不同经营方式下杨树人工林叶面积分布与动态研究

本文对集约经营与粗放经营杨树人工林分的叶面积垂直分布及其片节动态变化进行了对比研究，结果表明：集约经营林木叶面积的数值明显大于粗放经营林木的叶面积数值，林木个体间叶面积差异显著且与林木个体的胸径大小成正比。整个生长季节内，集约经营林木冠层叶面积数值较大，林分生长季节较长；而粗放经营林木叶面积相对较小，生长季较短。叶面积较大，生长季较长是导致集约经营林分生产力较高的主要原因之一，通过研究，建立了生长季不同时期内，集约经营与粗放经营林木累积叶面积指数垂直分布的拟合模型，可用于对叶面积的估算。该研究为进一步探讨杨树人工林冠层光分布、计算冠层光合产产量奠定了基础，为我国杨树人工林的速生、丰产提供了科学依据。     

刺槐蒸腾耗水的动态研究(英文)

推算林分的年总耗水量 ,首先要确定林分的年总蒸腾量 .该文以LI 16 0 0稳态气孔仪在 1998～2 0 0 0年生长季各月典型天气的实测数据和常规气象数据为基础 ,应用Penman Monteith方程模拟了刺槐林的日蒸腾过程 ,并以此计算了生长季的连续日蒸腾 .刺槐林林冠层截留净辐射由实测数据 (Rn)与叶面积指数 (LAI)建立的消光系数 (k)数学关系确定 ,冠层整体气孔阻力由实测数据拟合经验式计算 .经与实测结果对比 ,模拟的相对误差平均在 12 %以内 .由全年连续日蒸腾模拟计算 ,1998年至2 0 0 0年 5月～ 10月生长季不同密度刺槐林的平均总蒸腾量分别为 192 4 6 ,187 0 7和 195 5 9mm .     

北京山区典型林分生长季叶面积指数动态变化

叶面积指数为植物冠层表面的物质循环与能量循环提供了结构化的定量信息,是估算植被冠层功能的重要参数,也是生态系统中最重要的结构参数之一。以北京西山实验林场作为研究区域,以研究区域内5种典型林分作为研究对象,对各林分生长季内的叶面积指数进行测量,结果表明:1)不同植物类型叶面积指数表现为乔木林草本层灌木林;4种乔木林分类型生长季平均叶面积指数大小排序为栓皮栎林刺槐林油松林侧柏林。2)在整个生长季,4种乔木林分的叶面积指数表现出了一致的变化趋势,均表现为"增大—平稳—减小"的趋势;3)灌木林叶面积指数的变异系数均高于乔木林与草本层,就整个生长季,5种林分的变异系数为灌木林(荆条)栓皮栎林刺槐林侧柏林油松林。     

不同密度杉木林内辐射与叶面积垂直分布对生长的影响

本文通过对八年生杉木人工林内太阳辐射与叶面积垂直分布的观测与研究，用计算机绘图与计算，得出以下结果：杉木叶面积的垂直分布和叶面积指数与密度有关，冠形因密度增加由圆锥形变为圆柱形，叶面积指数是先上升后下降。杉木林内辐射的消减随密度的增加而加剧，在冠层深3／4处趋于平缓，当郁闭度低于0．85时，林冠下辐射有回升，用累加叶面积指数分层计算林冠消光系数，可减少因叶片分布不均匀而产生的误差，用累计值计算辐射，可简化观测与计算，用辐射吸收率、叶面积指数，杉木个体生物量及年均生物增长量4个指标对杉木林生长作综合评价，八年生杉木林生长的较适宜工是2m×1m。     

帽儿山地区森林叶面积指数生长季动态研究

叶面积指数(Leaf area index)是描述叶片生长过程的重要参数之一。为探讨我国帽儿山地区落叶阔叶林长时间序列叶面积指数变化规律,利用LAI-2200对帽儿山林场老爷岭试验站12块样地生长季叶面积指数进行测量,使用生长方程对离散LAI值进行拟合,计算不同时间的叶面积指数生长速率和生长季累积叶面积指数,分析不同立地条件下不同林分叶面积指数生长情况,对其动态变化规律进行研究。研究表明:生长季4月到8月,12块落叶阔叶林叶面积指数均随时间呈单峰变化。以杨树有优势树种的样地用Mitscherlich生长方程拟合其LAI效果最优,以色木和白桦为优势树种的样地采用logistic生长方程拟合效果最优,其它样地Gompertz生长方程拟合最优,各样地生长方程拟合R2均高于0.962。杨树林叶面积指数增速最快,胡桃楸林增速缓慢,6月初到8月中下旬为冠层LAI生长速度趋于平稳。空间位置相近的阴阳坡样地叶面积指数生长规律差别较大,最高累计叶面积指数相差17.6%。此研究结果为帽儿山地区阔叶林叶面积指数动态变化规律提供数据基础,为该地区林冠植被的空间异质性及其造成影响,以及提升日步长碳循环、水循环生态机理模型精度提供更为准确的数据支持。     

