Productivity of beef cattle grazing Brachiaria brizantha cv. Marandu with and without nitrogen fertilizer application or mixed pastures with the legume Desmodium ovalifolium

The use of forage legumes to contribute biologically fixed nitrogen (N) to pastures is an alternative to increase beef cattle production in tropical regions. The objective was to compare the impact of the introduction of a legume with that of N fertilizer application on forage and animal production in Brachiaria pastures. This two-year study assessed three pasture treatments: (1) mixed Marandu palisadegrass (Brachiaria brizantha [syn. Urochloa brizantha] cv. Marandu) and the legume “ovalifolium” (Desmodium ovalifolium) cv. Itabela (Mixed), (2) Marandu palisadegrass pastures with 150 kg N ha⁻¹ (Fertilized), and (3) Marandu palisadegrass without N fertilizer (Unfertilized). Rotational stocking with a variable stocking rate was used with a target herbage allowance of 1.0 kg forage kg body weight⁻¹. The pre-grazing green herbage mass was similar for Fertilized and Mixed pastures, with 54% and 63% more mass than Unfertilized pasture, respectively (p < .001). Cattle that grazed the fertilized pasture had the greatest average daily gain (ADG; p = .017). The stocking rate and liveweight gain per area were greatest for the Fertilized and Mixed pastures (p < .001 and p < .001, respectively). No differences between treatments were found for DM forage intake (p = .555). Organic matter digestibility was lowest (p < .001) for the Mixed pasture. The inclusion of the ovalifolium legume in the Marandu pasture had the same impact on beef cattle production as annual fertilization with 150 kg N ha⁻¹. The potential and environmental benefits of ovalifolium are discussed.     

Production of beef cattle grazing on Brachiaria brizantha (Marandu grass)—Arachis pintoi (forage peanut cv. Belomonte) mixtures exceeded that on grass monocultures fertilized with 120 kg N/ha

Mixed grass/legume pastures are an alternative to grass monocultures for increased beef cattle production in tropical climates. The objective of this study was to evaluate the productivity of beef cattle grazing either a mixed pasture of Brachiaria brizantha cv. Marandu grass and Arachis pintoi (forage peanut) cv. Belomonte or a Marandu monoculture, under rotational stocking. Five trials were conducted over a period of nine years in north-eastern Brazil where the sward structure (forage, grass and legume mass) and animal performance were compared for a mixed Marandu grass/forage peanut pasture, and a Marandu grass monoculture with 120 kg N ha⁻¹ y⁻¹. Stocking rate was adjusted to maintain forage allowance at 4% body weight/day. A block design was used with four replicates, and warm and cool seasons within each trial were considered, using repeated measurements over time. In the warm season, the forage mass in the mixed pastures was 17% greater than in the monoculture (p = .049), and the stocking rate, average daily gain and liveweight gain per ha were 16.4%, 20.0% and 28.7% greater (p = .004, p < .001 and p < .001 respectively). The average daily gain showed a positive linear relationship with the legume proportion in the sward (p < .001). The mixed forage peanut/Marandu pasture sustained significantly greater beef cattle production (789 kg ha⁻¹ y⁻¹) compared to the N-fertilized grass monoculture (655 kg ha⁻¹ y⁻¹). Appropriately managed, mixed pastures of forage peanut/Brachiaria pastures are sustainable and have high potential for use in the humid tropics.     

Grazing legumes in Europe: a review of their status, management, benefits, research needs and future prospects

In many parts of Europe there has been a net decline in the use of forage legumes since the 1980s, despite the reputed value of legumes for low‐input livestock production systems. The political environment within which livestock farming in much of Europe operates (Common Agricultural Policy) is shifting the balance of economic advantage towards legumes and away from high usage of inorganic fertilizer. This has already been found for legume and grass–legume silages when compared with grass silages with a potential economic gain for farmers averaging 137 € ha^?1, corresponding to an annual benefit for the European livestock farming sector of as much as € 1300 million. Recent literature has shown that legume‐based grazing systems have the ability to reduce environmental problems by increasing the efficiency of N use and by avoiding a high transient surplus of soil mineral N. From the perspective of livestock nutrition, when forage legumes contain moderate levels of secondary compounds, such as condensed tannins and flavonoids, they offer considerable advantages including increased efficiency of N utilization within the digestive tract, reduced incidence of bloat hazard and higher resilience to parasites. Nevertheless, these benefits are partially counterbalanced in both temperate and Mediterranean regions by difficulties in establishment, maintenance and management under grazing. To gain knowledge on mixed grass–legume pastures, further research is required on: (i) the development of sustainable systems of livestock production which can maintain sward persistence and agricultural production under environmental stress; (ii) increasing knowledge of soil–plant–animal relations for a wide range of leguminous species, and under different soil types and climatic situations; and (iii) the benefits for consumers of food produced from low‐input livestock production systems.     

