Ultrastructure of pits inPinus banksiana lamb

Summary The cross-sectional view of pitting between various cell types inPinus banksiana Lamb. was studied at the ultrastructural level. Cell types inPinus banksiana include longitudinal tracheids, ray tracheids, ray parenchyma cells, buffer cells and epithelial cells. Two common characteristic features of bordered pit-pairs between longitudinal tracheids are an initial pit border and a thickened torus at the center of the pit membrane. The shape and size of the pit border and torus of bordered pit-pairs between two compression wood cells, and between the last-formed latewood longitudinal tracheid and first-formed earlywood longitudinal tracheid were different from those in the earlywood and latewood longitudinal tracheids. The pit border on the ray tracheid side varied in size and shape due to wall dentation. No initial pit border was found on the pit border of the ray tracheid side. The shape of bordered pit-pairs between two ray tracheids varied considerably due to irregularity of the dentate cell wall. The size of bordered pit-pairs in longitudinal tracheids was between 16 m to 20 m, which was twice the diameter of bordered pit-pairs in ray tracheids. Bordered pitpairs at the end wall of two ray tracheids appeared to be the smallest at 5 m, Pit aspiration occurred in the bordered pit-pairs with or without a torus. In the heartwood zone, some half-borders pit-pairs between tracheary and ray parenchyma cells showed an additional secondary wall on the ray parenchyma cell side. Plasmodesmata were found in the half-bordered pit-pairs as well in the simple pit-pairs. Blind pits were observed between a ray tracheid and a longitudinal tracheid. Bordered pit-pairs between two buffer cells were also observed. The possible functions of buffer cells were discussed.Use of transmission electron microscope provided by the Science Instrumentation Lab, Lakehead University and the technical assistance provided by Mr. A. MacKenzie, Director of Science Instrumentation Lab are gratefully appreciated     

Maximum effective lumen and pit pore sizes of the earlywood and the latewood of never dried loblolly pine sapwood

Summary Measurements of the air pressure required to initially displace a saturating liquid and allow a slow continuous stream of air bubbles to pass through wood cross sections of different thickness, together with the equilibrium surface tension of the saturating liquid, make it possible to calculate the maximum effective opening radii. Previous measurements were made for wood as a whole over complete annual rings. Measurements reported here were made separately for earlywood and latewood. Extrapolating plots of the maximum effective opening radius-cross section thickness, for thicknesses below the maximum fiber length, to zero thickness gave maximum lumen radii of 16 m for the earlywood and 10.3 m for the latewood. The values are only slightly greater than the calculated average values. Extrapolating the plots in the opposite direction to zero opening radius gave approximate maximum lumen or fiber lengths for the earlywood of 6 mm and for the latewood of 5 mm. The maximum effective opening radii for cross sections thicker than the maximum fiber length give maximum effective communicating pore radii. These values continue to decrease, with increasing thickness of the cross sections due to the decreasing probability of the largest openings falling in any one series path through the structures. The maximum effective pit pore radius for passage through fifty pits in series was 0.8 m for the earlywood and 0.28 m for the latewood.Paper No. 3787 of the Journal Series of North Carolina State University Agricultural Experiment Station, Releigh, N. C.     

Lignin in woody tissues after chemical pretreatment and bacterial attack

Summary Tangential latewood sections (60 m) of Scots pine sapwood were differently treated with chlorite. The subsequently incubated two strains of Bacillus polymyxa caused a weight loss up to about 25%, including a loss of lignin of about 42%.UV-microspectrophotometry of 1 m-cross-sections prepared from the cultured woody tissues demonstrated that bacteria caused neither quantitative nor qualitative changes of the remaining lignin.The lignin, which was dissociated from the pretreated woody cell wall by bacteria, could not be respired, suggesting that the lignin is a ballast to these bacteria that inhibits the dissimilation of the carbohydrates in the wood.We appreciate the financial support by Deutsche Forschungsgemeinschaft. Thanks are due to Mrs. R. Endeward for her assistance during the microspectrophotometric evaluations and to Mr. U. Engel and Dr. O. Faix for carrying out the IR-measurements     

Water circulation, groundwater outflow and nutrient dynamics in Mida creek, Kenya