上层冠层因子对马尾松肉桂复层林下层林分生物量的影响

采用标准地定位监测方法,选取不同栽培模式马尾松(Pinus massoniana)肉桂(Cinnamomum cassia)人工复层林上层林冠的叶面积指数、平均叶倾角、直接辐射透过系数、散射辐射透过系数、消光系数等5个冠层因子作为影响下层林分生长的光环境指标,利用主成分分析法研究影响肉桂林分生物量的主要光环境影响因子。结果表明,不同上层密度光环境下的肉桂各器官生物量变化明显;在主成分分析中,马尾松冠层叶面积指数、平均叶倾角和直接辐射透过系数三者的方差贡献率累计达到99.37%,是影响下层林生长的主导光环境因子;肉桂生物量的主成分回归模型具有良好的拟合度,从回归模型中可得出影响肉桂生物量的主要因子是上层林冠层的直接辐射透过系数。     

应用多重极化L─波段合成孔径雷达数据估测森林生物量、林冠结构和树种组成（上）

本文报道了应用多重极化Ｌ－波段合成孔径雷达（ＳＡＲ）数据估测森林生物量、冠层结构和树种组成的主要研究成果。ＳＡＲ数据与树高、胸径、植株密度、方位和样地几何形状等因素呈显著相关（Ｐ＝０．０５或Ｐ＝０．０１），特别能反映硬阔叶林分子其它森林树种组间林冠特征差异（包括分枝形态、树冠重量和面积）、主要树种组内、树种组间与林分结构的变化（包括树高、植株胸径和密度）以及一些低生物量和部分疏林地与潮湿土壤及枯枝落叶层的分布和裸露程度。     

不同林型对林下光照和辐射消减的影响

在中山市长江库区水源林市级保护区内选择4种不同的林分类型,进行每木检尺和冠层影像拍摄,处理得到林下光照因子和辐射消减因子,分析不同林型对林下光照和辐射消减的影响。结果表明:不同林型对林冠结构和林下光照影响显著,针叶林与针阔混交林的林下光照接近,林冠较差,沟谷季雨林与常绿阔叶林接近,林冠较好;辐射消减在不同林型间差异显著,针叶林和针阔混交林辐射消减较差,常绿阔叶林和沟谷季雨林辐射消减较好;林下光照与辐射消减相关性强,特别是两个林冠结构因子与辐射消减关系密切。在进行林分改造时,建议选择叶片较大较厚,冠幅大而优美的常绿阔叶树种进行更新,天然起源的林分与常绿阔叶林的林冠结构较好,有效维持了森林生态系统稳定。     

林分密度对Ⅰ-69杨树冠结构和光能分布的影响

密植的Ⅰ-69杨林分树冠分枝角度小,骨干枝比率低。当叶面积指数达到7.81、冠层最大叶面积密度达0.8～0.9时,是林分最适密度状态。光在冠层内的分布,受叶面积指数和叶片消光系数支配。密植的林分,光强衰减较快。以8%的相对光强作为临界光强,林龄6年时以高密度林分的生产能力最低;中密度林分最高。但林龄7年时,低密度林分的生产能力已经接近中密度林分。     

二十四年生美国西部黄松生长量和树冠叶面积同下木竞争的关系

利用控制初植密度和下层植物进行长期试验所得的资料,全面证实了我们的假设,即,林分生长与林冠的生长有直接关系,例如,生长量与冠幅成比例关系。无下层植物的黄松林分冠幅生长速度明显快于有下层植物的黄松林分。在叶面积指数非常低、特别是在林分的下层植物已被清除的情况下,森林生长率随着叶面积指数的增加而增加。然而,一旦叶面积指数超过2.0,森林生产率保持稳定,并且有下层植物和无下层植物的林分出现了差异。     

Assessing forest structure and function from spectral transmittance measurements: a case study in a Mediterranean holm oak forest

