Inverting improves establishment of Pinus contorta and Picea abies— 10‐year results from a site preparation trial in Northern Sweden

The effects on seedling survival and growth of a new scarification method, inverting, were evaluated over 10‐yrs after planting lodgepole pine and Norway spruce on a 2‐yr‐oId clear‐cut in northern Sweden. Inverting, which provides planting spots containing humus turves covered in loose mineral soil without making mounds or ridges, was compared with ploughing, mounding, disc trenching, and no scarification. Subplots with high or low planting positions were used to assess small‐scale topographical effects. For both species, the treatment ranking according to stem volume production after 10‐yrs was inverting > ploughing ≥ mounding = disc trenching > no scarification. Inverting improved seedling height growth by approximately 35% compared with mounding or disc trenching and by more than 100% compared with no scarification. High survival rates were also found following inverting, but only the no‐scarification treatment resulted in a statistically significant reduction in survival rates (ca. 25%) for both species. Further development of the inverting technique might give environmental advantages compared with conventional mechanical site preparation.     

Indications that site preparation increases forest ecosystem carbon stocks in the long term

Mechanical site preparation (MSP) causes a mixing disturbance of the soil, which may increase decomposition of soil organic matter and subsequent carbon (C) dioxide emissions to the atmosphere. MSP also promotes the establishment and growth of tree seedlings, and hence ecosystem C fixation. However, there are uncertainties regarding the net effects of MSP on C stocks at the ecosystem scale. To assess decennial effects of MSP on ecosystem C stocks, C stocks in soil, ground vegetation and trees at three experimental forest sites with Pinus contorta, Pinus sylvestris and Picea abies in Sweden were sampled and measured for ca. 25 years in a control and after three MSP treatments: disc trenching, mounding and ploughing. After 25 years, all of the MSP treatments resulted in larger ecosystem C stocks than the control treatment due to positive effects on the tree biomass C stock. The tree C stock was highest after ploughing, intermediate after mounding or disc trenching and lowest in untreated control plots at all experimental sites. The MSP treatments did not affect the soil C stocks down to 30?cm. We recommend mounding or disc trenching to promote C sequestration as they disturb sites’ ecological, aesthetic and recreational values less than ploughing.     

The influence of soil scarification on the turn‐over rate of slash needles and nutrient release

The effect of soil scarification on decomposition of green Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) needles and mineral nutrient release was investigated. The treatments studied were ploughing, disk trenching, mound on inverted humus and untreated control. The study was conducted on two clear‐felled sites with different ground moisture conditions, located in south Sweden. The needles decomposed and released their contents of N and Ca considerably faster on scarified areas than on untreated ones. For P, Mg and especially K, only minor differences between scarified and untreated areas were established. At the wet site, needle decomposition and nutrient mineralization were generally affected to the same extent by all three scarification methods studied. At the dry to fresh site, mounding and ploughing enhanced needle decomposition and nutrient mineralization more than disk trenching. The effect of different scarification methods on the long‐term production capacity of soils is discussed.     

Resource competition across the crop-tree interface in a maize-silver maple temperate alley cropping stand in Missouri

In order to improve the management of temperate alley cropping, it is important to study the growth and physiological responses of plants arising from competition across the crop-tree interface. Maize (Zea mays L.) was established between rows of seven-year-old silver maple (Acer saccharinum L.) trees in north-central Missouri, USA with four imposed treatments: (1) an unmodified control with a standard rate of N fertilization (179.2 kg N (as NH₄NO₃) ha⁻¹), (2) trenching with root barrier installed, (3) supplemental fertilization treatment (standard N + 89.6 kg ha⁻¹ N), and (4) a combination of trenching with root barrier and supplemental fertilization. Whereas soil N status had little effect on maize physiology and yield at the interface, competition for soil water was substantial in both years. Without a root barrier, soil water content, predawn and midday water potential, and midday net photosynthesis of maize plants adjacent to the tree row were reduced compared with those of plants in the alley center, but no differences across the maize crop were evident in the presence of a barrier. Grain yield of border row maize plants lacking an adjacent barrier was depressed compared with that for maize plants with a root barrier present (8.42 vs. 6.59 Mg ha⁻¹ in 1997; 5.38 vs. 3.91 Mg ha⁻¹ in 1998). However, the barrier did not completely restore yield to that in the alley center, suggesting that reductions in light near the tree row also limited production. Top ear height showed a similar pattern of response to the presence of a root barrier. Silver maple trees responded to root barrier installation with reduced annual diameter growth and reduced water status on some sample days. This revised version was published online in June 2006 with corrections to the Cover Date.     

