Growth characteristics and allometry of <Emphasis Type="Italic">Robinia pseudoacacia</Emphasis> as a silvopastoral system component

Demand for goat (Capra hircus) meat in the southeastern USA is steadily increasing as a result of preferences exhibited by ethnic communities. Feeding systems that include fodder trees can be developed to take advantage of the natural preference of goats for browse. Data were collected for 2 years on a 5-year old stand of Robinia pseudoacacia L. to evaluate growth characteristics for goat browse and to develop allometric relationships in a randomized complete block design (intra-row spacing 0.5 or 1 m and coppice height 0.25 or 0.50 m) replicated six times. Allometric equations were derived from destructive harvests of 68 trees. Main branch size was not affected by intra-row spacing when trees were coppiced at 0.25 m; however, when coppiced at 0.5 m, trees spaced at 1 m had more and larger branches (P < 0.05) than trees spaced at 0.5 m. Intra-row spacing did not influence tree herbage biomass when trees were spaced at 0.5 m whereas trees coppiced at 0.25 m and spaced at 1 m produced less woody and herbaceous biomass than trees spaced at 0.5 m (P < 0.05). A strong relationship (P < 0.0001) was found between tree herbage biomass and the number of main branches greater than 0.01 m (r ² = 0.80). The results, based on growth characteristics and tree herbage biomass production, suggest that Robinia pseudoacacia L. would be an excellent candidate as a silvopastoral component in the southeastern USA.     

Soil water competition in a temperate hedgerow agroforestry system in South Africa

An on-farm trial was conducted to determine dry matter production of four fodder tree species and their effect on soil water and maize production. The trees were planted in rows intercropped with maize. The four tree species selected were Acacia karroo Hayne (indigenous fodder tree), Leucaena leucocephala (Lam.) De Wit (nitrogen fixing), Morus alba L. (fodder and fruit), and Gleditsia triacanthos L. (fodder and fuel). Volumetric soil water was measured in the upper 0.3 m of soil in each row of the trial using the time domain reflectometry technique. The neutron probe technique was used for monitoring the water content deeper in the soil. Geostatistical methods were used to analyse treatment differences in the upper 0.3 m of soil. The soil water content did not differ significantly between the maize and tree rows indicating that competition for water in the upper horizon was not the reason for lower maize yields. However, at greater soil depths (75–125 cm) trees in the wide spacing used less water than those in the narrow spacing. Light interception was an important factor in reducing maize yields in the row nearest to the trees. High soil water values recorded during summer indicated that in the current cycle of good rainfall the plants in the agroforestry trial were not stressed. Thus the trees do not compete with the crops for soil moisture in good rainfall seasons. However, this study would need further evaluation for the competition for water for the low rainfall years. Since the trees have access to water at greater depths, they are likely to be more productive into the dry season than shallow rooted crops.     

Soil physical factors affecting the growth of sycamore (Acer pseudoplatanus L.) in a silvopastoral system on a stony upland soil in North-East Scotland

Tree growth and soil physical properties were compared on two grazed plots planted with sycamore at 5 and 10 m square spacing on an upland Scottish site. Both plots received fertilizer and stocking density of ewes and lambs was adjusted to maintain a constant sward height.Mean tree height in 1990 and height increment (1988–90) were significantly greater in the 5 m spaced plot (P<0.05). Although matric potential under the trees was generally greater than in the grassed rows between trees, mean penetration resistance (37–107 mm depth) was significantly greater (P<0.01) under the trees. Even when the soil was close to field capacity, less than 10% of penetrometer readings were < 1 MPa under the trees, in comparison to 44% in grassed areas between trees. This demonstrates that surface compaction due to preferential treading by sheep near the base of trees was sufficient to have seriously reduced tree root growth. Penetration resistance under the 5 m spaced trees was significantly less (P<0.05) than under the 10 m spaced trees. A technique for estimating the probability of root deflections by stones (in 35 mm depth intervals), Ps, from penetrometer readings was used. A significant relationship (P<0.1) was found between tree height increment and depth to Ps0.4 in the 10 m plot.     

