遮光对野鸦椿幼苗生长和光合生理特性的影响

为了解不同光照条件下野鸦椿幼苗的生长发育状况和光合生理特性,探讨野鸦椿幼苗生长发育的最佳光照条件。采用遮光方式模拟不同光照强度(全光照,遮光30%、60%和90%),研究野鸦椿幼苗的生长、生物量、光合色素含量、生理指标和光合日变化等光合特性的响应规律。结果表明:随着遮光程度的增加,野鸦椿幼苗的生长量均高于对照,以遮光60%的苗高、地径、根长、根幅的增幅最大;遮光30%、60%处理下其生物量积累较多,植株长势良好,叶色较深;遮光处理后,野鸦椿幼苗的叶绿素a、叶绿素b及叶绿素总量(Chla+Chlb)的含量增加,叶绿素a/b值降低;遮光处理使野鸦椿幼苗叶片可溶性蛋白含量降低,使叶片可溶性糖含量增加;遮光30%、60%处理条件下,叶片中MDA的含量显著低于全光照和遮光90%;随着遮光程度的加强,光合速率不断下降,以遮光30%处理下光合速率较高。因此,在育苗生产中,建议有效辐射强度保持在遮光率为30%~60%的范围,可培育出优良健壮的野鸦椿苗木。     

辽东栎苗木早期生长与光的关系

辽东栎 (Quercus liaotungensis Koidz)属于落叶栎类中的一种 ,又称为辽东柞、柴树、小叶青冈。喜光树种 ,耐寒、喜凉爽气候 ,耐干旱瘠薄 ,多生于向阳干燥山坡。辽东栎的分布区位于3 2°0 8′～ 44°0 5′N,1 0 3°0 1′～ 1 3 0°0 2′E,涉及 1 3个省区 ,但主要分布在东北的东部和南部及河北、山西、山东、甘肃、宁夏、青海、陕西、四川的山区 ,朝鲜半岛也有分布。垂直分布范围为海拔80 0～ 2 80 0 m。在华北北部多成大面积次生纯林或与黑桦 (Betula dahurica Pall.)等混生 ,北京地区在山坡形成杂木林或成纯林 ;在河北地区分布在阳光充…     

Effects of nursery shading on seedling quality and post-planting performance in two Mediterranean species with contrasting shade tolerance

In Mediterranean climates, seedlings are frequently shaded in the nursery to avoid heat damage and save water. However, the impact of this shading on the seedling quality and transplanting performance of Mediterranean species is not well known. We studied the effect of nursery shading on pre-planting features and post-planting performance of two Mediterranean tree species: the shade-intolerant pioneer Pinus halepensis and the shade-tolerant late-successional Quercus ilex. We grew one-year-old seedlings of both species under 100, 40 and 5% full sunlight. Shade had a low impact on the morphology and physiology of Q. ilex seedlings. In pines, only the deep shade treatment produced low quality seedlings with poor root development. In both species, transference to high light at planting in autumn did not impose any additional stress than that caused by frosts, but initial root growth was impaired in the two shaded treatments in pine. Post-planting growth and survival of oak seedlings showed no difference between treatments. Pine seedlings grown in deep shade showed higher mortality and lower growth after planting than those grown in full sun and intermediate light treatments, while intermediate light only reduced growth. For the nursery culture of Q. ilex seedlings, we advise using low light levels during summer to save water without impairing field performance. In P. halepensis, seedlings should be cultured under full sunlight conditions to maximize post-planting growth, but they can be cultured under intermediate light without impairing survival.     

Effects of shading and removal of plant parts on growth of Trema micrantha seedlings

Effects of artificial shading and removal of plant parts on growth of Trema micrantha (L.) Blume (Ulmaceae) seedlings were studied. Seedlings were grown in pots in a greenhouse in 45, 30, 10.6, 4.8 and 1.8% of full sunlight. Shading for 60 days had no effect on survival, but it influenced all growth parameters measured. Total biomass decreased with decreasing irradiance, reflecting reductions in dry mass of leaves, stems and roots. In response to shading, allocation of biomass to leaves increased, while allocation of biomass to roots decreased. Specific leaf area, leaf area ratio and leaf mass ratio increased with decreasing irradiance. Decreases in relative growth rate were caused by reductions in net assimilation rate rather than leaf area ratio. Photosynthetic efficiency, as determined by the Fv/Fm ratio (Fv = variable fluorescence, Fm = maximal fluorescence), was unaffected by the shading treatments. Partial removal of leaves, stem or roots did not affect seedling survival. Seedlings responded to removal of plant parts by compensatory growth. Topophysis was observed when the apex was removed: the lateral buds developed only as new plagiotropic lateral shoots; consequently, the decapitated plant ceased height growth and was unable to compete with its neighbors for light.     