Above- and belowground biomass,nutrient and carbon stocks contrasting an open-grown and a shaded coffee plantation

Coffee (Coffea canephora var robusta) is grown in Southwestern Togo under shade of native Albizia adianthifolia as a low input cropping system. However, there is no information on carbon and nutrient cycling in these shaded coffee systems. Hence, a study was conducted in a mature coffee plantation in Southwestern Togo to determine carbon and nutrient stocks in shaded versus open-grown coffee systems. Biomass of Albizia trees was predicted by allometry, whereas biomass of coffee bushes was estimated through destructive sampling. Above- and belowground biomass estimates were respectively, 140 Mg ha⁻¹ and 32 Mg ha⁻¹ in the coffee–Albizia association, and 29.7 Mg ha⁻¹ and 18.7 Mg ha⁻¹ in the open-grown system. Albizia trees contributed 87% of total aboveground biomass and 55% of total root biomass in the shaded coffee system. Individual coffee bushes consistently had higher biomass in the open-grown than in the shaded coffee system. Total C stock was 81 Mg ha⁻¹ in the shaded coffee system and only 22.9 Mg ha⁻¹ for coffee grown in the open. Apart from P and Mg, considerable amounts of major nutrients were stored in the shade tree biomass in non-easily recyclable fractions. Plant tissues in the shaded coffee system had higher N concentration, suggesting possible N fixation. Given the potential for competition between the shade trees and coffee for nutrients, particularly in low soil fertility conditions, it is suggested that the shade trees be periodically pruned in order to increase organic matter addition and nutrient return to the soil. An erratum to this article can be found at     

Nutrient stocks of short-term fallows on a high base status soil in the humid tropics of Papua New Guinea

In order to understand nutrient dynamics in tropical farming systems with fallows, it is necessary to assess changes in nutrient stocks in plants, litter and soils. Nutrient stocks (soil, above ground biomass, litter) were assessed of one-year old fallows with Piper aduncum, Gliricidia sepium and Imperata cylindrica in the humid lowlands of Papua New Guinea. The experiment was conducted on a high base status soil (Typic Eutropepts), and in Papua New Guinea such soils are intensively used for agriculture. Soil samples were taken prior to fallow establisment and after one year when the fallows were slashed and above ground biomass and nutrients measured. The above ground and litter biomass of piper was 13.7 Mg dry matter ha^-1, compared to 23.3 Mg ha^-1 of gliricidia and 14.9 Mg ha^-1 of imperata. Gliricidia produced almost 7 Mg ha^-1 wood. Total above ground biomass returned to the soil when the fallows were slashed was the same for piper and gliricidia (8 Mg ha^-1). Gliricidia accumulated the largest amounts of all major nutrients except for K, which was highest in the above ground piper biomass. Imperata biomass contained the lowest amount of nutrients. The largest stocks of C, N, Ca and Mg were found in the soil, whereas the majority of P was found in the above ground biomass and litter. Almost half of the total K stock of piper and gliricidia was in the biomass. During the fallow period, soil organic C significantly increased under gliricidia fallow whereas no net changes occurred in piper and imperata fallows. The study has shown large differences in biomass and nutrient stocks between the two woody fallows (piper, gliricidia) and between the woody fallows and the non-woody fallow (imperata). Short-term woody fallows are to be preferred above grass (imperata) fallows in the humid lowlands of Papua New Guinea because of higher nutrient stocks.     