The role of complex agroforestry systems in the conservation of forest tree diversity and structure in southeastern Ghana

The role of different agroforestry systems in the conservation of plant diversity and forest structure has not been directly compared in many agricultural dominated landscapes. In this study, we investigated tree diversity and forest structure in a complex agroforestry landscape traditionally grown for cocoa and mixed food crops and compared these to the natural forest in southeastern Ghana. The study was carried out using 36 25 m × 25 m plots. There was significant difference [95% Confidence Interval (95% CI)] in the native forest/non-crop tree species richness between the natural forest and the agroforest farmlands but species richness was similar between the cocoa and mixed food crops agroforests. The density of native forest/non-crop trees was significantly higher (P < 0.05) in the natural forest but similar between cocoa and mixed food crops agroforest. Similarly, the basal area of native forest/non-crop trees was significantly higher (P < 0.05) in the natural forest but comparable between cocoa and food crops agroforest. Of the 20 most abundant native forest/non-crop trees recorded, 12 of them showed significant responses (P < 0.05) to land use change with nine of the species significantly abundant in the natural forest relative to the agroforest systems. Eighteen native forest/non-crop trees species in the agroforestry systems were commonly recorded as being used; 100% of them being used as fuel wood with 83.3 and 77.8%, respectively, used as medicines and materials. The findings of this study suggests that although complex agroforestry systems are a poor substitute for the natural forest the heterogeneous mosaic landscape in which complex agroforestry forms part can be strategically managed to maximize the benefits of both sustainable agriculture production and conservation of plant diversity by acting as buffer between protected areas and intensively managed areas.     

Change in soil mineral nitrogen and respiration following tree harvesting from an agrisilvicultural system in Sweden

The effect of tree harvesting on soil mineral nitrogen and microbiological activity were investigated in an agrisilvicultural system consisting of wheat cultivated along the sides of a poplar plantation in Sweden from 1993 to 1995. Sampling for mineral nitrogen was carried out in three layers down to 90 cm at two distances, near (0.5–1.5 m) and far (4.0–5.0 m) from rows of standing, ST, and harvested trees, HT. Sampling for basic respiration and substrate-induced respiration was carried out in the 0–10 cm layer in 1993 and in the 0–10 cm and 10–20 cm layers in 1994 at the same distances from trees.There was a higher concentration of ammonium and lower concentration of nitrate closer to trees, indicating an efficient uptake of nitrate by trees and enhanced N mineralization close to trees. Shortly after tree harvesting, there were higher concentrations of nitrate and ammonium in the 0–30 cm soil layer near the harvested trees than near standing trees, suggesting a derease in nitrogen uptake by tree roots. The soil microbiological activity was lower in the harvested than the standing plots of trees, which is considered as an indication of the important role of root exudates in maintaining a larger microbial biomass close to trees.     

The establishment phase of a silvopastoral national network experiment in the UK

Silvopastoral agroforestry can satisfy some objectives required of European land-use systems: reduced agricultural production, increased timber production, increased product diversity and environmental enhancement. A national network experiment was set up on four sites, each representative of a UK grassland farming area, with three replicates of common treatments: sycamore (Acer pseudoplatanus L.) at two silvopastoral planting densities [100 (S100) and 400 (S400) stems ha^–1] protected by tree shelters and an agricultural control (ACONT) all with grazing sheep; a woodland control (WCONT, 2500 trees ha^–1) without grazing. Common management protocols were applied and common measurements recorded. Results are provided for the first six years. There were no significant differences between S100, S400 and ACONT in agricultural productivity, though there were significant differences between sites (P < 0.001). There were no significant differences in tree survival between the silvopastoral treatments and WCONT (mean 92.5% ± 0.74) but there was a difference between S100 and S400 (90.8 vs 94.7%: P < 0.001). There were significant differences between the sites (range 86.5–96.2%: P < 0.001) and between the first three years, when replacement of dead trees took place, (82.5, 95.1 and 96.9% for years 1, 2 and 3 respectively: P < 0.001). There were significant differences in the total height of the trees in years two to four between WCONT, S100 and S400 (113.5, 154.1 and 194.5 cm respectively in year four: P < 0.001). However, by year six WCONT and S100 were similar (180.7 ± 17.31 cm) while S400 were taller (219.0 ± 22.80 cm: P < 0.05). It is concluded that tree shelters maintained silvopastoral tree survival at the level of conventional woodland. Tree height extension was compromised on S100 where a higher animal:tree ratio resulted in greater animal activity and soil compaction around trees compared to S400. The site with poorly-drained soil proved to be unsuitable for sycamore-based silvopastoral agroforestry.This revised version was published online in November 2005 with corrections to the Cover Date.     