Two perennial legumes (Astragalus adsurgens Pall. and Lespedeza davurica S.) adapted to semiarid environments are not as productive as lucerne (Medicago sativa L.), but use less water

Perennial forage legumes, particularly lucerne (Medicago sativa L.), play a significant role in crop/livestock mixed farming systems in the semiarid region of the Loess Plateau of China as stock feed and a source of nitrogen for subsequent crops. However, there is evidence that lucerne reduces soil water deep in the soil profile, thereby reducing subsequent crop productivity. From 2004 to 2010, this study evaluated the forage productivity and water use of two locally adapted perennial legume species, milk vetch (Astragalus adsurgens Pall.) and bush clover (Lespedeza davurica S.), compared with lucerne. The 7‐year total and average annual forage yield of milk vetch were 56 and 8 t ha^?1 and bush clover was 42 and 6 t ha^?1, respectively, significantly lower than lucerne at 91 and 13 t ha^?1. However, despite lower water‐use efficiencies (16 and 12 kg ha^?1 mm^?1 for milk vetch and bush clover, respectively, compared to 22 kg ha^?1 mm^?1 for lucerne), the total 7‐year water use in milk vetch and bush clover was 3500 mm and 3490 mm, respectively, which was 135–140 mm less than lucerne. After 7 years, lucerne had extracted water from the upper 5 m soil, whereas bush clover used water mainly from the upper 2 m of the soil profile and milk vetch still had some water available below 3 m. We conclude that while the locally adapted forage legumes were not as productive as lucerne as a source of fodder in mixed cropping/livestock system in this region, they use less water, which may be advantageous in drier regions.     

Increasing the population of forage peanut in a mixed pasture by controlling the canopy height

The aim of this study was to determine a temporary strategy for increasing the legume population in an established pasture of brachiaria grass (Brachiaria brizantha) and forage peanut (Arachis pintoi). The treatments comprised four previous long‐term canopy heights (10, 20, 30 and 40 cm) from the establishment (the first 32 months after treatment initiation). From September 2014, all of the experimental units were kept at 10 cm height. The pastures were evaluated for their forage harvest, forage mass, botanical composition and stolon and tiller density. The tiller density and brachiaria grass mass were maintained in all pastures since the first season of evaluation (p > 0.10). The stolon density and legume mass were initially lesser at 30 and 40 cm than those at 10 and 20 cm, early in the experiment, but progressively increased over time. After 1 year under 10 cm height, the population and mass of legume were similar in all treatments. Therefore, reducing the canopy height to 10 cm at the beginning of the rainy season is an efficient method to increase the botanical composition of forage peanut in mixed pastures with brachiaria grass.     

Sustainable,low‐input,warm‐season,grass–legume grassland mixtures: mission (nearly) impossible?

Grazing lands in warm‐temperate and subtropical North America have become less diverse. Pastures are typically grass monocultures, while rangelands are generally managed for the grass components. Overstocking, selective herbicides, fire exclusion and heavy rates of nitrogen fertilizer have contributed to near exclusion of native, warm‐season legumes. The simplicity of managing grass monocultures, pasture production responses to nitrogen fertilizer and profitability of grass‐only systems have limited interest in legume‐based approaches. Changing economics and ecological concerns with ecosystem accumulation of industrial inputs contribute to an increasing interest in legumes. Unlike the development of temperate pasture legumes and recent research in the tropics, legumes tolerant of both freezing temperatures and hot weather have received less attention. Poor establishment, limited persistence and potential invasiveness limit currently available introduced species. Native, herbaceous, warm‐season legume species occur throughout warm‐temperate North America, but little attention has been directed to these plants as potential forage species. Some success with a few native legume species, primarily in the genus Desmanthus, suggests potential for expanded assessment of forage value of the many species available. Current assessments of native legumes, primarily for conservation purposes, provide an opportunity to expand evaluations of these species for pasture and rangeland potential while economics of livestock production and public interest in ecosystem health are supportive. Experiences with legumes of warm‐temperate origin in North America, along with results with temperate and tropical pasture legumes globally, provide a starting point for future efforts at incorporating greater legume diversity in pastures and rangelands of subtropical and warm‐temperate regions around the world.     