遮荫对珙桐幼苗光合特性及生长特征的影响研究

以不同遮荫条件(50%、25%和自然光)下适应生长1 a后的珙桐幼苗为研究对象,测定分析了珙桐幼苗的生理生化指标和生长的变化特征,结果表明:(1)不同的遮荫处理下,珙桐幼苗的净光合速率日变化曲线均为双峰型,有"午休"现象。但不同光强下出现峰值的大小和时间有差异。不同的光强下蒸腾速率日变化趋势也是不一致的,全天蒸腾速率大小顺序为一层遮荫(50%)两层遮荫(25%)全光照。(2)遮荫下珙桐的单叶面积、比叶面积和叶柄长均大于全光照下,表明珙桐利用弱光的能力很强,提高了对有限光资源的吸收。遮荫试验结果说明:适度的遮荫(50%透光率)有利于珙桐幼苗生长。     

6个山茶品种的耐晒性研究

为筛选出耐晒性较强的山茶花品种,以烈香、六角大红、克瑞墨、金花茶、五色赤丹、迪斯6个品种为试验材料,研究不同光照强度（全光照、遮阴率50%、遮阴率70%、遮阴率90%）对植物形态指标及叶绿素含量的影响。结果表明,从生长特性、叶绿素含量、生物量积累等方面来看,全光照条件下品种迪斯、克瑞墨的生长表现最好,不同程度的遮阴处理均影响各品种的生长。最终筛选出品种迪斯、克瑞墨作为耐晒性较强的山茶品种。     

Survival and growth of drought hardened <Emphasis Type="Italic">Eucalyptus pilularis</Emphasis> Sm. seedlings and vegetative cuttings

Forestry requires low mortality of transplanted seedlings. Mortality shortly after planting is often associated with inadequate hydration of transplants. Seedlings can be hardened to the drought conditions they may experience after transplanting by exposing them to controlled drought conditions in the nursery. Eucalyptus pilularis Sm. seedlings were drought hardened by providing nil (severe treatment) or half (mild treatment) the daily irrigation routinely received (control treatment) for up to two non-consecutive days per week during the last 4 weeks of growth in the nursery. Drought hardening reduced stem diameter, seedling leaf area, leaf area per root biomass and seedling quality measured by the Dickson quality index, but increased root:shoot ratio. Hardened seedlings had lower stomatal conductance and leaf water potential on the days they received less irrigation that the control treatment. Hardened seedlings had greater stomatal conductance and were less water stressed than seedlings experiencing drought for the first time indicating hardened seedlings had adjusted physiologically to drought. Survival after transplanting in the controlled drought environment in a glasshouse was enhanced by the hardening treatments. Non hardened seedlings that had had their upper leaves manually removed immediately prior to transplanting to reduce leaf area (top-clipped) had similar survival to hardened seedlings. Stomatal conductance and leaf water potential after transplanting were higher in hardened and top-clipped seedlings than unhardened control seedlings or vegetative cuttings. Survival in the field trial was over 95% for all treatments, possibly as rain fell within 4 days of planting and follow-up rain occurred in the subsequent weeks. Neither the hardened or top-clipped seedlings planted in the field trial had reduced growth, increased propensity to form double leaders or worse stem form than control seedlings when measured at age 3 years.     

Effects of shade on the morphology and physiology of amabilis fir and western hemlock seedlings

Successful regeneration of coastal montane sites harvested using alternative silvicultural systems may depend on the degree to which tree species can acclimate morphologically and physiologically to a variety of light environments. In a study to determine shade acclimation in montane conifers, one-year-old amabilis fir (Abies amabilis (Dougl.) Forbes) and western hemlock (Tsuga heterophylla (Raf.) Sarg.) seedlings were grown in a nursery under four shade treatments: full sunlight (0% shade), 60% and 30% shade using shade cloth, and 30% shade using lath slats. Shading influenced shoot development, foliar physiology and morphological characteristics of both amabilis fir and western hemlock but in general, the effects were small. Shade levels of 60% were required to induce significant acclimation, and western hemlock appeared to respond more positively than amabilis fir and therefore was considered more shade tolerant than amabilis fir. Light quality had little influence on growth and development, as indicated by a lack of significant differences in physiology or morphology between seedlings grown under shade cloth or lath slats. There were indications that adequate nutrition levels may mitigate the effects of shade on seedling morphology and physiology.     