Organic matter contribution to soil fertility improvement and maintenance in red Alder （Alnus rubra） silvopastoral systems

An investigation was conducted to quantify fine roots and roots nodules over the four seasons in forestry and agroforestry alder (Alnus rubra) stands in North Wales. Soil samples collected in each season were excavated at three sampling points (0.30 m, 0.57 m and 1.00 m distance from the base of each tree) from nine trees of the agroforestry and forestry plots. Result showed that the density of live fine root had significant differences in between seasons and treatments (P ＜ 0.001). The mean weight density of live fine root over the four seasons in agroforestry and forestry was 0.27±0.01 kg·m-3 and 0.54±0.03 kg·m-3, respectively. Weight density of dead root in each system remained constant throughout the year. The mean weight density of dead root was also significantly different (P ＜ 0.01) between forestry and agroforestry systems. Weight density of live and dead root nodule was both constant throughout the year and between the different sampling distances. The mean weight densities of live and dead root nodule over the four seasons were 0.09±0.03 kg·m-3 and 0.05±0.03 kg·m-3 in agroforestry and 0.08±0.02 kg·m-3 and 0.03±0.01 kg·m-3 in the forestry plots, respectively.     

Comparative productivity of Prosopis cineraria and Tecomella undulata based agroforestry systems in degraded lands of Indian Desert

Tree-crop interactions were monitored by measuring tree growth characters of Prosopis cineraria L. and Tecomella undulata L. and yields of Vigna radiata (L) in agroforestry systems in degraded lands of Indian Desert. Potential competition for resource between the trees and associated crop was analyzed by measuring soil water contents, soil organic matters and NH₄-N at different depths of soil layers i.e., 0–25 cm, 25–50 cm and 50–75 cm in the experimental plots. The plots size were 16 m × 18 m (D₁), 20 m × 18 m (D₂) and 32 m × 18 m (D₃) with tree densities of 208, 138 and 104 trees·ha⁻¹ after June 2002, respectively. Results showed that tree height increased by 3% to 7% during June 2002 to June 2004. Collar diameter increased by 30% and 11% in D₁, 23% and 19% in D₂ and 18% and 36% in D₃ plots, respectively, in P. cineraria and T. undulata in two years period. The increase in crown diameter was 9% to 18% in P. cineraria and 11% to 16% in T. undulata. Tree growth was relatively greater in 2002 than in 2003. Yield of V. radiata increased linearly from D₁ to D₃ plots. Lowest soil water content at 1 m distance from tree base indicated greater utilization of soil water within the tree rooting zone. Concentrations of soil organic matters and NH₄-N were the highest (p<0.05) in 0–25 cm soil layer. P. cineraria was more beneficial than T. undulata in improving soil conditions and increasing crop yield by 11.1% and thus more suitable for its integration in agricultural land. The yield of agricultural crop increased when density of tree species was appropriate (i.e., optimum tree density), though it varied with tree size and depended upon resource availability. The result indicated bio-economic benefits of optimum density of P. cineraria and T. undulata over traditional practices of maintaining random trees in farming system in arid zones. Biography: G. Singh (1961– ), male, Scientist E and Head, Division of Forest Ecology, Arid Forest Research Institute, New Pali Road, Jodhpur-342005, India.     

Soil fertility changes and response of maize and beans to green manures of leucaena,sesbania and pigeonpea

Three multipurpose tree species (MPTS)-leucaena (Leucaena leucocephala), sesbania (Sesbania sesban var. nubica) and pigeonpea (Cajanus cajan) were pruned at a height of 60 cm above the ground every two months, and resulting plant biomass was incorporated into the soil as green manure. For comparison, maize (Zea mays) stover was also incorporated into some plots, while some other plots were left fallow. Varying quantities of plant biomass which were incorporated into the soil over a period of 12 months caused large changes in major soil plant nutrients, and it substantially improved soil fertility. To test for improved soil fertility, test crops of maize and beans (Phaseolus vulgaris) were grown on the plots after six biomass incorporations of 4806, 13603, 16659 and 7793 kg. ha^–1yr^–1 for pigeonpea, sesbania leucaena and maize, respectively. Responses of the test crops indicated that sesbania and leucaena green manures improved maize stover, cobs and grain yields; and bean haulms and grain yields by 77.6% when compared to fallow plots. Residual effects of green manures still resulted in significant (P < 0.05) yield differences in the test cropin the third testing season. Economic significance of green manures in increasing food crop yields to small scale farmers is discussed.     