The role of the legume in the nitrogen cycle of productive and sustainable pastures

The role of the legume in the nitrogen (N) cycle was examined in grazed pastures receiving no N fertilizer of both temperate and tropical regions by simulating the fluxes of N through different processes of the cycle. The amounts of legume-fixed N required to balance the cycle without invoking a drain on soil organic N reserves (i.e. no net N mineralization) was estimated to vary from 38 to 53% of the above-ground herbage N or from 20 to 31% on a dry matter (DM) basis for tropical pasture systems with a range of pasture utilization of 10–40%. At higher pasture utilization levels of 50-70%, more typical of intensively grazed temperate pastures, the N input requirement in the absence of fertilizer N would be 57-67% of the aboveground herbage N or 35-45% DM. An examination of the role of each contributory process of recycling (viz. excreta returns, internal cycling or remobilization from senescing tissues, litter decomposition) suggests that variations in the amounts of internally cycled N would have the greatest impact on the requirement for biologically fixed N at low levels of pasture utilization (10-40%), while at high pasture utilization levels of 70%, variations in the recovery of excreta-N would have a major effect on the requirement for fixed-N to balance the cycle. The amounts of biologically fixed N required to sustain a range of herbage DM yields of 3-22 t DM ha ^?1 yr^?1 would range from 15 to 158 kg N ha^?1 yr^?1 for tropical pastures. For intensively managed temperate pastures producing 6-15 t DM ha^?1 yr^?1 with a N content of 3·5%, a range of fixation of 120-352 kg N ha^?1 yr^?1 is required. These simulations indicate how legume contents of 20-45% of herbage DM could contribute to productive and sustainable (in terms of N) pasture systems of both temperate and tropical regions     

Contribution of Medicago sativa to the productivity and nutritive value of forage in semi‐arid grassland pastures

The inclusion of legumes in semi‐arid native grasslands may promote the productivity and nutritive value of forage. This study was designed to assess the effect of legumes (the introduced legume Medicago sativa or the native legume Dalea purpurea) and soil P fertility (addition of 0, 50, or 200 P₂O₅ kg/ha at seeding) on the dry matter and nutrient content of native grasses mixtures, compared with the commonly used introduced forage grass Bromus biebersteinii grown with M. sativa. Plant harvests were performed in September 2008, July 2009 and September 2009. Plants nutrient content, δ¹⁵N value and dry matter were analysed. Results show that the M. sativa enhanced the N and P concentrations of native grass mixtures early in the summer, as well as the N concentration in Bouteloua gracilis in late summer of the driest year, 2009. The higher AM fungal diversity promoted by M. sativa was positively correlated with the dry matter and nitrogen uptake of M. sativa and with the P concentration of native grasses, in early summer. Overall, this study shows that M. sativa promoted beneficial AM fungal taxa and improved forage production in the semi‐arid prairies.     

Forage chain arrangement for sustainable livestock systems in a Mediterranean area

Mediterranean forage systems suffer from limited availability of fresh forage because of water deficits and extreme temperatures. Consequently, fresh forage is unavailable for at least 6–7 months a year, and farmers must buy feed to support livestock production. With the aim of overcoming these limitations, a 2‐year trial was conducted on three distinct sites in Sicily (at 10, 600 and 1200 m elevation) with thirty‐four varieties of forage species belonging to nine biennial/perennial and thirteen annual species. Results showed that by integrating grasses and legumes, species from environments with different climatic conditions enable the season of forage production to be extended from mid‐April to mid‐November. Quality traits of forage in different areas varied in relation to species and varieties. In general, the sown‐forage quality was better than in pastures and fallows in the same areas commonly used to feed animals. This also leads to a reduction in the use of supplementary feeds. Among the tested species, Lolium multiflorum and Medicago sativa emerged as the most promising for filling the forage‐deficit periods, and Trifolium spp. and Vicia sativa were found to be superior for increasing forage quality. The results are discussed in the context of adapting Mediterranean forage supplies for ensuring greater sustainability of livestock production in mountain, hill and plain areas. The proposed forage chain arrangement represents part of local potential adaptation to climate limitations and climate change.     