3个与杉木混植木荷种源幼苗对光照响应的差异

目的]探寻不同木荷种源混交林生产力差异的光合机制。[方法]以福建建瓯、江西信丰和浙江龙泉3个木荷代表性种源1年生和2年生幼苗为试验材料,在与杉木混植条件下,设置100%(L0)、50%(L1)和25%(L2)3种光照的光环境,研究3个木荷种源幼苗的生长性状、光合生理及叶绿素荧光特性等对不同光照响应的差异。[结果]1年生和2年生木荷幼苗的苗高、地径和干质量均表现出显著的种源差异,福建建瓯种源幼苗生长优于浙江龙泉种源和江西信丰种源。随着遮阳强度的提高,1年生和2年生2个发育阶段,福建建瓯种源幼苗比叶面积和叶绿素含量等明显增加,叶绿素a/b值、光饱和点、暗呼吸速率和Fo值等均显著降低,表现出较高的形态和生理塑性;相同遮阳处理下福建建瓯种源幼苗总叶面积、叶绿素含量和Fv/Fm值等指标显著高于浙江龙泉种源和江西信丰种源,而叶绿素a/b值、光饱和点、光补偿点和暗呼吸速率则明显低于两种源;长期遮阳和混植状态下福建建瓯种源幼苗可通过较高的生物量分配塑性,增加根系在土壤中分布来响应邻株竞争,进而促进整株干物质的积累;各光环境下,3个木荷种源2年生幼苗Fo、Fm、Fv/Fm和Fv/Fo等叶绿素荧光参数均较1年生幼苗明显升高,但各光环境间差异不显著。[结论]不同木荷种源混交林生产力差异与其对光照的塑性反应能力有关,并随幼苗年龄而变化。     

叶型对马尾松幼苗生长及叶绿素荧光特征的影响

【目的】研究叶型对马尾松幼苗生长及叶绿素荧光特征的影响,以期为马尾松苗木选择及栽培提供参考。【方法】以出圃时呈现为全初生条形叶、全次生针形叶和中间型的马尾松1年生幼苗为试验材料,分析其生长指标、光合色素含量及叶绿素荧光参数,研究初生叶和次生叶叶绿素荧光特征差异及对幼苗生长的影响。【结果】全次生叶幼苗出圃时苗木质量指数(QI)、干物质积累量及造林1年后净生长量均显著高于全初生叶幼苗;次生叶的叶长、体积和表面积等指标均显著大于初生叶;初生叶中叶绿素a、叶绿素b和总叶绿素含量均显著高于次生叶,两者间的类胡萝卜素含量无显著差异。初生叶的F_m、F_v/F_m、F_v/F_o均显著高于次生叶,F_o在2种叶型间无显著差异;初生叶的PSⅡ反应中心参数Y(Ⅱ)和F_v’/F_m’、淬灭参数q_P和NPQ、能量耗散参数Y(NPQ)和Y(NO)及光响应参数ETR_max、α、β和I_k也均高于次生叶,在高光强条件下,初生叶的非光化学淬灭和调节性能量耗散能力增强。由于次生针叶叶长、体积和表面积等形态指标均显著大于初生叶,因而具有次生针叶的幼苗的光合作用面积更大,干物质量积累较多,出圃时的苗木质量以及造林后的生长表现均高于全初生叶的幼苗。【结论】与次生叶相比,初生叶拥有较高的叶绿素含量,较耐光抑制,有较高的光能转化效率,但高光强条件下会通过热耗散的形式来减少多余光能对光合系统的损害;拥有次生叶的幼苗表现出较好的苗木质量及生长特性。     

遮荫对红松、红皮云杉苗木生长的影响

研究不同年龄红松，红皮云杉在全日照、70％、40％、30％及10％光照条件下的生长情况，结果表明，红松、红皮云杉苗木均在相对照度70％的条件下，树高生长量最大。全日照不利于苗木高生长。地径生长随着光照强度的增大而增大。相对照度70％条件下，红松、红皮云杉（小）平均单株生物量最高。红皮云杉（大）在全日照下平均株生物量最高。随着苗龄的增长对光的需要量增大。     

Rehabilitating Abandoned Pastures in Panama: Control of the Invasive Exotic Grass,Saccharum spontaneum L., Using Artificial Shade Treatments