Changes in the stand structure on permanent Picea abies plots managed with single‐tree selection

Eleven permanent plots, established in central and northern Sweden during the years 1913–59, were repeatedly treated with single‐tree selection for between 30 and 67 years. The plots were all dominated by Norway spruce (Picea abies (L.) Karst.), soil moisture was mesic, and the ground vegetation was dominated by Vaccinium myrtillus L. All trees higher than 1.3 m were numbered and cross‐calipered at the time of plot establishment, and at following treatments. In the study trees with DBH < 8.5 cm were excluded from the calculations. Stand densities generally decreased as a result of harvests and mortality together exceeding increment and ingrowth. The number of trees was generally declining over DBH at the time of plot establishment. Six plots maintained such distributions during the whole observation period. Changes in stand structure were mainly caused by the harvests being concentrated to trees with DBH less than 30 cm, and low levels of ingrowth. The low ingrowth was probably an effect of over cutting of saplings and small trees.     

Improving the Traditional Acacia Senegal-Crop System in Sudan: The Effect of Tree Density on Water Use, Gum Production and Crop Yields

The traditional Acacia senegal bush-fallow in North Kordofan, Sudan, was disrupted and the traditional rotational fallow cultivation cycle has been shortened or completely abandoned, causing decline in soil fertility and crop and gum yields. An agroforestry system may give reasonable crop and gum yields, and be more appealing to farmers. We studied the effect of tree density (266 or 433 trees ha⁻¹) on two traditional crops; sorghum (Sorghum bicolor) early maturing variety and karkadeh (Hibiscus sabdariffa), with regard to physiological interactions, yields and soil water depletion. There was little evidence of complementarity of resource sharing between trees and crops, since both trees and field crops competed for soil water from the same depth. Intercropping significantly affected the soil water status, photosynthesis and stomatal conductance in trees and crops. Gum production per unit area increased when sorghum was intercropped with trees in low or high density. However, karkadeh reduced the gum yield significantly at high tree density. Yields of sorghum and karkadeh planted within trees of high density diminished by 44 and 55% compared to sole crops, respectively. Intercropping increased the rain use efficiency significantly compared to trees and field crops grown solely. Karkadeh appears to be more appropriate for intercropping with A. senegal than sorghum and particularly recommendable in combination with low tree density. Modification of tree density can be used as a management tool to mitigate competitive interaction in the intercropping system.     

Using non-destructive testing to assess modulus of elasticity of Pinus sylvestris trees

Abstract

This study assessed variation in modulus of elasticity of trees and logs of Scots pine (Pinus sylvestris) trees. The study used 192 sample trees (c. 90–150 years) selected from 24 clear-felling forests in central and southern Sweden. Modulus of elasticity (MOE) assessed with transit-time technology on standing tree stems at 0.5–2.0m on the southern and northern side of each tree varied from 8.6 to 17.6 GPa. No systematic MOE difference was found between the southern and northern side of tree stems. The sometimes large MOE variations seen in some individual trees are probably a result of wood variation and wood defects. MOE assessed with resonance-based technology varied between 7.4 and 14.1 GPa for logs cut at similar height (<6.0 m). Models of MOE variation were derived from factors related to growth conditions at stand and tree level, with an R ² _adj of c. 0.46–0.62. The models indicate that growth and tree attributes associated with and/or creating less stem taper would yield trees with higher MOE.     

Root distribution of Pinus pinaster, P. radiata, Eucalyptus globulus and E. kochii and associated soil chemistry in agricultural land adjacent to tree lines

