杂草科学的发展与度规的变革

本文以杂草科学中有关科学度规的变革为切入点，阐述了人类与杂草作斗争的历史，分析了现代农业生产中除草方式的革命性变化及其困境，并探讨了将来的杂草问题及其对策、以及杂草科学的发展趋势。     

Development of lowland weed management and weed succession in Japan

Since the introduction of rice production in Japan, lowland areas have been managed for rice production with the purpose of better rice growth, as well as lesser weed infestation. Rice is cropped every year in lowland fields by repeated cultivation of a single crop, with high yields and without soil sickness usually being observed in upland fields. This is probably because the irrigation water supplies various nutrients for healthy rice growth and the drainage washes out and removes harmful factors. However, until recently, the wet or flooded conditions of lowland fields in the Asian monsoon region never have allowed humans to cultivate useful summer crops, except rice or some aquatic plants. Therefore, the management of lowland areas in the Asian monsoon region has been significantly different from European field management, where crop rotation has been the traditional standard practice. Paddy weeds are aquatic plants or hygrophytes that have adapted to lowland fields. Traditionally, tillage and puddling were practiced seasonally in lowland fields on a regular schedule every year. Rice cultivation technology was developed and supported by regional irrigation systems that created stable environments for typical paddy weeds to complete their life cycle. After the introduction of chemical weed control, rice fields became very severe habitats for these paddy weeds, where they could not grow and reproduce without strategies for survival under herbicide exposure. Even so, many of the traditional paddy weeds survived because of their accumulated or uneradicated seed banks, although several aquatic plants were listed as endangered or threatened species. The important weed species changed, sometimes rapidly and sometimes slowly, depending both on their reproductive system and their biological response towards field management and weed control systems. Very recently, the level of perennial weeds, herbicide‐resistant weeds, and weedy rice has increased in paddy fields that are highly dependent on herbicide use. In addition, several hygrophyte species have invaded paddy fields. In order to address these issues, the improvement and application of integrated weed management methods are expected to be critical.     

Ecological weed management by cover cropping: effects on weed growth in autumn and weed establishment in spring

Cover crops grown in the period between two main crops have potential as an important component of a system‐oriented ecological weed management strategy. In late summer and autumn, the cover crop can suppress growth and seed production of weeds, whereas the incorporation of cover crop residues in spring may reduce or retard weed emergence. Based on these two criteria, six cover crop species were evaluated for their weed suppressive potential in 2 years of experimentation in the Netherlands. Fodder radish, winter oilseed rape and winter rye had the strongest competitive ability in autumn; the competitive strength of Italian ryegrass was intermediate and white lupin and lucerne were poor competitors. Competitiveness was strongly correlated to early light interception. Surprisingly, doubling the recommended sowing density did not increase weed suppressive ability. Although a poor competitor in the fall, after incorporation in spring, lucerne had the strongest inhibitory effect on seedling establishment, followed by winter oilseed rape and white lupin. Winter rye and fodder radish did not affect seedling establishment, whereas Italian ryegrass was not evaluated because of re‐growth after incorporation. Competition in autumn and subsequent residue‐mediated suppression of weed establishment in spring varied among the cover crop species, with winter oilseed rape offering relatively strong effects during both periods.     

Incorporation of weed spatial variability into the weed control decision-making process

Floral surveys were carried out on a field of 28 m × 100 m on the nodes of a regular 2 m × 2 m grid, using a rectangular sampling area of 25 cm × 30 cm. In total, 765 units were sampled, each one characterized by the spatial co-ordinates and the number of seedlings of different weed species. The spatial representation of the weeds was obtained with kriging. Simulations were carried out for Amaranthus spp., which had the highest frequency and density (221 plants m^?2), and Portulaca oleracea L., a species that combined a more aggregated distribution with a medium–high density (27 plants m^?2). The results obtained clearly indicated that the usefulness of geostatistical procedures depends on the type of question posed by the user. If the goal is to estimate weed density and, consequently, crop yield loss, kriging appears to overburden the decision-making process, without improving the estimates obtained. This procedure becomes useful for obtaining weed infestation maps to be used for intermittent spraying applications. The reliability of these maps increases with the number of samples used for kriging. With the more aggregated species, at least 50 samples are required to obtain an infestation map. The reduction in the area to be treated depends on the threshold level adopted and on the number of samples used for kriging. With a threshold around the break-even point for most post-emergence treatments, this reduction varies from 10% to 40% with maps obtained from 50 and 175 samples respectively. The usefulness of infestation maps obtained with kriging for improving the decision-making process is strictly dependent on the weed patch dynamics: if these patches remain relatively stable over time, kriging can be carried out periodically without overburdening the decision-making process, whereas, if they are not stable, maps need to be drawn up each year, with a significant increase in costs.     