Optimizing the growth of forage and grain legumes on low pH soils through the application of superior Rhizobium leguminosarum biovar viciae strains

Climate variability and current farming practices have led to declining soil fertility and pH, with a heavy reliance on fertilizers and herbicides. The addition of forage and grain legumes to farming systems not only improves soil health but also increases farm profitability through nitrogen (N) fertilizer cost offsets. However, the formation of effective symbioses between legumes and rhizobia can be unreliable and is considered at risk when combined with dry sowing practices such as those that have been designed to obviate effects of climate change. This research was initiated to improve the robustness of the legume/rhizobia symbiosis in low pH, infertile and dry soils. Production from two cultivars of field pea (Pisum sativum) and two species of vetch (Vicia spp.), and symbiotic outcomes when inoculated with a range of experimental rhizobial strains (Rhizobium leguminosarum biovar viciae), was assessed in broad acre field trials which simulated farmer practice. New rhizobia strains increased nodulation, N fixation, produced more biomass and higher seed yield than comparator commercial strains. Strain WSM4643 also demonstrated superior survival when desiccated compared to current commercial strains in the laboratory and on seed when delivered as inoculant in peat carriers. WSM4643 is a suitable prospect for a commercial inoculant in Australia and other agricultural areas of the world where growing peas and vetch on soils generally considered problematic for this legume/rhizobia symbiosis. A particular advantage of WSM4643 may be that it potentiates sowing inoculated legumes into dry soil, which is a contemporary response by farmers to climate variation.     

Annual forage legumes in dryland agricultural systems of the West Asia and North Africa Regions: research achievements and future perspective

Forage legumes are vitally important to animal production in the dryland farming systems of the Mediterranean region. Of the diverse forage‐legume species adapted to the Mediterranean climate, vetches, (Vicia spp.), chicklings (Lathyrus spp.), annual medics (Medicago spp), clovers (Trifolium spp.) and species of the Lupinus, Lotus, Onobrychis, Hedysarum and Ornithopus genera are considered to be the most agronomically important and economically valuable species for the region. Adoption of perennial self‐regenerating medic (Medicago spp.) has been limited because of technical difficulties, but annual vetch (Vicia spp.) has the greatest potential as a viable animal‐feed source and a rotation crop with cereals. Some forage legumes survive harsh conditions by their unique underground growth habit, for example, V. amphicarpa and Lathyrus ciliolatus. Efforts to improve forage legumes have been based on both management/cultural factors and breeding. Research based on several long‐term barley‐ and wheat‐based rotation trials has demonstrated the viability of forage legumes, especially vetch, in the region's improved farming system. An additional benefit to such legumes is the enhancement of soil quality, that is, soil fertility, soil organic matter and soil physical properties. Thus, the development of forage legumes is essential to agricultural sustainability in the Mediterranean region and in other dryland cereal‐growing areas of the world where grazing livestock is a dominant enterprise. To build upon the considerable research conducted on forages, intensified efforts are needed to develop locally adapted forage cultivars, to provide economic assessment of forages in cropping systems and to promote technology transfer at the farm and community level.     