We quantified the extent and distribution of roots of four commonly planted tree species (Eucalyptus globulus Labill., Pinus radiata D. Don, P. pinaster Aiton and E. kochii Maiden & Blakely subsp. plenissima C.A. Gardner) in agricultural land adjacent to tree lines, and examined the effect of soil type and root pruning on root morphology. Root distribution in soil adjacent to tree lines was mapped by a trench profile method at 13 sites on the south coast of Western Australia. Soil samples were collected to determine water content and fertility. The lateral extent of tree roots ranged from 10 m for E. kochii to 44 m for P. pinaster. This equated to between 1.5 and 2.5 times tree height (H) for E. globulus and Pinus spp. to 4H for E. kochii. Root density declined logarithmically with distance from the trees and was greatest for P. pinaster and least for E. globulus (P < 0.001). The rate of decrease in root density with distance from the trees was greatest for the Pinus spp. and least for E. kochii (P < 0.05). Root density was generally greatest in the top 0.5 m of the soil profile and decreased with increasing depth. This decrease was relatively gradual in the deep sands, but abrupt in clay subsoil. Root dry mass in the sandy top soil beyond 0.5H ranged between 1.0 and 55.5 Mg km(treeline) (-1) for 6-year-old E. kochii and 50-year-old P. pinaster, respectively. Soil water content generally increased with distance from the trees (P < 0.001). There was no evidence of reduced soil fertility in the top 1.4 m of the soil profile adjacent to the trees. Two to four years after trees had been root pruned, both the lateral extent and vertical distribution of roots were similar for pruned and unpruned trees. The density of roots < 2 mm in diameter was greater for root-pruned trees than for unpruned trees (P < 0.05). We conclude that the study species can compete with agricultural crops based on the lateral extent of their roots and the occurrence of greatest root density within 0.5 m of the soil surface.     

Millet production under pruned tree crowns in a parkland system in Burkina Faso

As a tree management tool, three treatments of crown pruning (total-pruning, half-pruning and no-pruning) were applied to Vitellaria paradoxa (karité) and Parkia biglobosa (néré) in agroforestry parkland systems in Burkina Faso. The area under each tree was divided into four concentric tree influence zones (Zones A: up to 2 m from the tree trunk, B: up to half of the radius of the tree crown, C: up to the edge of the tree crown and D: up to 2 m away from the edge of the tree crown). Millet production under these zones and outside was assessed during two cropping seasons over the study period of three years and the results showed that tree crown pruning had significant effect on millet production and the highest millet grain yield and total dry matter were produced under total-pruned trees (507 ± 49 and 2033 ± 236 kg ha⁻¹ year⁻¹, respectively). Light transmission, transpiration and soil nutrient status under the trees were also analysed in relation to millet production. The results of the analysis showed that total-pruned trees gave the highest millet production due to the reduction by crown pruning of the effects of large tree crowns on PAR transmission below crowns and rates of transpiration by trees. Soil was more fertile closer to the tree trunks than outside tree crowns. This may also be one of the reasons why millet overall performed better under Zone B than outside tree crowns. The higher production of millet under Zone B than under Zone A, the zone closer to the tree trunk, may be due to lower light intensity and more intense competition for water between trees and crops under Zone A. It was concluded that at least in the short term millet production could be improved by crown pruning of both karité and néré, but long term effects may depend on the ability of the trees to maintain the amelioration of soil fertility and on how quickly the trees recover from pruning. This revised version was published online in June 2006 with corrections to the Cover Date.     

Insect (Salebria sp.) infestation and impact on Vitellaria paradoxa C.F. Gaertn. fruit production in agroforestry parklands

Vitellaria paradoxa, known as Shea tree or karité, is one of the dominant trees of agroforestry parklands in Sahelian region with an immense commercial value. In this study, we examined infestation of fruit-bearing shoots and fruits on two sites in western Burkina Faso across different reproductive phenophases, crown heights and aspects. The number of infested and healthy shoots and fruits were counted on 60 reproductively active trees randomly selected from on-year shea trees in each site, using a metal frame (0.5 × 0.5 m) to mark a fixed area of the canopy for sampling. The shoot and fruit borer was identified as Salebria sp. (Lepidoptera:Pyralidae), which was not reported earlier. The proportion of infested trees and shoots varied significantly between sites and among reproductive phenophases (P < 0.0001). The site close to the natural forest had the highest infestation rate than distantly located site. Among reproductive phenophases, the proportions of infested trees and shoots were the highest during flowering and fruit set, respectively. Fruit infestation also varied significantly between sites, crown heights and aspects (P < 0.001). The site close to the natural forest and the crown with north-east aspect were heavily infested while the bottom part of the crown had the lowest infestation. As a whole, 49–80% of trees, 24–31% of shoots and 4–15% of fruits were found infested, depending on the site. These infestation levels combined with precocious flower and fruit abscissions would pose serious economic losses that justify a call for immediate pest control initiatives.     