The relationship between physical hydrodynamic processes and nutrients dynamics was investigated in Mida creek, a groundwater influenced mangrovefringed creek in Kenya between March 1996 and May 1997. The research involved spot and timeseries measurement of nitrate–nitrite, ammonia, silicates, phosphates, salinity, temperature, sealevel as well as tidal currents at seven stations located in the front, middle and backwater zones of the creek. Groundwater level as well as total dissolved solids' concentration, salinity, temperature and nutrients' concentration were also measured once every month in shallow wells (watertable<5m) located in the upper region of the creek. Results of the study show that nutrient concentrations vary with the tide and that, though there is no river drainage, they are of the same magnitude as in mangrove creeks with substantial river runoff. The peak concentrations of NH ₄ ⁺ –N (5.45M), NO ₂ –NO ₃ (5.63M), PO ₄ ^3– –P (0.58M) and SiO ₃ ^2– –Si (81.36M) in the creek occurred during flood tide, 2–3h before high waters. The (NO ₂ ^– + NO ₃ ^– )–N concentrations declined rapidly during ebb tide, reaching the minimum levels during low water. Contribution of groundwater seepage to the net nutrients flux (particularly on nitrite–nitrates) is largest in dry seasons. The study shows that groundwater outflow sustains the mangroves during periods of severe salinity stress and nutrients deficiency in dry seasons. This is essentially by limiting salinity increase and by boosting nutrient supply in dry seasons.     

Collapse and internal checking in the latewood of Eucalyptus regnans F.Muell

Summary Tangential shrinkage was measured on longitudinal-tangential slices of separate earlywood and latewood from one board of Eucalyptus regnans F.Muell. at various temperatures. Large amounts of collapse shrinkage were measured in the latewood slices, and lesser amounts in the earlywood slices. Collapse shrinkage in the earlywood was found only when the slices were dried at temperatures above a minimum temperature (the collapse threshold temperature). End-coated board sections approximately 200 mm long were rapidly dried at dry-bulb temperatures below the collapse threshold temperature for earlywood. Incipient internal checks were found in the latewood of these boards. Board sections dried at higher temperatures showed internal checks starting in both early and latewood. A non-linear drying simulation model was modified to take the heterogeneous nature of wood into account. This model predicted that internal checks would form in this wood even if it were dried sufficiently slowly to avoid surface checking.Symbols D Diffusion coefficient - e_c Creep strain - e_i Instantaneous strain - e_m Mechano-sorptive strain - e_n Net strain - e_u Unconfined shrinkage strain - EW Earlywood - FSP Fibre saturation point - LW Latewood - L-T Longitudinal-tangential slice - MC Moisture content (kg water/kg dry wood) - q Moisture concentration - R-T Radial-tangential slice - t Time - y Depth in board The author is pleased to acknowledge the assistance of Emeritus Professor A. R. Oliver, Associate Professor P. E. Doe, University of Tasmania, and the Australian Furniture Research and Development Institute     

Micropropagation of western hemlock (Tsuga heterophylla [Raf.] Sarg.) from embryonic explants

An in vitro propagation protocol is described for western hemlock, an important forestry species in Canada. For shoot bud induction, embryonic explants were placed initially on one-third strength Schenk and Hildebrandt medium containing 5 M N⁶-benzyladenine (BA) or 5 M BA in combination with either 5 M kinetin or 5 M 2-isopentenyl adenine for 14 days. The explants were transferred to basal medium without cytokinins for 3 weeks, and then to basal medium containing 0.05% activated charcoal. Elongating shoots were subcultured every 4 weeks on charcoal medium. Shoots, 10 mm in stem height, were rooted either in agar or sterilized peat/perlite (1:1). Up to 70% of the shoots formed roots when they were transferred to the latter, moistened with 1/2 strength Gresshoff and Doy medium containing 5 M -naphthaleneacetic acid. About 90% of the plantlets survived transfer to greenhouse conditions.     

The effects of drying on the structure and permeability of the wood of Abies grandis

Summary The effects of air-drying and solvent-drying on the sapwood of Abies grandis have been investigated by a new method for the determination of the size and number of conducting tracheid lumina and pit membrane pores which involves the measurement of gaseous permeability at various mean pressures. Both earlywood and latewood tracheids (83% of the total) were found to be conducting in solvent-dried wood, but in air-dried wood only latewood tracheids (32% of the total) were conducting. In solvent-dried wood there were on average 27,000 pit membrane pores per conducting tracheid compared with only 600 in air-dried wood. In both, the average pit membrane pore radius was about 0.1 m.Liquid permeabilities have been predicted from the calculated radii and numbers. The liquid permeability of solvent-dried wood was 31 times greater than that of air-dried wood in which the lumina were responsible for 13% of the total resistance to flow. The lumina were responsible for 39% of the resistance in solvent-dried wood and it is suggested that in first-formed earlywood the lumina may cause more than half the total resistance.A new method is described for the cleaning of direct carbon replicas of wood. In this the cellulose is removed by cellulase instead of sulphuric acid, and no wax backing is required. This provides much cleaner replicas. Electron micrographs have been obtained of both earlywood and latewood dried by the two methods.The authors wish to thank Mr. A. R. Sayers for preparing the computer programme used in this work, Dr. R. Ph. C. Johnson for his help and advice regarding the electron microscopy and Professors Matthews and Weatherley for their advice and encouragement.     