Development of chemical weed control and integrated weed management in China

More than 200 species of weeds are infesting main crop fields in China, among which approximately 30 species are major weeds causing great crop yield losses. About 35.8 million hectares of crop fields are heavily infested by weeds and the annual reduction of crop yields is 12.3–16.5% (weighted average). Along with rural economic development, approximately 50% of the main crop fields undergo herbicide application. Chemical weed control has changed cultural practices to save weeding labor in rice, wheat, maize, soybeans and cotton. At the same time, continuous use of the same herbicides has caused weed shift problems and weed resistance to herbicides. Consequently, integrated weed management in main crops is being developed.     

Measurement of weed species diversity in crop/weed competition studies

Data on dry matter accumulation by weeds in the presence or absence of two densities of vining peas was used to compare six diversity indices as measures of changes in the structure and behaviour of the weed flora. Analysis of variance was carried out for each index and the residuals were examined for skewness, kurtosis and heteroscedasticity. Indices based on species number (richness) and species evenness were found to be effective and reliable over the wide range of data examined and permitted new information to be derived. They showed that while increasing competition from the pea crops suppressed weed growth and reduced the number of species recorded at successive sampling dates, there was no selection pressure and species evenness was not reduced.     

Modelling herbicide dose and weed density effects on crop:weed competition

The effects of a range of herbicide doses on crop:weed competition were investigated by measuring crop yield and weed seed production. Weed competitivity of wheat was greater in cv. Spark than in cv. Avalon, and decreased with increasing herbicide dose, being well described by the standard dose–response curve. A combined model was then developed by incorporating the standard dose–response curve into the rectangular hyperbola competition model to describe the effects of plant density of a model weed, Brassica napus L., and a herbicide, metsulfuron‐methyl, on crop yield and weed seed production. The model developed in this study was used to describe crop yield and weed seed production, and to estimate the herbicide dose required to restrict crop yield loss caused by weeds and weed seed production to an acceptable level. At the acceptable yield loss of 5% and the weed density of 200 B. napus plants m^–2, the model recommends 0.9 g a.i. metsulfuron‐methyl ha^–1 in Avalon and 2.0 g a.i. in Spark.     

Weeds —influences of weed vegetation in ipm and non-chemical weed control

Weed vegetation can have both detrimental and beneficial effects on populations of aphids and their natural enemies. Examples are given of these effects in relation to aphid and virus problems of vegetable crops. Important factors in these relationships are discussed and possible lines of future research are indicated.     

Site-specific weed control technologies

Site-specific weed control technologies are defined as machinery or equipment embedded with technologies that detect weeds growing in a crop and, taking into account predefined factors such as economics, take action to maximise the chances of successfully controlling them. In this study, we describe the basic parts of site-specific weed control technologies, comprising weed sensing systems, weed management models and precision weed control implements. A review of state-of-the-art technologies shows that several weed sensing systems and precision implements have been developed over the last two decades, although barriers prevent their breakthrough. Most important among these is the lack of a truly robust weed recognition method, owing to mutual shading among plants and limitations in the capacity of highly accurate spraying and weeding apparatus. Another barrier is the lack of knowledge about the economic and environmental potential for increasing the resolution of weed control. The integration of site-specific information on weed distribution, weed species composition and density and the effect on crop yield, is decisive for successful site-specific weed management.     

Predicting the competitive effects of weed and crop density on weed biomass, weed seed production and crop yield in wheat

Competition between winter-sown wheat and Viola arvensis Murray or Papaver rhoeas L. was studied in two experiments in two successive years. The effects of varying crop and weed density were modelled in terms of weed biomass over time, weed seed production and crop yield. Biomass model parameters, representing maximum weed biomass and intra- and interspecific competition, were obtained for different assessment dates, enabling biomass levels to be predicted during the two growing seasons. Weed biomass declined, and its maximum level was reached earlier, with increasing crop density. Intraspecific competition was higher in the absence than in the presence of crop, increasing with time and with weed density. Halving the wheat population increased June biomass of V. arvensis by 74% and of P. rhoeas by 63%. Crop yield losses with increasing weed density were greater with low than with medium and high crop populations. P. rhoeas was significantly more competitive than V. arvensis in both years. Weed biomass in 1989 responded more to reductions in crop density following the milder winter of 1988/89 than in the previous year; however crop yields were less affected in 1989 due to summer drought, restricting late weed growth and competition. Weed seed production was related to weed biomass; the progressive lowering of crop density increased seed production, and both species were very prolific in the absence of crop. By combining models, seed production could be derived for a given competitive effect on the crop. Threshold weed populations, based on low weed levels that are not economic to control, could then be equated with the accompanying weed seed production.     