Aboveground biomass and nitrogen in four short-rotation woody crop species growing with different water and nutrient availabilities

We quantified the effect of water and nutrient availability on aboveground biomass and nitrogen accumulation and partitioning in four species from the southeastern United States, loblolly pine (Pinus taeda), slash pine (Pinus elliottii), sweetgum (Liquidambar styraciflua), and sycamore (Platanus occidentalis). The 6-year-old stands received five levels of resource input (control, irrigation with 3.05 cm water week⁻¹, irrigation + 57 kg N ha⁻¹ year⁻¹, irrigation + 85 kg N ha⁻¹ year⁻¹, and irrigation + 114 kg N ha⁻¹ year⁻¹). Irrigation significantly increased foliage, stem, and branch biomass for sweetgum and sycamore, culminating in 103% and 238% increases in total aboveground biomass. Fertilization significantly increased aboveground components for all species resulting in 49, 58, 281, and 132% increases in total aboveground biomass for loblolly pine, slash pine, sweetgum, and sycamore, respectively. Standing total aboveground biomass of the fertilized treatments reached 79, 59, 48, and 54 Mg ha⁻¹ for loblolly pine, slash pine, sweetgum, and sycamore, respectively. Fertilization increased foliar nitrogen concentration for loblolly pine, sweetgum, and sycamore foliage. Irrigation increased total stand nitrogen content by 6, 14, 93, and 161% for loblolly pine, slash pine, sweetgum, and sycamore, respectively. Fertilization increased total nitrogen content by 62, 53, 172, and 69% with maximum nitrogen contents of 267, 212, 237, and 203 kg ha⁻¹ for loblolly pine, slash pine, sweetgum, and sycamore, respectively. Growth efficiency (stem growth per unit of leaf biomass) and nitrogen use efficiency (stem growth per unit of foliar nitrogen content) increased for the sycamore and sweetgum, but not the loblolly or slash pine.     

Plant growth, biomass production and nutrient accumulation by slash/mulch agroforestry systems in tropical hillsides of Colombia

Planted fallow systems under ‘slash and mulch’ management were compared with natural fallow systems at two farms (BM1 and BM2) in the Colombian Andes. The BM1 site was relatively more fertile than the BM2 site. Planted fallow systems evaluated included Calliandra calothyrsus CIAT 20400 (CAL), Indigofera constricta (IND) or Tithonia diversifola (TTH). During each pruning event slashed biomass was weighed, surface-applied to the soil on the same plot and sub-samples taken for chemical analyses. While Indigofera trees consistently showed significantly greater (p < 0.05) plant height and collar diameter than Calliandra trees at both study sites, only collar diameter in Indigofera was significantly affected at all sampling times by differences between BM1 and BM2. After 27 months, TTH presented the greatest cumulative dry weight biomass (37 t ha^–1) and nutrient accumulation in biomass (417.5 kg N ha^–1, 85.3 kg P ha^–1, 928 kg K ha^–1, 299 kg Ca ha^–1 and 127.6 kg Mg ha^–1) among planted fallow systems studied at BM1. Leaf biomass was significantly greater (P < 0.05) for CAL than IND irrespective of site. However, CAL and IND biomass from other plant parts studied and nutrient accumulation were generally similar at BM1 and BM2. At both sites, NAT consistently presented the lowest biomass production and nutrient accumulation among fallow systems. Planted fallows using Calliandra and Indigofera trees had the additional benefit of producing considerable quantities of firewood for household use. This revised version was published online in June 2006 with corrections to the Cover Date.     