Tangential pitting in black spruce tracheids

Tangential pit features were studied in a 55-year old black spruce (Picea mariana (Mill) B.S.P.) tree by means of light and electron microscopy.It was found that tangential pitting is lacking from the greatest part of the growth ring, except for the last four tangential rows of latewood tracheids and the first row of early wood tracheids. The average number of pits per tangential wall of a 3.55-mm-long tracheid is 234, 144, 28, 4 and zero, respectively, in the last 5 tangential rows of latewood tracheids, starting at the growth-ring boundary.On the average, tangential pits measure 5.4 m in diameter, possess oval to elliptical apertures, and are randomly distributed uniformly over the tangential tracheid wall. All tangential intertracheid pits are bordered and in that respect are similar to those in the radial walls. Although most of the pits contain membranes with tori, some at the growth-ring boundary lack tori and exhibit randomly oriented microfibrillar structure.     

The distribution of lignin in sprucewood as determined by ultraviolet microscopy

The distribution of lignin in black spruce has been determined quantitatively by the study of 0.5 m transverse sections in a UV microscope. The average lignin concentration in the compound middle lamella was about twice that in the secondary wall. The lignin concentration of the middle lamella at the cell corners of adjacent tracheids was nearly four times that in the secondary wall but the volume of the secondary wall was much greater than the volume of the middle lamella. Thus, for earlywood, 72% of the total lignin was in the secondary wall leaving only 28% in the compound middle lamella and cell corner middle lamella regions. The corresponding values for latewood were 82% and 18% respectively. Use of oblique longitudinal sections of 0.1 m thick permitted the resolution of the compound middle lamella. The lignin concentration in the true middle lamella was found to be equal to that in the cell corner middle lamella and the primary wall lignin content to be about twice that in the secondary wall.

---

Zusammenfassung Die Verteilung des Lignins in Fichtenholz wurde quantitativ durch Untersuchung von 0,5 m dicken Querschnitten unter dem UV-Mikroskop bestimmt. Die mittlere Ligninkonzentration war in der Mittelschicht etwa doppelt so hoch wie in der Sekundärwand. Die Ligninkonzentration der Mittelschicht war in den an die Tracheiden anstoßenden Zellecken annähernd viermal höher als in der Sekundärwand, wogegen das Volumen der Sekundärwand wesentlich größer war als das der Mittelschicht. Dagegen befand sich beim Frühholz 72% des gesamten Lignins in der Sekundärwand und nur 28% fanden sich in der Mittelschicht selbst und in ihren Zelleckbereichen. Die entsprechenden Werte für Spätholz betragen 82% bzw. 18%. Die Anwendung von schrägen Längsschnitten von 0,1 m Dicke erlaubte die Auflösung der Mittelschicht. Die Ligninkonzentration in der Mittllamelle war gleich groß wie in der in den Zellzwickeln befindlichen Mittellamelle und der Ligningehalt der Primärwand war etwa doppelt so groß wie derjenige in der Sekundärwand.

     

Stiffening phenomena in annual rings of coniferous wood

Summary The structure of coniferous wood is characterized by pronounced differences in density between earlywood and latewood. This could result in disadvantages in wood strength especially against bending. A summary of the observations in this paper leads to the following hypothesis: Coniferous trees meet this danger by forming numerous stiffenings especially in broad annual rings with a high percentage of earlywood. Such stiffenings may be produced by more or less slight fluctuations of density or chemistry within annual rings in radial as well as in tangential directions. The question remains open which anatomical or chemical reasons, upon sandblasting, locally cause radial or radially branched ribs and craters in cross sections, waviness in radial sections and cloudy structures in tangential sections.Paper presented to the IUFRO-Division 5. Conference in Madison, Wisc. 1983.     