Modelling the correlation between plant phenology and weed emergence for improving weed control

The efficiency of weed control practices could be improved if their timing is linked to weed emergence dynamics. A study was conducted in a pre‐alpine valley in northern Italy to evaluate whether the phenological phases of some perennial plants could serve as reliable indicators of time of weed emergence and thus be an alternative to bioclimatic models for supporting management decisions. Weed emergence was observed from 2003 to 2004 in five sites at different altitudes. The emergence dynamics of the main weeds were modelled with a Gompertz model. The phenology of 10 common shrubs and a tree was monitored by visual assessment. The flowering and fruit‐ripening phases of the most useful shrubs were modelled with a lognormal model. Correlation analysis between the two functions was used to study the correspondence between plant phenology and weed emergence. Flowering and fruit‐ripening phases of the shrubs were well described by the lognormal model. The correlation analysis between the lognormal model and Gompertz model showed correspondences between the phenological phases and emergence dynamics of the main weeds. The proposed method can be used to examine shrub phenology–weed emergence correlations and consequently for supporting weed management, under certain conditions.     

Effects of weed harrowing and undersown clover on weed growth and spring cereal yield

Weed competition and nutrient scarcity often restrict organic cereal production, especially where the availability of livestock manure is limited. While harrowing of annual weeds and legume cover crops can be used, these methods are both executed in early spring and may hinder each other. Two cycles of a 2‐year crop rotation were carried out in south‐east Norway (60°42′N, 10°51′E, altitude 250 m) with weed harrowing and undersown cover crops (WHCC) at two fertiliser rates (40 and 100 kg nitrogen ha^?1). The effect of the WHCC treatments was measured by weed density and species, weed biomass, changes in weed seedbank and grain yield. The weed density depended on the interaction between WHCC, fertiliser and year. On average, pre‐emergence weed harrowing reduced weed density by 32% and weed biomass by 49%, while pre‐ and post‐emergence weed harrowing reduced weed density by 59% and weed biomass by 67% compared with the untreated control. Spergula arvensis became more abundant at low rather than at high fertiliser rates. On average, white clover cover crop sown after pre‐emergence weed harrowing resulted in the highest yields for both oat (+12.1%) and wheat (+16.4%) compared with the untreated control. Despite differences in weed population density and biomass among WHCC treatments within years, the weed biomass, weed density and seedbank increased for all WHCC treatments over the 4‐year period. More research is required into improving the efficacy of mechanical and cultural weed suppression methods that organic systems rely on.     