ABSTRACT

The exotic grass, Saccharum spontaneum L., has invaded abandoned agricultural lands in the Panama Canal Watershed for decades. The grass aggressively competes with regenerating tree seedlings preventing natural forest regeneration. To estimate effectual light level for controlling the grass, the growth of S. spontaneum was measured under a range of artificial shading conditions. Five shade treatment subplots–light intensities of 100% (full sunlight), 50, 25, 15, and 5%–were established at each of eight plots in land adjacent to the Panama Canal Watershed. Each site was cleared of S. spontaneum and then the regrowth was harvested. The regrowth of the grass was harvested and measured four times every one and half months for six months. The biomass of S. spontaneum was significantly less in lower light conditions than in full sunlight. The results showed that comparing growth of the grass at each harvest date, except for the first harvest date, there were significant differences between full sunlight and light intensities of 5, 15 and 25%. When compared by different harvest dates for each light level, only full sunlight showed a significant difference in biomass of the grass. The study demonstrates that shading is an effective method for controlling S. spontaneum. The results can be applied to developing reforestation strategies for abandoned lands occupied by S. spontaneum.     

苗期遮阴对凤丹白牡丹光合特性的影响

在大田条件下研究不同遮阴程度对凤丹白(牡丹)苗期光合特性和叶绿素荧光参数的影响。结果表明:在夏季晴天、自然光照条件下,凤丹白苗期的光合速率日变化表现为"双峰型",有明显的"光合午休"现象,净光合速率显著低于遮阴处理;当年生苗遮阴75%条件下,净光合速率没有出现"光合午休"现象,2年生苗遮阴60%条件下,净光合速率显著高于其他处理。自然光照条件下,2年生苗的最大净光合速率和表观量子效率显著高于1年生苗。经过遮阴处理的F_v/F_m均显著高于全光照植株。试验结果说明:夏季大田中午强光、高温条件下,凤丹白幼苗光合作用受光抑制显著,遮阴可有效减轻光抑制,综合比较各参数可知,遮阴60%最适合凤丹白播种苗的生长。     

Growth and photosynthetic responses of Fraxinus mandshurica seedlings to various light environments

To determine light requirement and adaptability of Fraxinus mandshurica seedlings, the seasonal variations of photosynthetic variables were measured in 3-year-old seedlings grown under four light levels (100%, 60%, 30%, and 15% of full sunlight) with a LI-6400 portable photosynthesis system. The leaf chlorophyll content, special leaf weight, annual height and basal diameter increment of seedlings were also observed. The maximum and minimum values of net photosynthetic rate, maximum rate of carboxylation, an...     

Effects of water stress on growth,dry matter allocation and water-use efficiency of a leguminous species, <Emphasis Type="Italic">Sophora davidii</Emphasis>

The adaptation responses to different water conditions and the drought tolerance of Sophora davidii seedlings were assessed in a greenhouse experiment. Two-month-old seedlings were subjected to the following water supplies for 95 days: 100, 80, 60, 40 and 20% of field water capacity. The seedlings at 100% FC had the greatest productivity, height, basal diameter, branch number, leaf number and leaf area. Water supply <80% FC was the threshold of drought-initiated negative effects on seedling growth, yield and physiological processes; these parameters were severely reduced at 20% FC, however, there was no plant death during the experiment. Moreover, water stress decreased leaf relative water content, specific leaf area, leaf area ratio, and water-use efficiency (WUE), whereas it increased the biomass allocation to roots, which resulted in a higher root:stem mass ratio under drought. The S. davidii seedlings tolerated drought by maintaining high leaf relative water content and by reducing branching and leaf expansion. However, low productivity and WUE at 20% FC suggested that seedlings did not produce high biomass under severe drought. Therefore, prior to introducing S. davidii in forestation efforts, a water supply >40% FC is recommended for seedlings to maintain growth and productivity. These results provide insights into limitations and opportunities for establishment of S. davidii in arid regions.     

