly decreased as the BSCs developed. Significant and positive correlations were observed between chlorophyll a and microbial biomass C. Total P and N were positively associated with chlorophyll a and microbial biomass C, whereas there was a significant and negative correlation between microbial biomass and available P. The growth of cyanobacteria and microorganism contributed C and N in the soil, which offered substrates for enzyme activities thus increasing enzyme activities. Probably, improvement in enzyme activities increased soil fertility and promoted the growth of cyanobacteria, eukaryotic algae and heterotrophic microorganism, with the accelerating succession of BSCs. The present research found that microalgal-microbial biomass and enzyme activities played important roles on the contents of nutrients in the successional stages of BSCs and helped us to understand developmental mechanism in the succession of BSCs.     

Effects of artificially cultivated biological soil crusts on soil nutrients and biological activities in the Loess Plateau

Biological soil crusts （BSCs） play an important role in the early succession of vegetation restoration in the Loess Plateau, China. To evaluate the effects of artificially cultivated BSCs on the soil surface micro-envir- onment, we obtained natural moss crusts and moss-lichen crusts from the Loess Plateau of Shaanxi province, and subsequently inoculated and cultivated on horizontal and sloping surfaces of loess soil in a greenhouse. The chemical and biological properties of the subsoil under cultivated BSCs were determined after 10 weeks of cul- tivation. The results indicated that BSCs coverage was more than 65% after 10 weeks of cultivation. Moss crust coverage reached 40% after 5 weeks of cultivation. Compared with the control, soil organic matter and available nitrogen contents in moss crust with the horizontal treatments increased by 100.87% and 48.23%, respectively; increased by 67.56% and 52.17% with the sloping treatments, respectively; they also increased in moss-lichen crust with horizontal and sloping treatments, but there was no significant difference. Available phosphorus in cultivated BSCs was reduced, soil pH was lower and cationic exchange capacity was higher in cultivated BSCs than in the control. Alkaline phosphatase, urease and invertase activities were increased in artificially cultivated BSCs, and alkaline phosphatase activity in all cultivated BSCs was obviously higher than that in the control. Numbers of soil bacteria, fungi and actinomycetes were increased in the formation process of cultivated BSCs. These results indicate that BSCs could be formed rapidly in short-term cultivation and improve the mi- cro-environment of soil surface, which provides a scientific reference for vegetation restoration and ecological reconstruction in the Loess Plateau. China.     

Effects of biological soil crusts and drought on emergence and survival of a Patagonian perennial grass in the Monte of Argentina

Biological soil crusts are widely distributed in arid and semiarid regions. They have an important eco- logical role, especially by modifying physical and chemical properties of soils. Biological crusts may also modify seed germination and seedling establishment. The effects vary widely according to the type of crust and the vas- cular plant species. The objective of this study was to determine the effect of moss-dominated biological soil crusts on the emergence, biomass and survival of Poa ligularis Nees ex Steud. under different irrigation regimes. We col- lected seeds of P. ligularis and biological soil crusts composed of two species of mosses： Syntrichia princeps （De Not.） Mitt and Ceratodon purpureus （Hedw.） Brid. from an area in the Monte of Argentina. The result showed that seedling emergence of P. ligularis was higher in treatments with bare soil than in soil covered by crusts, and also in those with watering to field capacity. Mean emergence time was higher in treatments with bare soil and watering to field capacity. Seedling biomass also showed significant differences between treatments. These results suggest that biological soil crusts dominated by mosses do not promote P. ligularis emergence, although they would not affect its survival.     

Microbiotic Crust Influence on Unsaturated Hydraulic Conductivity

Microbiotic crusts occur extensively in rangeland soils . Developed by filaments of cyanobacteria and algae , and thalli of lichen and moss entanglement of soil particles , they create a physical discontinuity in the surface profile with greater concen trations of clay , silt , and potentially hydrophobic organic matter. These conditions potentially contribute to variability in soil hydrology of arid land and should be considered in the development of hydrologic and erosion models. However, there is limited manipulative research examining the functional relationships between soil and microbiotic crusts . We investigated the influence of cyanobacterial-dominated microbiotic crust on measured hydraulic conductivity (K) in a sandy loam soil at a southeastern Utah site . Using a tension infiltrometer , we determined K under three surface treatments: undisturbed , chemically killed (representing dead microphytes within the crust), and removed (scalped) microbiotic crusts. We applied treatments to spatially interspersed intact surface soils within shrub interspaces . Microbiotic crusts at this site and in this stage of successional development had no discernible influence on K. This finding supports results from research conducted in a variety of soils from sandy to silt dominated with a range of microbiotic development . Because this research was site and time specific , and because the role of microbiotic crusts in the environment continues to be debated , additional research is warranted to deter mine how stage of development of microbiotic crust influences soil hydrology .     

黄土地表生物结皮对土壤贮水性能及水分入渗特征的影响

采用双环法对山西省偏关县3种结皮覆盖下(苔藓藻结皮、藻结皮和薄层藻结皮)土壤的贮水性能和渗透特征进行了对比研究.结果表明:不同结皮覆盖下土壤的贮水能力受结皮厚度和孔隙度状况的影响较大,0～10 cm土层饱和贮水量为502.69～525.80 t/hm2 ,滞留贮水量为169.71～198.29 t/hm2;初渗速率的变化范围为5. 19～11.10 mm/min,无结皮最高,苔藓藻结皮最低;稳渗速率变化范围为1.6 7～2.67 mm/min.采用的3种入渗模型(Kostiakov模型、Horton模型和Ph ilip模型)中Horton模型的拟合值更接近于实测值, 决定系数R2在0.98～0.99,更适用于描述本研究区具有生物结皮土壤的入渗特征.     