Annual changes in structural attributes and seasonal dynamics in water content, photosynthetic rate and light-use efficiency (LUE) were assessed by spectral transmittance for 4 years (1999-2003) in six stands of a Mediterranean holm oak forest. Green biomass, total biomass and leaf area index (LAI) were determined. In 1999, seasonal dynamics of net carbon dioxide (CO2) exchange and water content were measured. We recorded photosynthetically active radiation (PAR) transmittance and hyperspectral transmittance in the 400-1100 nm region and derived reflectance-based vegetation indices. Transmittance over the PAR region derived from either ceptometer or spectroradiometer measurements (PART and TPAR, respectively) was related to green and total biomass. Both PART and TPAR were also related to LAI (r=0.79 and r=0.70, respectively, P <0.001) and were appropriate for comparison among stands, whereas subtle changes in LAI within a stand were better assessed by the transmittance amplitude in the red edge region (TRE) (within a stand, r=0.77-0.99, P <0.001). Spectral transmittance-based indices successfully captured physiological processes that occurred on temporal (seasonal) and spatial scales. The transmittance-based water index (TWI) was related to both foliage and canopy water content (r=0.69, P <0.001). Estimates of foliage and canopy water content improved in dense (closed) stands (r=0.84 and r=0.87, respectively, P <0.001) compared with low-density stands. Under non-drought conditions, transmittance-based photochemical reflectance index (TPRI) was related to LUE (r=0.58, P <0.05) and net CO2 exchange (r=0.72, P <0.01), and the combined TPAR x TPRI index greatly improved these relationships (r=0.93 and r=0.84, respectively, P <0.01), indicating that both structural and physiological adjustments modified CO2 fixation capacity in these forest stands. Our novel approach to the study of transmitted radiation provides a tool for estimating structural and functional variables such as LAI, LUE and water content, which are key determinants of terrestrial productivity.     

Canopy light transmittance in Douglas-fir--western hemlock stands

We measured vertical and horizontal variation in canopy transmittance of photosynthetically active radiation in five Pseudotsuga menziesii (Mirb.) Franco-Tsuga heterophylla (Raf.) Sarg. (Douglas-fir-western hemlock) stands in the central Cascades of southern Washington to determine how stand structure and age affect the forest light environment. The shape of the mean transmittance profile was related to stand height, but height of mean maximum transmittance was progressively lower than maximum tree height in older stands. The vertical rate of attenuation declined with stand age in both the overstory and understory. A classification of vertical light zones based on the mean and variance of transmittance showed a progressive widening of the bright (low variance and high mean) and transition (high variance and rapid vertical change) zones in older stands, whereas the dim zone (low variance and mean) narrowed. The zone of maximum canopy surface area in height profiles, estimated by inversion of transmittance profiles, changed from relatively high in the canopy in most young stands ("top-heavy") to lower in the canopy in older stands ("bottom-heavy"). In the understory, all stands had similar mean transmittances, but the spatial scale of variation increased with stand age and increasing crown size. The angular distribution of openness was similar in all stands, though the older stands were less open at all angles than the younger stands. Understory openness was generally unrelated to transmittance in the canopy above. Whole-canopy leaf area indices, estimated using three methods of inverting light measurements, showed little correspondence across methods. The observed patterns in light environment are consistent with structural changes occurring during stand development, particularly the diversification of crowns, the creation of openings of various sizes and the elaboration of the outer canopy surface. The ensemble of measurements has potential use in distinguishing forests of differing ages that have similar stature.     

Photosynthetic utilization efficiency of absorbed photosynthetically active radiation by Scots pine and birch forest stands in the southern Taiga

Absorption and utilization of photosynthetically active radiation (PAR) were investigated in Scots pine (Pinus sylvestris L.) and birch (Betula pendula Roth.) stands that were 41 years old at the end of the experimental period. Canopy depth of the Scots pine stand was about half that of the birch stand (6.5 versus 11.0 m), but absorption of PAR was similar in the two stands. The Scots pine forest canopy, with a leaf area index of 8.9, absorbed 90% of the incoming PAR (APAR), whereas the birch forest canopy, with a leaf area index of 5.9, absorbed 92% of APAR. During maximum foliage development, the upper Scots pine canopy absorbed more PAR than the upper birch canopy (75 versus 66%). The upper, middle and lower layers of the Scots pine canopy contained 37, 48 and 15% of the total needle surface area, respectively. The corresponding distribution of foliage surface area in the three layers of the birch canopy was 50, 30 and 20%, respectively. Measurements of photosynthetic rate were combined with estimates of leaf area index and stand phytomass to determine rates of primary production on a sunny day, a cloudy day, and on an annual basis. The energy equivalents of short- and long-term carbon gain were used with determinations of APAR to calculate photosynthetic utilization efficiency. Throughout the growing season, photosynthetic utilization efficiency of APAR in the upper canopy layer of the Scots pine forest was almost twice that in the lower canopy layer. In the birch forest, photosynthetic utilization efficiency was greater in the lower canopy layer than in the upper canopy layer. In all cases, utilization efficiency was higher in the birch stand than in the Scots pine stand (52 versus 29 J kJ(-1)). Taking account of respiration of the non-photosynthetic parts of each stand (night respiration of needles or leaves; respiration of branches, trunk and roots), estimated utilization efficiency of APAR for net primary production was 11 J kJ(-1) for Scots pine and 19 J kJ(-1) for birch. Solar conversion ratios, expressed as whole-plant net primary productivity per unit of APAR for the growing season, were 0.81 g MJ(-1) for Scots pine and 0.93 g MJ(-1) for birch.     