Modelling agroforestry systems of cacao (Theobroma cacao) with laurel (Cordia alliodora) or poro (Erythrina poeppigiana) in Costa Rica

Predictive models were developed for Cordia alliodora branch and Theobroma cacao branch or leaf biomass,based on branch basal areas (r² 0.79) but the model of C. alliodora leaf biomass, although significant, was of very low accuracy (r² = 0.09) due to annual leaf fall. At age 10 years, shade tree stem biomass accounted for 80% of the total above-ground biomass of either tree. However, between the ages of 6 and 10 years, the biomass increment of T. cacao branches (3–4t.ha^–1.a^–1) was similar to that of the shade tree stems. During the same period, the net primary productivity was 35 and 28 t.ha^–1.a^–1, for the Erythrina poepigiana and and C. alliodora systems, respectively.Cocoa production under either of the shade trees C. alliodora or E. poeppigiana was 1000 kg.ha^–1.a^–1 (oven-dry; ages 6–10 yr). During the same period, C. alliodora timber production was 9 m³.ha^–1.a^–1 whilst the leguminous shade tree E. poeppigiana does not produce timber. Litterfall over the same 5 years, including crop and/or shade tree pruning residues, averages 11 and 23 t.ha^–1.a^–1, respectively. The main difference was due to E. poeppigiana pruning residues (10t.ha^–1.a^–1).Soil organic material reserves (0–45 cm) increased over 10 years from 198 to 240 t.ha^–1 in the E. poeppigiana plots and from 168–184 t.ha^–1 in the C. alliodora plots. These values, together with the productivity indices presented, provide evidence that the systems are sustainable.For economic reasons, the use of C. alliodora is recommended under the experimental conditions. however, on less fertile soils without fertilization, the greater biomass and hence nutrient return to the soil surface under E. poeppigiana, might make this the preferable shade tree.     

Selective control of weeds in an arable crop by mulches from some multipurpose trees in Southwestern Nigeria

The use of agroforestry systems in which pruning from trees is used to mulch the companion crops is an important area of research in the tropics. However, previous studies mostly evaluated the contribution of mulch to soil improvement and rarely examined the effect of mulch on weeds. Field experiments were conducted during the 1995 and 1996 growing seasons to investigate the effects of mulch from three woody fallow species on weed composition, biomass and maize grain yield. Treatments consisted of mulch from Leucaena leucocephala, Gliricidia sepium, and Senna siamea applied at rates of five and three tons dry matter ha^–1 at planting and three weeks after planting (WAP), respectively, an unmulched treatment that received 90 kg N ha^–1 of inorganic fertiliser, and an unmulched control plot that received no fertiliser. In both years and sampling dates, plots mulched with G. sepium and S. siamea had significantly lower weed density and biomass than the control plot in each of the sampling times and year of study. There was no significant difference in either weed density or biomass between the plot mulched with L. leucocephala and the unmulched plots. Mulches from G. sepium and S. siamea reduced weed density and weed biomass, while L. leucocephala was less effective in reducing weed biomass and weed density. Weed reduction by the mulches was in the order G. sepium S. siamea > L. leucocephala. Sedges were the dominant species in all the treatments except in G. sepium plots, where Talinum triangulare and other broadleaved species were dominant.This revised version was published online in November 2005 with corrections to the Cover Date.     

Weed biomass dynamics in planted fallow systems in the humid forest zone of southern Cameroon