On movement of water through wood — The diffusion coefficient

Summary It is demonstrated that there can be only one driving potential for the movement of water through wood and this will be a function of wood state. On the assumption that the driving potential is the partial pressure of water vapour, a theoretical expression is derived for the diffusion coefficient. Such expression is fitted to diffusion coefficients for Scots pine and a remarkably good fit is obtained.Symbols a reciprocal mean radius of curvature of a capillary meniscus; also taken to be the radius of the corresponding exposed liquid surface, m - b spacing between flow paths in the cell wall, m - D diffusion coefficient for water in wood with vapour pressure as the driving potential, kg/ms Pa - D_a diffusion coefficient for water vapour through air, kg/ms Pa - D diffusion coefficient for water in wood with the driving potential - D diffusion coefficient for water in wood with the driving potential - D₀ diffusion coefficient for water in wood with vapour pressure as the driving potential, which is associated with leakage paths through the wood, kg/ms Pa - D_f diffusion coefficient for water in wood with vapour pressure as the driving potential, corresponding to fibre saturation and with no leakage paths, kg/ms Pa - D_c diffusion coefficient for water in wood with vapour pressure as the driving potential, which is associated with the constriction of the vapour flow as it approaches the cell wall, kg/ms Pa - D diffusion coefficient for water in wood with moisture content as the driving potential, kg/ms - diffusivity for water vapour in air, m²/s - F flux of water, kg/m² s - p partial pressure of water vapour, Pa - R specific gas constant for water, J/kg K - r fractional relative humidity - T temperature, K - x length coordinate in direction of flow, m - the dimensionless ratio D_f/D_c evaluated at r=1/e - arbitrary driving potential for movement of water in wood - cell spacing in the direction of water flux, m - density of liquid water, kg/m³ - coefficient of surface tension, N/m - arbitrary driving potential for movement of water in wood - fractional moisture content     

Structure of cellulose microfibrils and the hydration effect in <Emphasis Type="Italic">Cryptomeria japonica</Emphasis>: a small-angle X-ray scattering study

The nanometer scale structure of cell walls in sugi wood (Cryptomeria japonica D. Don) and the hydration dependence were examined by the small angle X-ray scattering technique. Disk-shaped scattering patterns were observed for sugi wood. The radial average of two-dimensional data from the cross section could supply the scattering intensity with statistical accuracy much higher than that obtained from the sector average of the streak-shaped scattering pattern, and both the scattering intensities provided similar structural information. The scattering patterns from the cross section of the wood are characterized by rhombic or cross-shaped isointensity curves for the lower q region and by circularly symmetric isointensity curves for the higher q region. This shows that the disk-shaped scattering has two different kinds of scattering origins. The microfibril radii in the cell wall were determined by fitting the model scattering function of cylindrical fibrils to the scattering data. Values of 12.3 ± 0.3 and 12.2 ± 0.3 were obtained for the fibril radii of the neighboring earlywood and latewood, respectively, in dry specimens. A drastic structural change of the cell walls was detected with increasing water content from 40% to 100%. A low q rise in the scattering intensities below 0.1^–1 became weak and changed into a flat pattern, and the rhombic isointensity curves changed to cross-shaped patterns in the two-dimensional scattering from the cross section. The calculated radii R increased from 12.2 ± 0.3 to 13.3 ± 0.1.Part of this report was presented at the 52nd Annual Meeting of the Japan Wood Research Society, Gifu, Japan, April 2002     

The measurement of tensile creep in thin wood strips

Summary Tensile creep of 100 m thick wood strips cut parallel to the grain was measured using a high resolution high accuracy creep apparatus. Rigorous attempts were made in both design and use of the apparatus to reduce errors to a minimum: the standard deviation of the random errors about a smoothed curve was assessed as 0.34 m. By comparing elongation of strips under load with control strips at zero load viscous and visco-elastic elongation may be accurately measured.     