Analysis of the sensitivity of absorbed light and incident light profile to various canopy architecture and stand conditions

We analyzed the effect of simplifying assumptions in canopy representation of radiation transfer models, comparing modeled diffuse non-interceptance and photosynthetic photon flux density with measurements at different layers of complex pine-broadleaved canopy with large seasonal variation of leaf area index. The most detailed model included clumping of trees (i.e.,?stand density) and a vertical specification of leaf angle distribution and shoot clumping. A less detailed model replaced the vertically specified variables with their means. The most parsimonious model accounted for neither shoot clumping nor stand density. The vertical specification of shoot clumping and leaf angle distribution only slightly improved vertical and seasonal openness and light estimates over using mean values. Further simplification had little effect on total absorbed light but was more risky for estimates of the vertical distributions of openness and light absorbed by the canopy, which will affect photosynthesis estimates due to the non-linearity of photosynthetic light response. Including woody surfaces in winter, when leaf area was low, was essential for reproducing the measurements correctly. A sensitivity analysis showed that ignoring (i)?shoot clumping could result in a substantial overestimation of total absorbed light with errors increasing with decreasing leaf area and (ii) stand density in sparse stands could lead to substantial overestimation of total absorbed light, and the effect is largely independent of leaf area. Also, (iii) the effect of changing leaf angle distribution increased with decreasing leaf area, and was larger and more persistent along the leaf area range with increasing shoot clumping. Overall, accounting for the effect of tree clumping on absorbed light is most important in stands composed of species where leaves are not very clumped (e.g., broadleaved). However, even in forests with highly clumped shoots (i.e., coniferous), an accurate estimation of absorbed light distribution in stands requires incorporation of stand density in the model.     

Canopy development in young sugi (Cryptomeria japonica) stands in relation to changes with age in crown morphology and structure

In young sugi (Cryptomeria japonica D. Don) stands, crown shape (crown length/crown diameter) ratio, average branch inclination, and spatial density of foliage in the crown increased with stand age. Within crowns, foliage distribution increased from the apex downward and, until crown closure, reached a maximum near the crown base. After crown closure, the maximum occurred near the middle of the crown. In each stand, foliage distribution in the canopy showed almost the same vertical change over time as it did in individual crowns. The vertical distribution of foliage in the canopy moved upward with stand age, accompanied by an increase in canopy depth and leaf mass. The shape of the vertical distribution was almost symmetrical between the upper and lower halves in the closed stands, although slightly skewed downward. The logarithm of average spatial density decreased linearly as cumulative leaf mass increased with distance from the top of the canopy. The total cross-sectional area of the crowns exceeded the stand area from the middle of the canopy downward in the closed stands because of crown overlap. However, partly because of changes in crown morphology and structure, the increase in leaf mass with stand age did not always cause more severe crown competition.     

Leaf area dynamics of a boreal black spruce fire chronosequence

Specific leaf area (SLA) and leaf area index (LAI) were estimated using site-specific allometric equations for a boreal black spruce (Picea mariana (Mill.) BSP) fire chronosequence in northern Manitoba, Canada. Stands ranged from 3 to 131 years in age and had soils that were categorized as well or poorly drained. The goals of the study were to: (i) measure SLA for the dominant tree and understory species of boreal black spruce-dominated stands, and examine the effect of various biophysical conditions on SLA; and (ii) examine leaf area dynamics of both understory and overstory for well- and poorly drained stands in the chronosequence. Overall, average SLA values for black spruce (n = 215), jack pine (Pinus banksiana Lamb., n = 72) and trembling aspen (Populus tremuloides Michx., n = 27) were 5.82 +/- 1.91, 5.76 +/- 1.91 and 17.42 +/- 2.21 m2 x kg-1, respectively. Foliage age, stand age, vertical position in the canopy and soil drainage had significant effects on SLA. Black spruce dominated overstory LAI in the older stands. Well-drained stands had significantly higher overstory LAI (P < 0.001), but lower understory LAI (P = 0.022), than poorly drained stands. Overstory LAI was negligible in the recent (3-12 years old) burn sites and highest in the 70-year-old burn site (6.8 and 3.0 in the well- and poorly drained stands, respectively), declining significantly (by 30-50%) from this peak in the oldest stands. Understory leaf area represented a significant portion (> 40%) of total leaf area in all stands except the oldest.     