Cattle select against the invasive grass tall fescue in heterogeneous pastures managed with prescribed fire

Tall fescue (Festuca arundinacea) is a Eurasian forage grass extensively planted in the United States. However, an endophytic fungus in tall fescue, Epichloë coenophiala, causes health problems in cattle. We predicted that cattle prefer to graze alternative forages when available. We also predicted that cattle use tall fescue more intensively in recently burned areas, as fire can increase forage quality. We tested these predictions in four diverse‐forage pastures in Iowa, comparing use by cattle of tall fescue and four alternative forages (non‐fescue cool‐season grasses, native warm‐season grasses, non‐leguminous forbs and legumes) to their availabilities at the pasture scale. We also examined how tall fescue influences the distribution of grazing at a fine scale (0.1‐m² quadrats). Tall fescue was the most abundant forage (46% of plants), but composed only 26% of grazed vegetation. In contrast, legumes composed 12% of available forage but 25% of grazed vegetation. Other forages were used in proportion to availability. At a fine scale, total grazing frequency (proportion of plants grazed) was lower in quadrats containing abundant tall fescue, and higher in quadrats with abundant warm‐season grasses. Grazing frequency of tall fescue and other cool‐season grasses was greatest in recently burned quadrats, but total grazing frequency did not increase after burning. Our results show that although cattle graze tall fescue, particularly following burns, they limit their use of this grass. Given that tall fescue is underused, creates health risks for cattle, and degrades wildlife habitat quality, it may be advisable to reduce tall fescue in pastures.     

Liveweight gain of beef cattle in Brachiaria brizantha pastures and mixtures with Stylosanthes guianensis in the Brazilian savannah

The use of stylosanthes in mixed grass-legume pastures may minimize the decline in forage quality and quantity that occurs in monoculture grass pastures, even though the availability of commercial cultivars in Brazil is still limited. The objective of the study was to evaluate the liveweight (LW) gain of young Nellore bulls in a mixed pasture of Brachiaria brizantha (cv. Paiaguás) with the latest release Stylosanthes guianensis cv. Bela. The study was conducted in Planaltina, FD, Brazil, from September to August in two consecutive years, right after seeding. The experimental design was a randomized complete block design with two treatments (mixed or monoculture Paiaguás pastures) and three replicates. The average daily gain (ADG) was on average greater in mixed pastures (0.436 vs. 0.350 kg head⁻¹ day⁻¹ in mixed and grass pastures respectively), particularly in the dry period (0.344 vs. 0.183 kg head⁻¹ day⁻¹). The benefit of mixed over monoculture grass pastures throughout the year was 22 kg LW/head and 55 kg LW/ha. The presence of stylosanthes increased the crude protein (CP) concentration in mixed pastures (120 g/kg) when compared to monoculture grass pastures (109 g/kg), probably influencing the ADG of bulls. The increase of liveweight gain, predominantly in the dry season, makes the high-protein stylosanthes cv. Bela an alternative to buffer the seasonal deficit of forage quality in newly seeded grass pastures.     

“Summer sowing”: A successful innovation to increase the adoption of key species of annual forage legumes for agriculture in Mediterranean and temperate environments

This paper reports on the evaluation of “summer sowing,” an innovative approach to increase the adoption of recently domesticated species of hard seeded annual legumes in Mediterranean and temperate Agriculture. The research revealed that several species of annual legumes whose seed can be readily harvested on‐farm and which possess natural hard seed dormancy, may be sown into dry soil in late summer without additional processing. These studies proved that the hard seed dormancy was broken down sufficiently in the soil over 4–6 weeks to produce robust legume pastures with more than 150 seedlings per m² following the first winter rains, in replicated field sites established across wide agro‐ecological zones in Western Australia (WA) and New South Wales (NSW). Ornithopus sativus Brot., O. compressus L. and Trifolium spumosum L. were suitable for summer sowing based on both hard seed breakdown patterns and subsequent seedling survival in WA. While in NSW, in addition to these legumes, Biserrula pelecinus L., T. vesiculosum Savi. and T. glanduliferum Boiss. were also suitable for summer sowing. A 1.5‐ to 10‐fold increase in herbage production was achieved relative to conventionally sown T. subterraneum L. This development represents a step change in forage legume development for renovated pastures in these environments. Importantly, the experiments revealed differences in G x E effects on seedling establishment, total herbage production and seed yield in different climatic zones. The summer sowing approach is presented as a revolutionary method for pasture renovation that overcomes significant barriers to adoption.     