Spiral angle of elementary cellulose fibrils in cell walls ofPicea abies determined by small-angle x-ray scattering

Summary The spiral angle of the elementary cellulose fibril in the wood cell wall, often called microfibril angle, is of primary importance for the mechanical properties of wood. While there are a number of methods to estimate this angle, x-ray diffraction (XRD) techniques have recently obtained a lot of attention because of their ability to provide information averaged over a significantly large specimen volume. Here, we present results from a related method, small-angle x-ray scattering (SAXS). The advantage of SAXS is that, unlike XRD, it does not require any assumption on the orientation of the cellulose crystal axis with respect to the fibril axis. Full three-dimensional scattering patterns were collected using an area detector by rotating the sample around one axis. The distribution of fibrillar orientations was seen to reflect the typical cross-sectional shape of the tracheids (square or circular). In the stem, the spiral angle was found <5° in earlywood and 20° in latewood. In branches the angle was 30° in the upper part and 40° in the lower part, which strongly supports the idea that the spiral angle has primarily mechanical function.We thank W. Laube, F. Einramhof, H. Königshofer and H. Löppert for their technical assistance. Financial support from the Fonds zu Förderung der Wissenschaftlichen Forschung (Proj. P10729-BIO) is gratefully acknowledged     

A simple model for density of annual rings

Summary A simple model for density distribution within an annual ring is proposed. The model originates from the concept that a tree has two growing modes early and late wood mode respectively and that during a growing season there is transition from the first to second. An annual ring is described by six parameters. The use of the model is discussed and a number of possible measures are suggested. The advantages and limitations of the model is also discussed.     

Five heavy metals in propagules of ten mangrove species of China

Five heavy metals in the propagules of 10 mangrove species in China have been investigated. The results revealed that the levels of five heavy metals in most of the propagules were lower than the background levels of the soil. The levels of copper, manganese, zinc, cadmium, and lead in the propagules varied at about 2.1–7.8 g/g, 3.9–28.0 g/g, 5.7–60.0 g/g, 0.014–0.057 g/g, and 0.018–0.038 g/g, respectively. On average, the levels of five heavy metals were in the order Zn > Mn > Cu > Cd > Pb. The biological absorption coefficients were 0.02–1.30 for copper, manganese, zinc, and cadmium and about 0.0007–0.0020 for lead in propagules, showing the following order: Zn > Cd > Cu > Mn > Pb. The above results indicate that the five heavy metals have low accumulation in most of the propagules of 10 mangrove species and are at safe levels for the effective utilization of mangrove resources.     

Tissue culture micropropagation of Douglas-fir

A procedure was developed for plantlet production from embryos of Douglas-fir [Pseudotsuga menziesii (Mirb.) Franco]. The Medium for Conifer Morphogenesis, used at full strength, and supplemented with 10 M benzyladenine for 17 days produced an average of 6.8 shoots on more than 90% of the embryos. The percentage of shoot-forming embryos as well as the average number of shoots per embryo varied significantly among eight seedlots. For secondary multiplication, 89% of the adventitious shoots produced axillary buds on MCM with 5 M benzyladenine. However, 0.5 M BA was more suitable for the elongation of axillaries. Rooting ranged from 0–87% depending upon the treatment. The highest percentage was obtained with a 7-week incubation in peat:perlite containing 1/5-strength medium, 1% sucrose and 2.7 M naphthaleneacetic acid, followed by 5 weeks on peat:perlite with 1/5-strength major and minor salts, full iron and organics, 1% sucrose and 0.1% charcoal.     

Cross sectional dimensions of wood pulp fibers from softwood fiber counts

Summary There are, on the average, about 300 fibers or tracheids per centimeter in both the tangential and radial directions in cross sections of softwoods when the counts are made over complete annual rings with the extremes varying from about 200 fibers per centimeter for redwood to about 400 fibers per centimeter for Alaska yellow cedar. Fiber widths, including half of surrounding middle lamella, range from about 50 m for redwood to 25m for Alaska yellow cedar, averaging about 33 m. Average lumen widths vary but slightly with changes in the specific gravity of the wood whereas the double cell wall thickness varies directly with the specific gravity. Effects of pulping to different pulp yields on the fiber dimensions are calculated with the use of two newly developed equations. The theoretical thickness of completely collapsed pulp fibers is equal to their double cell wall thickness. Complete collapse rarely if ever occurs as it requires double fractures of the cell wall. In practice, collapse of fibers is incomplete so their thickness is usually two or more times the theoretical minimum.Emeritus Robertson Professor of Wood and Paper Science, North Carolina State University, Raleigh, N.C., U.S.A.     