Interest in planted fallow systems has focused on soil fertility improvement, neglecting other potential benefits of such systems. It is important to quantify other processes responsible for crop yield increases under planted fallows, such as weed control. The suppressive potential on weeds of Flemingia macrophylla [(Willd.) Merrill] and Pueraria phaseoloides (Roxb.) Benth, planted fallows was evaluated in field trials in three villages in southern Cameroon. In each village, experiments were set up in 4–5 year-old bush fallow dominated by Chromolaena odorata (L.) R. M. King & H. Rob. and 20 year-old secondary forest. Total aboveground biomass production of P. phaseoloides was 7.45 Mg ha⁻¹, 4.2 times higher than F. macrophylla (1.78 Mg ha⁻¹ ; P < 0.05). The high biomass of P. phaseoloides resulted in a significantly greater reduction in total weed biomass compared to Flemingia macrophylla in both wet and dry seasons. In the wet season (11 and 18 MAP), there were significant fallow system × land use and fallow system × village interactions for total weeds and broadleaf weeds. P. phaseoloides in bush (0.55 Mg ha⁻¹), and P. phaseoloides at Ngoumou (0.09 Mg ha⁻¹) had the lowest total weeds in the wet seasons. After the dry season, the lowest total weed mass was consistently recorded in P. phaseoloides while the highest was in the natural regrowth. The population of grasses was always higher in the F. macrophylla system than in P. phaseoloides system throughout the wet and dry seasons. Grass biomass in the P. phaseoloides-forest LUS was the least (0.01 Mg ha⁻¹), 58 times lower than in F. macrophylla-bush (0.58 Mg ha⁻¹). Biomass production of P. phaseoloides was highly significantly correlated with total weed biomass (r = −0.64; P = 0.004) while no relationship was found between biomass production of F. macrophylla and total weed biomass (r = −0.08, P = 0.747). It was concluded that P. phaseoloides was a suitable leguminous species for weed control. But for F. macrophylla, its low biomass production coupled with a compact plant architecture compromised it as an appropriate species for weed control in a planted fallow system.     

Direct seeding of Quercus robur L. for reforestation: The influence of mechanical site preparation and sowing date on early growth of seedlings

Compared with conventional planting, direct seeding of Quercus spp. is a method that can reduce the costs of reforestation considerably. However, interference from natural vegetation and predation on acorns contribute to regeneration failures. Mechanical site preparation has the potential to reduce both these problems. To study the influence of sowing date and of different mechanical site preparation treatments on early seedling growth, an experiment was carried out in storm-felled areas, formerly Norway spruce forests, in southern Sweden from May 2006 to September 2008. Five treatments were applied and acorns were planted in May and July 2006. The treatments were: disc trenching, patch scarification, topsoil removal, mounding and an undisturbed control. A delay of less than two months in seeding resulted in the equivalent of one year's growth reduction with respect to oak seedling's height and biomass. This could be explained by delayed emergence of seedlings, and by the dry conditions during summer seeding. The seedlings only exhibited a weak positive biomass growth response following disc trenching, patch scarification and top soil removal. In the latter case this might be explained by increased soil compaction and extensive removal of the humus layer. Site preparation by mounding resulted in good vegetation control, an increased biomass growth response and deeper seedling roots. In this treatment, we found high relative light levels near seedlings, reduced soil moisture and dry bulk density of the soil and higher soil temperatures. One or more of these environmental factors could have influenced the growth response. Whether this increased early growth rate persists throughout young stand development is, however, uncertain.     

Biomass and production of fine and coarse roots of trees under agrisilvicultural practices in north-east India

An understanding of the rooting pattern of tree species used in agroforestry systems is essential for the development and management of systems involving them. Seasonal variation, depth wise and lateral distribution of biomass in roots of different diameter classes and their annual production were studied using sequential core sampling. The investigations were carried out in four tree species under tree only and tree+crop situations at ICAR Research Farm, Barapani (Meghalya), India. The tree species were mandarin (Citrus reticulata), alder (Alnus nepalensis), cherry (Prunus cerasoides) and albizia (Paraserianthes falcataria). The contribution of fine roots to the total root biomass ranged from 87% in albizia to 77% in mandarin. The bulk of the fine roots (38% to 47%) in the four tree species was concentrated in the upper 10 cm soil layer, but the coarse roots were concentrated in 10–20 cm soil depth in alder (46%) and albizia (51%) and at 0–10 cm in cherry (41%) and mandarin (48%). In all the four tree species, biomass of both fine- and coarse-roots followed a unimodal growth curve by showing a gradual increase from spring (pre-rainy) season to autumn (post rainy) season. Biomass to necromass ratio varied between 2 to 3 in the four tree species. The maximum (3.2) ratio was observed during spring and the minimum (2) in the rainy season. In alder and albizia, the fine roots were distributed only up to 1 m distance from the tree trunk but in the other two species they were found at a distance up to 1.5 m from the tree trunk. The annual fine root production varied from 3.6 Mg ha^–1 to 6.2 Mg ha^–1 and total production from 4.2 to 8.4 Mg ha^–1 in albizia to mandarin, respectively. Cherry and mandarin had a large number of woody roots in the surface layers which pose physical hindrance during soil working and intercultural operations under agroforestry. But the high biomass of roots of these two species may be advantageous for sequential or spatially separated agroforestry systems. However, alder and albizia have the most desirable rooting characteristics for agroforestry systems.This revised version was published online in November 2005 with corrections to the Cover Date.     