Land-use impacts on the composition and diversity of the Baikiaea– Guibourtia–Pterocarpus woodlands of north-western Zimbabwe

Variation in floristic composition of recovering Baikiaea–Guibourtia–Pterocarpus woodlands was studied in different development stages (from early regrowth to mature woodland) under different land-use systems (protected areas, timber harvesting, pole and firewood collection, and abandoned crop fields), in the Gwayi and Tsholotsho areas in north-western Zimbabwe. A total of 150 nested circular plots were sampled representatively in four different development stages related to the land-use systems. The DBH (stem diameter at 1.3 m above ground level) and tree height were recorded by species for all stems of tree species with DBH ≥ 15 cm in a 30-m-radius plot (0.283 ha) and for trees with DBH 5.0–14.9 cm in a 11.3-m-radius plot (0.04 ha) (both centred around the same midpoint). Stems with DBH < 5 cm were counted by species in an inner sub-plot of 5.65 m (0.01 ha) radius. Tree data (stem DBH ≥ 5 cm) and regeneration data (stem DBH < 5 cm) by stem counts per species per plot, were used separately to run TWINSPAN (TWo-way INdicator SPecies ANalysis) classifications of species assemblages. Importance values were calculated for all tree species per community. Shannon–Wiener diversity indices were calculated for each community and tested for differences using one-way ANOVA in SPSS version 21. Detrended correspondence analysis (DCA) implemented in the CANOCO ordination software was used to determine the extent of variation amongst the identified communities. The classification identified 12 tree communities and 13 regeneration communities, clustering plots from different land uses together. Baikiaea plurijuga was the most important tree species in all tree communities, except where Combretum collinum, C. apiculatum, Commiphora mossambicensis and Pterocarpus angolensis were dominant. Pterocarpus angolensis showed low importance in most communities, except for communities from undisturbed sites (mostly mature trees) and abandoned fields (mostly young trees). Baikiaea plurijuga was most important in most regeneration communities, except in communities dominated by Baphia massaiensis, C. collinum, C. apiculatum and P. angolensis. Species diversity differed significantly (p < 0.05) amongst tree communities. The DCA ordination showed little variation amongst the communities. The cumulative contribution of environmental factors explaining variation in species composition was 22.6% for tree communities and 26.1% for regeneration communities, suggesting that recovery from disturbance after different land uses may explain more of such variation.     

Comparative performance of seventeen upperstorey tree species associated with crops in the highlands of Uganda

Trials were established at three sites in Uganda to test the suitability of multipurpose trees (MPTs) as upperstorey in crop lands to provide poles, small timber and fuelwood. The three sites were Kachwekano District Farm Institute (1°16 S, 29°57 E, 2000 m.a.s.l.) in Kabale District, Kabanyolo University Farm (0°28 N, 32°27 E, 1250 m.a.s.l.) in Mpigi District and Bushenyi District Farm Institute (0°34 S, 30°13 E, 1610 m.a.s.l.) in Bushenyi District. The MPTs were planted in single rows at intra spacing of 2 m and each plot contained seven or nine trees. On both sides of the tree row, crops were raised. Data on crop yields were collected every season, while data on the growth of the trees were collected four times each year.In terms of tree growth,Grevillea robusta, Casuarina cunninghamiana andAlnus acuminata performed well with height growth of 1.8–2.4 m per year at Kachwekano, while at BushenyiGrevillea robusta, Casuarina junghuhniana, Cupressus lusitanica andCedrela serrulata averaged 1.6–2.0 m height per year. At Kabanyolo,Melia azedarach, Cassia siamea, Jacaranda mimosifolia, Grevillea robusta andMaesopsis eminii had height increments ranging from 1.8–2.7 m annually. Crop yields were affected by the presence of the MPTs, withMaesopsis eminii being the most competitive (averaging 60% reduction, over five seasons). The crop rows nearest to the tree line were the most affected. OnlyAlnus acuminata seems to have had a positive effect on crop yields. The installation of a root mesh to reduce tree root competition for nutrients and water in four species increased yields in plots with MPTs by 5% (Melia azedarach) to 152% (Maesopsis eminii), but the control plot still had significantly higher bean yields, suggesting that shading could also be important. In the case of maize, suppression seems to be due mainly to root competition because after its elimination yields obtained thereafter did not differ significantly from those of the control except for the Maesopsis plots. The negative influence of the MPTs could, therefore, be minimized by periodically pruning the tree crowns and roots.     