Shrub modulates the stoichiometry of moss and soil in desert ecosystems,China

Desert mosses, which are important stabilizers in desert ecosystems, are distributed patchily under and between shrubs. Mosses differ from vascular plants in the ways they take up nutrients. Clarifying their distribution with ecological stoichiometry may be useful in explaining their mechanisms of living in different microhabitats. In this study, Syntrichia caninervis, the dominant moss species of moss crusts in the Gurbantunggut Desert, China, was selected to examine the study of stoichiometric characteristics in three microhabitats (under living shrubs, under dead shrubs and in exposed ground). The stoichiometry and enzyme activity of rhizosphere soil were analyzed. The plant function in the above-ground and below-ground parts of S. caninervis is significantly different, so the stoichiometry of the above-ground and below-ground parts might also be different. Results showed that carbon (C), nitrogen (N) and phosphorus (P) contents in the below-ground parts of S. caninervis were significantly lower than those in the above-ground parts. The highest N and P contents of the two parts were found under living shrubs and the lowest under dead shrubs. The C contents of the two parts did not differ significantly among the three microhabitats. In contrast, the ratios of C:N and C:P in the below-ground parts were higher than those in the above-ground parts in all microhabitats, with significant differences in the microhabitats of exposed ground and under living shrubs. There was an increasing trend in soil organic carbon (SOC), soil total nitrogen (STN), soil available phosphorous (SAP), and C:P and N:P ratios from exposed ground to under living shrubs and to under dead shrubs. No significant differences were found in soil total phosphorous (STP) and soil available nitrogen (SAN), or in ratios of C:N and SAN:SAP. Higher soil urease (SUE) and soil nitrate reductase (SNR) activities were found in soil under dead shrubs, while higher soil sucrase (STC) and soil β-glucosidase (SBG) activities were respectively found in exposed ground and under living shrubs. Soil alkaline phosphatase (AKP) activity reached its lowest value under dead shrubs, and there was no significant difference between the microhabitats of exposed ground and under living shrubs. Results indicated that the photosynthesis-related C of S. caninervis remained stable under the three microhabitats while N and P were mediated by the microhabitats. The growth strategy of S. caninervis varied in different microhabitats because of the different energy cycles and nutrient balances. The changes of stoichiometry in soil were not mirrored in the moss. We conclude that microhabitat could change the growth strategy of moss and nutrients cycling of moss patches.     

Precipitation and soil particle size co-determine spatial distribution of biological soil crusts in the Gurbantunggut Desert,China

Biological soil crusts(BSCs) are bio-sedimentary associations that play crucial ecological roles in arid and semi-arid regions. In the Gurbantunggut Desert of China, more than 27% of the land surface is characterized by a predominant cover of lichen-dominated BSCs that contribute to the stability of the desert. However, little is known about the major factors that limit the spatial distribution of BSCs at a macro scale. In this study, the cover of BSCs was investigated along a precipitation gradient from the margins to the center of the Gurbantunggut Desert. Environmental variables including precipitation, soil particle size, soil p H, electrical conductivity, soil organic carbon, total salt, total nitrogen, total phosphorus and total potassium were analyzed at a macro scale to determine their association with differing assemblages of BSCs(cyanobacteria crusts, lichen crusts and moss crusts) using constrained linear ordination redundancy analysis(RDA). A model of BSCs distribution correlated with environmental variables that dominated the first two axes of the RDA was constructed to clearly demonstrate the succession stages of BSCs. The study determined that soil particle size(represented by coarse sand content) and precipitation are the most significant drivers influencing the spatial distribution of BSCs at a macro scale in the Gurbantunggut Desert. The cover of lichen and moss crusts increased with increasing precipitation, while the cover of cyanobacteria crusts decreased with increasing precipitation. The cover of lichen and moss crusts was negatively associated with coarse sand content, whereas the cover of cyanobacteria crusts was positively correlated with coarse sand content. These findings highlight the need for both the availability of soil moisture and a relatively stable of soil matrix, not only for the growth of BSCs but more importantly, for the regeneration and rehabilitation of disturbed BSC communities in arid and semi-arid lands. Thereby, this study will provide a theory basis to effectively increase soil stability in desert regions.     

Topographic differentiations of biological soil crusts and hydraulic properties in fixed sand dunes,Tengger Desert

Biological soil crusts(BSCs) play an important role in surface soil hydrology. Soils dominated with moss BSCs may have higher infiltration rates than those dominated with cyanobacteria or algal BSCs. However, it is unnown whether improved infiltration in moss BSCs is accompanied by an increase in soil hydraulic conductivity or water retention capacity. We investigated this question in the Tengger Desert, where a 43-year-old revegetation program has promoted the formation of two distinct types of BSCs along topographic positions, i.e. the moss-dominated BSCs on the interdune land and windward slopes of the fixed sand dunes, and the algal-dominated BSCs on the crest and leeward slopes. Soil water retention capacity and hydraulic conductivity were measured using an indoor evaporation method and a field infiltration method. And the results were fitted to the van Genuchten–Mualem model. Unsaturated hydraulic conductivities under greater water pressure(–0.01 MPa) and water retention capacities in the entire pressure head range were higher for both crust types than for bare sand. However, saturated and unsaturated hydraulic conductivities in the near-saturation range(–0.01 MPa) showed decreasing trends from bare sand to moss crusts and to algal crusts. Our data suggested that topographic differentiation of BSCs significantly affected not only soil water retention and hydraulic conductivities, but also the overall hydrology of the fixed sand dunes at a landscape scale, as seen in the reduction and spatial variability in deep soil water storage.