Pasture production,persistence of legumes and lamb growth in summer‐dry hill pastures

Pasture legumes that persist under challenging agroecological conditions are crucial to ensure high lamb growth rates in dryland pastures. Pasture and lamb production from binary and diverse mixtures (Mix) of tall fescue (TF) with white clover (Whc), balansa clover (Bc), subterranean clover (Sc) and birdsfoot trefoil (Bft) were compared in a summer‐dry hill site in Corvallis, Oregon over a two‐year period. In 2018, all pasture combinations provided similar lamb liveweight gains (LWG, mean 177 g/day) in the first half of spring. Lambs in TF‐Bc and TF‐Mix pastures grew 31 to 41 g/day faster than those grazing TF‐Sc, TF‐Whc and TF‐Bft in the second half of spring (p < .05). Overall, TF‐Bc and TF‐Mix had higher (p < .05) legume contents (32% and 37% respectively) compared to other pasture combinations. In spring 2019, lambs that grazed the TF‐Mix and TF‐Whc pastures had higher LWG than those on other pastures (p < .05). The superior lamb growth rates were associated with the higher legume content and pasture quality maintained into the late spring period. Overall, the legume content of all pastures decreased over the course of the two‐year trial, with the decline being substantial for balansa clover. The present study confirmed that a high legume content of pastures leads to greater lamb growth rates. Total annual yields of pastures that had greater legume contents were superior to others (p < .05). Thus, a combination of self‐regenerating annual clovers with perennial legumes in pasture mixtures may ensure a higher legume content and longer persistence in dryland hill pastures.     

Evaluation of forage legumes for introduction into natural pastures of semi‐arid rangelands of Kenya

To address the potential of legumes to contribute to improved quality and quantity of natural pastures in the semi‐arid rangelands of Kenya, five legume species were introduced and evaluated in a small‐plot field experiment over three growing seasons. The investigated species were glycine (Neonotonia wightii), siratro (Macroptilium atropurpureum), dolichos (Lablab purpureus cv. Rongai), velvet bean (Mucuna pruriens) and shrubby stylo (Stylosanthes scabra cv. seca). Treatments included two cutting heights (ground level and 15 cm) and two cutting intervals (at 2 and 4 months). The mean dry matter (DM) yields of glycine and siratro were highest when the legumes were harvested at ground level at 2‐month intervals (10·31 and 7·81 t ha^?1 year^?1 respectively). Mean DM yield of stylo was highest when the legume was harvested at 15 cm after 4 months (3·52 t ha^?1 year^?1). These three legumes also produced high organic matter through litter fall, which contributed to soil fertility. Evidence from a supporting pot experiment showed effective nodulation and potential for N fixation. These legumes also possessed deep tap roots and withstood heavy defoliation. These three legumes were selected for further integration with grasses in natural pastures. The DM yields of dolichos and velvet bean when harvested at 15 cm after 2 or 4 months were low (2·48 and 1·91 t ha^?1 year^?1), and these species were considered inappropriate for further investigation.     

Phenotyping of Urochloa humidicola grass hybrids for agronomic and environmental performance in the Piedmont region of the Orinoquian savannas of Colombia

In the low fertility acid soils of the Orinoquian savannas of Colombia, Urochloa humidicola cv. Tully or Humidicola is one of the most widely planted tropical forage grasses for improving livestock productivity. Low nutritional quality of this grass limits sustainable livestock production in this region. In this study, we conducted a phenotypic evaluation under field and greenhouse conditions of one of the first hybrid populations of U. humidicola generated from the forage breeding program of CIAT. Our objective was to identify a set of new hybrids of U. humidicola that combine improved productivity and nutritional quality plus the biological nitrification inhibition (BNI) trait/ability to reduce nitrogen (N) losses via leaching and nitrous oxide (N₂O) emissions. To this end, we tested 118 hybrids (planted in pots) in the greenhouse for over 6 months and measured potential nitrification rates (NR) using soil microcosm incubation. NR values observed ranged from 0.27 to 5.75 mg N-NO₃⁻ kg soil⁻¹ day⁻¹. Later, 12 hybrids with different levels of NR were selected and field-tested in the Orinoquia region over a 4 years period (2013–2017) for dry matter production, nutrition quality (crude protein, in vitro digestibility and fibres content) and NR in each year. In the rainy season of 2018, two hybrids with superior agronomic performance and contrasting field level NR (Uh08/1149 and 0450) were subjected to analysis of soil-borne N₂O emissions after fertilization during 13 days. The NR values recorded were not directly correlated with the forage quality parameters evaluated, however, the two grasses with the lowest NR values were among those with the highest biomass production, crude protein content, and N uptake. The grass hybrid Uh08/1149 and the germplasm accession CIAT 16888 were found as materials with superior forage value, with production of 14.1 and 14.6 tons dry matter ha⁻¹ year⁻¹ (up to 8% higher than the cv. Tully), crude protein of 11.5 and 9.1% per cut (up to 20% higher than the cv. Tully), and N uptake of 31.6 and 25.7 kg N ha⁻¹ cut⁻¹ (up to 30% higher than the cv. Tully). Additionally, these two grasses are likely to exhibit high-BNI ability, with potential to improve N use efficiency in managed pastures.     