Dimensional instability of cement bonded particleboard: Modelling CBPB as a composite of two materials

Previous papers have quantitatively indicated that the total movement of cement bonded particleboard (CBPB) is equal to the sum of the movement of its components. This paper examined the efficacy of the law of mixtures when applied to the movement of a wood-cement composite under internal swelling or shrinkage stresses. Abundant data generated in companion papers were first manipulated to develop the universal formulae for predicting the movement of components. In conjunction with previous numerical results from image analysis of the structure of CBPB, and the orientated elasticity and stress algorithms, the models for theoretically predicting mass and dimensional changes of CBPB were derived. Validation studies were conducted and these demonstrated an excellent agreement of the theoretical predictions with experimental data for both mass and dimensional changes of CBPB due to internal swelling or shrinkage stresses during adsorption and desorption. The success also implied that CBPB can be treated as a composite and its properties can be well derived by the law of mixtures even though CBPB is an unusual type of composite having a very high volume fraction of wood chips, but a very high mass fraction of cement paste.Notation E_RT Mean transverse modulus of elasticity of wood - E_L Longitudinal modulus of elasticity of wood - E_p Modulus of elasticity of cement paste - E_wa Modulus of elasticity of embedded wood chips at angle - E Modulus of elasticity of wood chips at direction - E Modulus of elasticity of wood chips at direction - G_LRT Mean transverse shear modulus of wood - L(T)_cp Length/width (thickness) change of CBPB at angle - L(T)_p Length (thickness) change of cement paste - m_pf Mass fraction of cement paste in unit mass of CBPB - m_wf Mass fraction of wood chips in unit mass of CBPB - M_cpj Mass change of CBPB at the various conditions tested - M_pj Mass change of cement paste at corresponding conditions - M_wj Mass change of wood chips at corresponding conditions - M(L; T)_w/P Mass, length or thickness changes of wood chips or cement paste at various conditions - t Duration of exposure - _LRT Mean transverse Poissons ratio of wood - V_pf Volume fraction of cement paste in unit mass of CBPB - V_wf Volume fraction of wood chip in unit mass of CBPB - _cp Density of CBPB - _k Density of wood chip or cement paste - _cp Overall stresses of CBPB at angle - _L Stress in the longitudinal direction of wood - _RT Mean stress in the transverse direction of wood - _p Stress of cement paste - _w Stress of the wood chips at angle - Stress of the wood chips at direction - Stress of the chip at direction - _cp Strain in CBPB - _p Strain of cement paste - _WL Strain in the length of wood chips - _WT Strain in the thickness of wood chips - _w Strain in wood chips - Angle between the longitudinal direction of wood chips and surfaces or edges of CBPB - Angle between wood chips and edges (length direction) of CBPB - Angle between wood chip and vertical coordinate - A, B, C Coefficients related to the feature of materials and exposure conditions The senior author wishes to thank Professor W.B. Banks of University of Wales, Bangor for his constructive discussions and assistance and the British Council for partly financial support.     

Zur Tagesdynamik des Harzdruckes und der Harzabsonderung sowie ihre Bedeutung für den Befall der Kiefer durch den TriebwicklerRhyacionia buoliana Schiff. (Lep., Tortricidae)

Zusammenfassung Es wurden in Kiefernkulturen Untersuchungen über die Tagesdynamik des Harzdruckes und der Intensität der Harzabsonderung geführt und die Ergebnisse in Beziehung zum Befallsgrad des KiefernknospentriebwicklersRhyacionia buoliana Schiff. gesetzt.Im Verlauf eines Tages variierte der Harzdruck wenig, die Intensität der Harzabsonderung dagegen stark. Die höchste Harzfluß-Intensität fällt in den Zeitraum zwischen 13 und 16 Uhr. Die Flugzeit des Wicklers, nach 18 Uhr, liegt somit außerhalb der Zeit des stärksten Harzflusses.Weder der Index der Harzfluß-Intensität noch jener des Harzdruckes zeigten eine Beziehung zur Stärke des Wicklerbefalls. Sie sind daher als Kriterien der Widerstandsfähigkeit einer Kulturkiefer gegenRh. buoliana, zumindest unter den Verhältnissen der Süd-Ukraine, nicht verwendbar.

---

Summary Studies on the diurnal rhythm of resin pressure and resin secretion in connection with the degree of pine-infestation byRhyacionia buoliana Schiff.In the course of one day pines in the age of 15 years showed only a small variation in resin pressure but a great one in secretion of resin. The maximal intensity of secretion was observed early in the afternoon. The moths ofRh. buoliana are flying after this time when the secretion of resin is low.There was no connection between resin pressure and secretion of resin on the one hand and the degree of pine-infestation byRh. buoliana on the other. The two factors are not suitable for indicating the degree of pine-infestation by this tortricid.

---

- . / 0 26 / /., , . , 13 15 . , , , , .