Ailanthus triphysa at different density and fertiliser levels in Kerala, India: tree growth, light transmittance and understorey ginger yield

Ailanthus triphysa (Family – Simaroubaceae) growth is known to vary in response to different stocking and fertiliser levels. Understorey productivity related to these differences remain elusive, yet are important for optimising the combined production of tree and crop components. A split plot experiment to evaluate the effect of different stocking levels and fertiliser regimes on ailanthus growth, stand leaf area index (LAI) and understorey PAR (photosynthetically active radiation) transmittance was started at Vellanikkara, India in June 1991. Main plot treatments included four densities (3,333, 2,500, 1,660 and 1,111 trees ha⁻¹), replicated thrice. Four fertiliser levels (0:0:0, 50:25:25, 100:50:50 and 150:75:75 kg N:P₂O₅:K₂O ha⁻¹) formed the sub plot treatments. Ginger (Zingiber officinale) was planted as an understorey crop in May 1994 with contiguous treeless control plots. Soil nutrient availability before and after ginger was assessed. Higher densities stimulated ailanthus growth modestly, while fertiliser response of tree and ginger was inconsistent. PAR transmittance below the canopy was related to tree density, LAI and time of measurement. Midday PAR flux having low standard deviations is ideal for evaluating canopy effects on understorey light availability. Ginger in the interspaces exhibited better growth compared to sole crop. Highest rhizome yield was observed in the 2,500 trees ha⁻¹ stocking level, which is optimum for below five year-old ailanthus stands on good sites. It represents 52% mean daily PAR flux or 73% midday PAR flux. Ailanthus+ginger combinations improved the site nutrient capital when ginger was adequately fertilised, despite treeless controls having relatively higher initial soil nutrient availability. This revised version was published online in June 2006 with corrections to the Cover Date.     

不同类型毛竹林植物物种多样性研究

通过对不同类型毛竹林植物物种多样性的研究,结果表明:毛竹林内物种是丰富的,但较大部分物种是脆弱的;乔木层植物物种多样性指数与林下木本植物物种多样性指数相关显著,与草本植物物种多样性指数相关不显著,乔木层物种多样性指数小于林下植物物种多样性指数;木本植物物种多样性指数,竹阔混交林＞竹针混交林＞毛竹纯林,粗放经营竹林＞中等集约经营竹林＞集约经营竹林;林下草本植物物种多样性指数,竹针混交林＞毛竹纯林＞竹阔混交林,中等集约经营竹林＞粗放经营竹林＞集约经营竹林;竹阔混交林向毛竹纯林转型时,林下木本植物部分丧失,而草本植物将可能增加,也可能减少.     

Leaf distribution in large trees and stands of the floodplain forest in southern Moravia

Vertical distributions of leaf dry mass (M(d)) and leaf area (A(f)) were related to relative irradiance (I(r); I(r) above the stand = 1) in closed-canopy, old-growth stands of the floodplain forest in southern Moravia composed largely of Quercus, Fraxinus and Tilia species. Foliage area and mass at any given canopy height were converted to solar equivalent leaf area (A(s)) and mass (M(s)) by multiplying actual values at a given level in the canopy by the relative irradiance at that position. Stand leaf area index (LAI) was 5 (7 including shrub and herb layer), and solar equivalent parameters reached about 25% of that amount. In all species, vertical profiles of both relative irradiance and leaf dry mass to area ratio (LMA) were sigmoidal and the two variables were linearly related. The dominant, upper canopy species had a larger proportion of solar equivalent foliage than suppressed understory species. For individual trees of all species, the upper canopy had a larger proportion of solar equivalent foliage than the lower canopy. Light compensation points at both the leaf and whole-tree level were defined according to leaf or tree position, size and structure. I conclude that optimization of A(s) for forest stands may be used as a basis for determining thinning schedules and evaluating tree survival after damage to tree crowns by various factors.