Nodulation and root growth of forage legumes sown into tall fescue swards

Establishing forage legumes into endophyte‐infected tall fescue (Festuca arundinacae Schreb.) pastures is problematic, especially in well‐established stands. A oversowing field experiment determined if this problem was because of poor nodulation. Four renovation techniques, clipped sward (treatment A), herbicided + rye seeding in the previous autumn (treatment B), herbicided in the autumn and spring (treatment C) and herbicided to suppress the sward (treatment D), were investigated to determine their effect on nodulation and root growth of lucerne (Medicago sativa L.), red clover (Trifolium pratense L.) and white clover (T. repens L.) at 16, 22 and 29 d after sowing the legumes. A pot experiment was also conducted under optimal growth conditions and using the same soil to determine the nodulation and root growth potentials of these legume species. At adequate rhizobial populations (>6 × 10⁴ cfu g^?1 soil), substantial nodulation of all species occurred by 29 d after sowing in treatments C and D, whereas nodulation of clovers was usually reduced in treatment A. Total root lengths for all sampling dates, species and treatments were severely restricted, especially under treatment A. A general correspondence of nodulation with root growth was observed for all species, with high correlations (r ≥ 0·85) between these variables for all legume species and treatments, suggesting that soil moisture, and possibly competition for light, were the limiting factors. These results demonstrate that weak stands of forage legumes, typically found when sown into tall fescue swards, are probably not because of inadequate nodulation. Rather, inhibition of root growth by detrimental physical/chemical conditions or allocation of limited photosynthate to shoots instead of roots is suggested.     

Ensiled sorghum and soybean as ruminant feed in the tropics,with emphasis on Cuba

The concept is presented of combined cultivation of legumes and sorghum–maize and their use as conserved ruminant feed in tropical regions, with special reference to Cuba. Good yields are obtained during the rainy season through intercropping (alternate rows of either sorghum or maize and soybean). When followed by ensiling, this provides high‐quality ruminant feed for the tropical dry season. Soybean compensates for the low crude protein content of sorghum, whereas sorghum allows good silage quality in combination with legumes. The paper reviews and updates recent studies assessing combined sorghum–soybean cultivation and ensiling as well as determination of their feed value. The high nutritive quality and forage potential when these crops are intercropped demonstrate that silage from these plants can be used successfully in ruminant diets in Cuba and other tropical areas. Perspectives for new studies in this field are suggested, particularly with legume species that are more adapted to specific tropical regions and/or with higher forage yield. In addition, it is suggested that there is a need to assess the supplementation impact on meat and milk production at the farm level, as well as its environmental impact, when ruminants are fed combined silages from whole plants of sorghum–legumes.     

PASTURE DEVELOPMENT IN THE BEEF CATTLE REGIONS OF ARGENTINA

An account is given of the climate, vegetation and native pastures of the beef-cattle regions of Argentina. Though some of these native pastures have a satisfactory carrying capacity, most of them are defective in quality or low in productivity; there is considerable scope for increasing production through the establishment of sown pastures. Trials have been made with a wide range of grasses and legumes, principally of African and European origin. Though the pattern of species to be used in these regions is imperfectly resolved, the authors present a map showing tentative boundaries for temperate, sub-tropical and tropical species. Work to date suggests that pasture development can probably proceed over great areas without the use of fertilizers, and this adds to the attractiveness of pasture programmes. A problem of no less importance than the technical aspects of pasture improvement is that of gaining a greater investment of capital into the development of cattle estancias.