Chemical manipulation of weed emergence

Field studies were conducted over a 4-year period to determine whether weed emergence could be increased by incorporating germination stimulants into the soil. Sodium azide (4.5–36 kg azide ha^?1), ammonium nitrate (34–67 kg nitrate ha^?1), and butylate (0.2–0.4 kg ai ha^?1) were used. Sodium azide was the most consistent stimulant of indigenous weed emergence over all experiments and seasons, but there was great variability in its effects and the highest application rate sometimes inhibited emergence. Consistent stimulation of broad-leaf weed emergence occurred following September applications of 4–5 kg azide ha^?1 of sodium azide. Sodium azide at 4.5 kg azide ha^?1 increased grass plus broad-leaf weed emergence 43% based on summer applications in three growing seasons. No general recommendation for stimulating the emergence of weeds can be made because stimulation was a function of rate of application, season of the year, and post-treatment environment. Influence de traitments chimiques sur la levée des mauvaises herbes Des études portant sur une durée de quatre ans ont été conduites afin de déterminer si l'incorporation dans le sol de stimulants de germination pouvait augmenter la levée des mauvaises herbes. Les produits essayés ont été le nitrure de sodium (4, 5–36 kg de nitrure ha^?1), le nitrate d'ammonium (34–67 kg nitrate ha^?1), et le butylate (0, 2–0, 4 kg ma ha^?1). C'est avec le nitrure de sodium que la stimulation de la levée des graines en place a été observée le plus régulièrement, au cours de plusieurs expériences effectuées en différentes saisons; cependant, une grande variabilité de l'effet de ce produit a été notée, et les doses d'application les plus élevées ont parfois inhibé les levées. Appliqué en septembre à la dose de 4, 5 kg nitrure ha^?1, le nitrure de sodium stimule d'une manière régulière la levée des dicotylédones. Des traitements d'été opérés durant trois années ont montré qu'à cette dose, il augmente la levée des mono et des dicotylédones de 43%. Il n'est cependant pas possible de donner des recommandations générales pour la stimulation des levées de mauvaises herbes, car cette stimulation dépend de la dose appliquée, de la saison pendant laquelle elle est opérée, et des conditions qui régnent après le traitement. Chemische Beeinflussung des Auflaufens von Unkräutern In einer über vier Jahre laufenden Feldstudie wurde untersucht, ob durch das Einarbeiten von Keimungsstimulatoren das Auflaufen von Unkräutern beschleunigt werden kann. Zu die-sem Zweck wurden Natriumazid (4, 5–36 kg azid ha^?1), Ammoniumnitrat (34–67 kg nitrat ha^?1) und Butylat (0, 2–0, 4 kg ai ha^?1) verwendet. Natriumazid war in allen Versuchen das am regelmässigsten wirksame Auflaufstimulans für die an den verschiedenen Standorten vorkommenden Unkräuter, wobei die höchsten Applikationsraten hin und wieder das Auflaufen hemmten. Applikationen von 4, 5 kg azid ha^?1 Natriumazid im September erhöhten die Auflaufrate regelmässig; sie wurde in drei Vegetations-perioden bei grasartigen und breitblättrigen Unkräutern um 43%, vergleichen mit den Sommerapplikationen, gesteigert. Es kann keine all-gemein gültige Anwendungsempfehlung gegeben werden, da der Stimulationseffekt stark von Dosierung, Jahreszeit und den Umweltsbedingungen nach der Applikation abhängt.     

Using plant population biology in weed research: a strategy to improve weed management

The aim of this paper is to show the need for developing plant population biology studies in order to improve weed management when traditional approaches in weed science have failed, as shown by recent reviews on herbicide resistance. It is also suggested that the usual weed definitions do not reflect the new aims of weed ecology. The results of studies on the genetic, demographic and spatial variability of weed populations and their regulation by pests and pathogens are reported, and are discussed in the specific case of clonal perennial species.     

Critical periods for weed competition

Abstract

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THE SAFETY OF THE HERBICIDES 2,4-D AND 2,4.5-T. D. J. Turner. Her Majesty's Stationery Office, London, 1977. ISBN 0 11 710149 4. Pp. 56. Price £1.20

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Modelling the effects of herbicide dose and weed density on rice‐weed competition

The effects of herbicide dose on rice‐weed competition were investigated to develop a combined model, which can be utilised to estimate an optimum herbicide dose for a given weed density in paddy rice cultivation. Field studies were conducted in Suwon for rice‐Echinochloa crus‐galli competition and Iksan for rice‐Eleocharis kuroguwai during 2007. The competitive effect of the weeds E. crus‐galli and E. kuroguwai decreased with increasing doses of flucetosulfuron and azimsulfuron, respectively, in the same manner as the standard dose–response curve. The combination of the rectangular hyperbolic model and the standard dose–response curve adequately described the complex effects of herbicide dose and weed competition on rice yield. Parameter estimates were used with the model to predict rice yield and estimate the doses of flucetosulfuron and azimsulfuron required to restrict rice yield loss caused by E. crus‐galli and E. kuroguwai, respectively, to an acceptable level. For a rice yield of 5.0 t ha^?1, the model recommended flucetosulfuron doses of 8.7, 13.4 and 20.1 g a.i. ha^?1 when infested with E. crus‐galli at 12, 24 and 48 plants m^?2 respectively. For a rice yield of 5.2 t ha^?1, the model recommended azimsulfuron doses of 3.9, 7.5 and 12.6 g a.i. ha^?1 when infested with E. kuroguwai at 24, 48 and 96 plants m^?2 respectively. The theoretical outputs of the combined model appear robust and indicate there are opportunities for reduced herbicide use in the field. These now require evaluation under field conditions.     

Multivariate analysis in weed science

Weed species in arable ecosystems are adapted to the conditions produced by agricultural management. In order to give preventive and curative guidelines to weed management one needs to predict the composition of the weed vegetation after certain (soil) cultivation measures. Dormancy mechanisms of seeds in the seedbank and the environmental conditions in the furrow before and after cultivation are usually such that the vegetation composition is heterogeneous. Multivariate analyses enable us to describe and classify weed vegetation and to analyse its relationship to environmental factors.