Phosphorus Uptake by Birch from Various Depths in the Soil

Potential phosphorus (P) uptake from various depths by treesin a 30-year-old birch stand located at Brathens, Banchory,Scotland, on a brown podzolic soil, was studied using ³²P injectedat 0, 10, 25 and 50 cm depth at 40 points within a 0.5 m radiusof each tree. Two experiments were carried out, the first using8 trees, the second using 16 trees. The ground vegetation hadbeen hand weeded within a 3 m radius of each tree to removecompetition for ³²P uptake. Considerable variation was found in the amount of ³²P in leafsamples within and between replicate trees. Because of this,no statistically significant differences in ³²P uptake weredetected at the 5% level in the first experiment. In the secondlarger experiment, in which a significant (P <lt; 0. 001)depth effect was detected, the results show that ³²P was mainlytaken up from the 0–10 cm depth, with less than 1% ofuptake coming from the 50 cm depth. These results are discussed in relation to the findings fromother studies that the total phosphorus content of surface soilsincreases under developing birch stands. It is concluded thatthese increases are unlikely to result from recycling of P derivedfrom lower soil horizons, but instead from a raising of soilbulk density due to higher rates of organic matter decompositioninduced by birch.     

Soil moisture and water use by pastures and silvopastures in a sub-humid temperate climate in New Zealand

Soil moisture content from 0 to 2 m depth was monitored under 2–6 year old radiata pine (Pinus radiata) with three understoreys of bare ground, lucerne (Medicago sativa) and ryegrass/clover (Lolium perenne/Trifolium spp.) and under adjacent open-grown lucerne and ryegrass/clover pastures. By the fifth year soil moisture depletion/recharge pattern under the trees alone was similar to that under open pasture and under trees with pasture understoreys. Maximum plant available moisture storage was 207–223 mm in the top meter of this Templeton silt loam soil but only 69–104 mm at 1–2 m depth where coarse textures often predominated. Lucerne reduced soil moisture content (SMC) to lower levels during drier summers and extracted more water from 1 to 2 m depth than ryegrass/clover. Evapotranspiration (ET) during early summer when soil moisture was high was close to the Penman potential evapotranspiration (E _p ), but the difference increased when SMC in the top meter dropped below 200 mm. The silvopasture treatments had higher ET in winter than pasture alone but this was still less than E _p . Soil moisture deficits (SMD) at the end of each summer were sufficiently large to require slightly higher than normal winter rainfall and ET < E _p to recharge the soil to field capacity before the next summer. The soil moisture results, taken together with root and growth data, suggest that trees and understorey pastures are complementary in the first three or four growing seasons but this balance subsequently declines in favor of the pine trees. Management options, to extend the period that understorey pastures are productive, include reducing tree stockings, more vigorous pruning, using competitive understoreys and changing from pines to deciduous trees. Research on new silvopastoral combinations is suggested.     

Growth, productivity and quality of Megathyrsus maximus under cover from Gliricidia sepium

The effect of tree canopy on the growth, productivity and forage quality of Megathyrsus maximus and changes in soil properties were evaluated over three seasonal periods. Four adjacent plots (15 m × 17 m each) in a tropical secondary deciduous forest having 12 years of growth and dominated by Gliricidia sepium were randomly assigned to two treatments: removal of trees (SCA) in two of the plots and leaving trees intact (COA) in the other two. In all plots, M. maximus was planted with 50 cm spacing among plants. Tree removal significantly increased the incident photosynthetically active radiation (PAR, P < 0.001) and grass size (12.5 % in height, P < 0.01, and 16.5 % in clump diameter, P < 0.05), but did not significantly affect any other variable. Season significantly affected grass height (P < 0.003), tiller number (P < 0.001), clump diameter (P < 0.001), net CO₂ assimilation rate (P < 0.001), forage biomass production (P < 0.003), and acid detergent fiber content (P = 0.033). Primary soil changes after 1 year of establishment of the grass were the decline by 3 % in organic carbon (P = 0.03), and qualitative changes in soil structure, regardless of tree presence. Results are consistent with the ability of M. maximus to tolerate shade. We conclude that under the conditions of the study there was no evidence for a negative effect of tree canopy on M. maximus mediated by a reduction in PAR.