Fine root dynamics of shaded cacao plantations in Costa Rica

Root turnover may contribute a significant proportion of recycled nutrients in agroforestry systems and competition between trees and crops for nutrients and water may depend on temporal fine root regrowth patterns. Fine root biomass ( 2 mm) and fine root productivity were measured during one year in plantations of cacao (Theobroma cacao) shaded by Erythrina poeppigiana or Cordia alliodora planted on a deep alluvial soil in Turrialba, Costa Rica. Fine root biomass of approximately 1.0 Mg ha^–1 varied little during the year with maximum values at the beginning of the rainy season of 1.85 Mg ha^–1 in the cacao-C. alliodora system compared to 1.20 Mg ha^–1 for cacao-E. poeppigiana. Fine root productivity of C. alliodora and E. poeppigiana (maximum of 205 and 120 kg ha^–1 4 week^–1, respectively) was greatest at the end of the rainy season, while for cacao it was greatest at the beginning of the rainy season (34–68 kg ha^–1 4 week^–1), which suggests that if nutrient competition occurs between the shade trees and the cacao, it could be minimized by early fertilization during the beginning of the rains immediately after pruning the shade trees. Annual fine root turnover was close to 1.0 in both systems. Assuming that fine root biomass in these mature plantations was constant on an annual basis, nutrient inputs from fine root turnover were estimated as 23–24 (N), 2 (P), 14–16 (K), 7–11 (Ca) and 3–10 (Mg) kg ha^–1 year^–1, representing 6–13% and 3–6% of total nutrient input in organic matter in the C. alliodora and E. poeppigiana systems, respectively.     

Initial seedling emergence of hairy birch and silver birch on abandoned fields following different site preparation regimes

Experiments were carried out at four sites in Sweden to investigate the possibility of establishing hairy birch (Betula pubescens Ehrh.) and silver birch (Betula pendula Roth) on abandoned fields using natural regeneration and (or) direct seeding. The effects of six soil preparation methods (no preparation, ordinary ploughing, rotary cultivation, deep ploughing, inverted ground, removal of top-soil) and five additional treatments (no treatment, herbicide, peat litter, wood-ashes, slaked lime) on seedling emergence percentages (SEP) and vegetation cover percentages (VCP) were studied. SEP-values were estimated in June (SEP(J)) and October (SEP(O)). The experimental designs used were: 1) split-plot design with whole plots in a randomized complete block design (RCBD); 2) RCBD. The SEP-values observed with no preparation were close to 0% while the SEP-values obtained with mechanical soil preparation methods mostly were much higher (p0.014). Seedbeds with top-soil in the surface, created by ordinary ploughing or rotary cultivation, and seedbeds with mainly bare mineral soil in the surface, created by deep ploughing, inverted ground or removal of top-soil, obtained SEP-values of equal merit on silty soils, reaching SEP(O)-values up to 15%. The latter seedbeds obtained the best results on sandy soil, with as high SEP(O)-values as 47% after removal of top-soil. Seedbeds with top-soil in the surface were quickly colonized by ground vegetation, reaching VCP-values between 70% and 100%. Herbicide spraying with glyphosate and application of peat litter to the seedbed surface promoted seedling emergence. However, herbicide spraying before soil preparation was of little effect when followed by mechanical soil preparation.     