不同基质对印度紫檀幼苗生长的影响

在黄心土中添加椰糠、泥炭土、塘泥、过磷酸钙和钙镁磷肥进行不同体积比或质量比混合作为育苗基质,培育印度紫檀容器苗,旨在筛选出适宜的育苗基质。结果表明,育苗基质对印度紫檀容器苗苗高、地径、侧根数、侧根长、根瘤菌、生物量、叶面积和叶绿素相对含量等指标的影响非常显著。综合分析评价各项指标,筛选出较适合印度紫檀容器育苗的培育基质为黄心土70%+塘泥30%、黄心土99.5%+钙镁磷肥0.5%、黄心土50%+椰糠50%和黄心土99.75%+钙镁磷肥0.25%。     

Effects of light availability and soil flooding on growth and photosynthetic characteristics of Genipa americana L. seedlings

Genipa americana L. (Rubiaceae) is a late successional neotropical fruit tree used in riparian forest restoration programs. We analyze the effects of light availability and soil flooding on the growth and photosynthetic characteristics of G. americana seedlings under nursery and natural light conditions. Two light levels (full sunlight and shade), and two levels of soil water (flooded and control) were used in the experiment. Flooding induced significant changes in the total seedling biomass (P < 0.01). The differences among water treatments were 70 and 10% at full sunlight and shade, respectively. These changes were explained by alterations in the maximum quantum efficiency of the photosystem 2 (Fv/Fm), light-saturated net photosynthetic rates (A _sat) and intrinsic water use efficiency (WUE). G. americana has high physiological plasticity in relation to the light availability and flooding, although significant interactive effects between high light exposure and soil flooding were observed in several photosynthetic and growth variables. The results highlight the importance of the synergistic effects between physical environmental variables on the establishment and growth of G. americana seedlings. In the practical point of view we can indicate that, in degraded riparian forests subjected to soil flooding, seedlings of this species should be planted under partially shaded environments.     

Effects of artificial shade on early performance of white spruce seedlings planted on clearcuts

Chlorophyll fluorescence, chlorophyll content, growth, and mortality of white spruce (Picea glauca [Moench] Voss) seedlings were monitored for 2 years after planting under three scenarios of artificial shade: no-shade (control), shade in summer only, and shade all year. The shade frames allowed 50–60% light transmission, with limited effects on air temperature, relative humidity, soil temperature, and soil moisture around seedlings. Based on fluorescence yield and chlorophyll content measurements, summer-only shade reduced photoinhibition and photooxidation, especially in summer and fall; extending to all year shading did not further reduce either photoinhibition or photooxidation. Shade tended to reduce seedling diameter and mortality, but after 2 years the cumulative effect on mortality was not statistically significant. Study results support the establishment of white spruce seedlings under partial forest canopy, especially on sites with harsh environmental conditions.     

遮阴处理对金蒲桃生长量及光合参数的影响

以2 a生金蒲桃(Xanthostemon chrysanthus)幼苗为材料,研究了全光照,50%遮阴,70%遮阴,90%遮阴4种遮阴处理对金蒲桃生长特性、生物量分配、光合生理变化的影响,结果表明:与自然光全处理对比,遮光极显著(P<0.01)地抑制了金蒲桃幼苗生物量的积累,遮光处理组叶片数量减少,叶生物量比下降,茎生物量比增加;光补偿点、光饱和点、暗呼吸速率和最大光合速率降低,光能利用效率低于全光照叶,强光下遮阴叶的净光合速率保持稳定.上述结果说明:遮阴处理后,金蒲桃在生长特性、生物量分配和光合参数上表现出对遮阴弱光的适应,表明金蒲桃是一种能广泛应用于各种光照条件的优良景观树种.     

Optimizing nursery and plantation methods to grow <Emphasis Type="Italic">Cedrela odorata</Emphasis> seedlings in tropical dry agroecosystems

Cedrela odorata (Spanish cedar) is a valuable multi-purpose tree which Central American rural communities and farmers give priority to introducing in pastures and home gardens. In order to propose realistic methods for C. odorata production in local nurseries in the dry tropical region of Nicaragua, we studied: (a) the ability of locally collected C. odorata seeds to germinate, (b) seed response to storage under ambient conditions or under cold storage, (c) the effects of irradiance and watering during cultivation on seedling morphology and post-transplantation survival, and (d) the effects of competition from grasses on C. odorata seedlings transplanted to pastures. Seed germination ranged from 55 to 66% and remained constant after 6 months of storage under ambient conditions or cold storage. C. odorata seedling morphology was sensitive to irradiation and watering in the nursery growing period. Deep shade reduced seedling biomass and leafiness and increased specific leaf area and root-to-shoot ratio. Water shortage increased root mass ratio and root-to-shoot ratio and decreased leaf mass ratio. Post-transplantation success was favored by weeding, and was the highest for seedlings grown under deep shade and water restrictions.