Role of weeds in the management of nitrogen in a young Pinus radiata plantation

Pinus radiata trees were grown on a podzolized sandy soil at a second rotation site under the following treatments: total weed control, total weed control plus ammonium nitrate, strip weed control and no weed control. During the first two summers after planting the differences in needle water potential between trees under no, strip or total weed control were very small. Despite similar rates of net N-mineralization in strip and total weed control treatments, which averaged 64 kg ha^–1 yr^–1 in the 0–15 cm soil depth, weeds in the strip weed control treatment reduced soil mineral-N concentrations by 50–80%, leaching of N by the end of the first growing season by 45%, foliar-N concentrations by 4–14% and stem biomass at 20 months after planting by 46%. Although N-uptake by above-ground vegetation (trees plus weeds) was 49% higher in the strip weed control treatment, the amount of N apportioned to trees during the first 20 months after planting was reduced from 15.5 to 9.0 kg ha^–1. These effects of weeds were even more pronounced in the no weed control treatment. Since weeds had little effect on the needle water potential of trees and the annual rates of N-mineralization, but adversely affected N-uptake by trees, results indicate that weeds directly competed with trees for N, and thereby aggravated N-deficiency in trees. Application of ammonium nitrate after complete weed control increased foliar-N concentrations, and N-uptake and growth of trees, but also induced severe stem deformation.     

Biomass production of tree fallows and their residual effect on maize in Zimbabwe

The rotation of maize (Zea mays) with fast-growing, N₂-fixing trees (improved fallows) can increase soil fertility and crop yields on N-deficient soils. There is little predictive understanding on the magnitude and duration of residual effects of improved fallows on maize yield. Our objectives were to determine the effect of fallow species and duration on biomass production and to relate biomass produced during the fallow to residual effects on maize. The study was conducted on an N-deficient, sandy loam (Alfisol) under unimodal rainfall conditions in Zimbabwe. Three fallow species — Acacia angustissima, pigeonpea (Cajanus cajan), and Sesbania sesban — of one-, two-, and three-year duration were followed by three seasons of maize. Pigeonpea and acacia produced more fallow biomass than sesbania. The regrowth of acacia during post-fallow maize cropping provided an annual input of biomass to maize. Grain yields for the first unfertilized maize crop after the fallows were higher following sesbania (mean = 4.2 Mg ha^–1) than acacia (mean = 2.6 Mg ha^–1). The increased yield of the first maize crop following sesbania was directly related to leaf biomass of sesbania at the end of the fallow. Nitrogen fertilizer did not increase yield of the first maize crop following one- and two-year sesbania fallows, but it increased yield following acacia fallows. Nitrogen fertilizer supplementation was not required for the first maize crop after sesbania, which produced high-quality biomass. For acacia, which produced low-quality biomass and regrew after cutting, N fertilizer increased yield of the first post-fallow maize crop, but it had little benefit on yield of the third post-fallow maize crop.This revised version was published online in November 2005 with corrections to the Cover Date.     

Management of pigeon pea in short fallows for crop-livestock production systems in the Guinea savanna zone of northern Ghana

Crop and livestock production in the Guinea savanna zone of northern Ghana has been declining over the past years as a result of increasing pressure on land. To sustain soil productivity, pigeon pea(Cajanus cajan), a leguminous perennial crop was evaluated for its potential as a short duration fallow crop for fodder and grain, and maize (Zea mays)production. It involved comparing a natural fallow (i.e., control) and four improved fallows of pigeon pea pruned annually at 30 cm, 60 cm and 90 cm from the ground, and unpruned pigeon pea over a two-year period. After this time, the land was cleared manually and planted to maize. The highest mean annual biomass of pigeon pea over the two-year period of 6.1 t ha⁻¹ dry matter (DM) was obtained by pruning at 60 cm. The highest leaf litter production and pigeon pea seed yield was obtained from the no pruning treatment. The mean maize grain yield from the improved fallow (3.02 t ha⁻¹) in the first year after clearing was significantly (P < 0.05) greater than that of the natural fallow (1.54 t ha⁻¹). Considering the biomass of pigeon pea from pruning, pigeon pea seed yield and maize grain yield after the pigeon pea, pruning pigeon pea at 60 cm is the most promising regime for crop-livestock production systems. This revised version was published online in June 2006 with corrections to the Cover Date.     

Substitution of nitrogen requirement of maize through leaf biomass of Leucaena leucocephala: agronomic and economic considerations

In a 3-year field study, the effects of substitution of nitrogen requirement of maize through Leucaena leaves were studied on runoff, soil loss, maize and wheat yield and economic returns. The treatments were (1) 80 kg N ha^-;1 all through Leucaena leaves (80 L), (2) 40 kg N through Leucaena leaves + 40 kg N ha^-;1 through fertilizer (40 L + 40 F), (3) 20 kg N through Leucaena leaves + 60 kg N ha^-;1 through fertilizer (20 L + 60 F), (4) 80 kg N ha^-;1 all through fertilizer (80 F), and (5) control (No fertilizer). Green Leucaena leaf biomass (containing 3.3% N on dry basis) was incorporated every year in 15 cm top soil two weeks before sowing of summer maize.Other treatments being almost equal, runoff was reduced marginally in treatment 20 L + 60 F which was attributed to better crop growth in this treatment. Mean minimum soil loss (6.202 t ha^-;1) also occurred in treatment 20 L + 60 F. Soil loss in 80 L was 13% less than in 80 F. Maize yield was at par in 80 L and 80 F. However, mean maximum yield of maize was obtained with 20 L + 60 F.Residual effect of incorporation of Leucaena leaves to maize crop was observed on wheat yield. The mean yield differences were statistically at par in all the treatments except control. The total mean net returns were statistically at par in 80 L and 80 F. However, significantly higher mean net returns (Rs 6811 ha^-;1; one US$ = Rs30) were obtained with 20 L + 60 F. Substitution of N through Leucaena leaves even in small quantity may be helpful to small holders, particularly where chemical fertilizers are in short supply or too expensive.     

Productivity and nutrient cycling in young Acacia senegal farming systems on Vertisol in the Blue Nile region, Sudan

Acacia senegal, the gum arabic-producing tree, is the most important component of traditional dryland agroforestry systems in the␣Sudan. The spatial arrangement of trees and the type of agricultural crop used influence the interaction between trees and crops. Tree and crop growth, gum and crop yields and nutrient cycling were investigated over a period of 4 years. Trees were grown at 5 × 5 m and 10 × 10 m spacing alone or in mixtures with sorghum or sesame. No statistically significant differences in sorghum or sesame yields between the intercropping and control treatments were observed (mean values were 1.54 and 1.54 t ha⁻¹ for sorghum grain and 0.36 and 0.42 t ha⁻¹ for sesame seed in the mixed and mono-crop plots, respectively). At an early stage of agroforestry system management, A. senegal had no detrimental effect on crop yield; however, the pattern of resource capture by trees and crops may change as the system matures. A significant positive relationship existed between the second gum picking and the total gum yield. The second gum picking seems to be a decisive factor in gum production and could be used as an indicator for the prediction of the total gum yield. Soil organic carbon, N, P and K contents were not increased by agroforestry as compared to the initial levels. Soil OC was not increased by agroforestry as compared to sole cropping. There was no evidence that P increased in the topsoil as the agroforestry plantations aged. At a stocking density of 400 trees ha⁻¹ (5 × 5 m spacing), A. senegal accumulated in its biomass a total of 18.0, 1.21, 7.8 and 972 kg ha⁻¹ of N, P, K and OC, respectively. Agroforestry contributed ca. 217 and 1500 kg ha⁻¹ of K and OC, respectively, to the top 25-cm of soil during the first four years of intercropping.