THE EFFECT OF LEVEL OF NITROGEN FERTILIZER ON BEEF PRODUCTION FROM GRAZED PERENNIAL RYEGRASS/WHITE CLOVER PASTURES

In a 3 × 3 factorial experiment, the effects of three levels of nitrogenous fertilizer (0, 336 and 672 kg/N per ha) and three grazing intensities on live-weight gains and carcass attributes were measured.
The live-weight gains of the animals (per head) over the season were greater at the low and moderate grazing intensities than at the high; this effect occurred mainly from July to October each year. Gains were low on the no-nitrogen swards from July 1962 to the end of the grazing season and throughout 1963: this was related to the very high clover content of the swards.
In all years, as grazing intensities increased, the carcass weights per animal became lower. Carcass balance, carcass length, depth of longissimus dorsi muscle, fat thickness over the longissimus dorsi, weight of kidney suet and channel fat, and weight of the alimentary tract also reflected the effect of treatment on weight gains.
The results are discussed in relation to the botanical composition and consumption of the herbage, presented in Part I.     

THE EFFECTS OF LEVEL OF NITROGEN FERTILIZER ON BEEF PRODUCTION FROM GRAZED PERENNIAL RYEGRASS/WHITE–CLOVER PASTURES

The treatments in this summer–grazing experiment were designed as a 3X3 factorial; 3 levels of nitrogenous fertilizer, supplying 0, 336 and 672 kg N/ha per annum, and 3 stocking rates. The fertilizer was applied in 6 to 8 dressings at intervals of 3 to 4 weeks. The animals used were fattening Hereford X Friesian steers. The stocking rates were calculated on the basis of data available on herbage production under N and irrigation treatments and varied according to the level of N input. The experiment was laid out in 6 randomized blocks, which were grazed in rotation. Blocks were removed from the grazing cycle in spring and early summer of each of the 3 years (1962—-4) over which the experiment was conducted and cut for conservation: yields were recorded. Details are given of: grazing management; results of studies of soil–N levels; the quantities of herbage removed for conservation: the quantity, chemical and botanical composition of the herbage available for grazing; and herbage consumption.     

THE EFFECT OF NITROGEN, STOCKING RATE AND GRAZING METHOD ON THE OUTPUT OF PASTURE GRAZED BY BEEF CATTLE

An experiment was conducted in 1971 and 1972 to study the effects of two levels of fertilizer N (50 and 504 kg/ha) on the productivity of pastures grazed by young beef cattle. Two stocking rates were imposed at the lower N level and three at the higher. In addition to rotational grazing, set-stocking and an integrated grazing-conservation system were included. The responses/kg N were approximately 1 kg live-weight gain, 19–24 Meal ME and 8–9 kg DM. Maximum yields of 1200–1300 kg live-weight gain/ha and 25,000–27,000 Meal ME/ha were recorded. Productivity of set-stocked pasture was similar to rotational grazing at the high N level, but lower at low N. The integrated system yielded comparable results to high-N rotational grazing. Performances and herbage intakes per animal reflected the stocking rates imposed.     

EFFECT OF HIGH FERTILIZER NITROGEN AND STOCKING RATES ON LIVEWEIGHT GAIN PER ANIMAL AND PER HECTARE

The effect of increasing annual fertilizer N application rate from 400 to 800 kg/ha (357 to 714 lb N/ac) and stocking rate from 5.0 to 7.5 animals per forage ha (2.02 to 3.04 animals per forage ac) on herbage availability, digestible OM intake and live weight gain was investigated in a grazing experiment repeated in 3 consecutive years. A 21×1 day rotational paddock grazing system was used in which the grazing area was increased in the ratio of 1:1.5:3 on two predetermined occasions. Increasing the fertilizer N application rate increased herbage availability by 12% but failed to increase live weight gain. Increasing stocking rate decreased herbage availability per ha and per animal, and also live weight gain per animal, but increased live weight gain per forage ha by 17.6%.     

A FURTHER INVESTIGATION OF THE EFFECT OF NITROGEN AND STOCKING RATE ON THE PRODUCTIVITY OF PASTURE FOR BEEF CATTLE

The liveweight gain was measured of beef cattle on pastures stocked at 5.9 to 11.8 cattle/ha and receiving 50, 182 or 329 kg N/ha in a 12-week season. N levels and stocking rates were so adjusted that similar yields of herbage per animal were expected. Results showed that similar liveweight gains per animal, averaging 1.03 kg/day, occurred at all N levels. Liveweight gain/ha ranged from 547 to 1037 kg in 12 weeks and the corresponding quantities of metabolizable energy utilized were 9890 and 17590 Meal. The results confinn earlier work (4, 7).     

THE EFFECT OF NITROGEN, STOCKING RATE AND FREQUENCY OF GRAZING BY BEEF CATTLE ON THE OUTPUT OF PASTURE

Two successive grazing experiments were conducted over 12 weeks on perennial ryegrass pastures with 50 and 44 young cattle to study the effect of N fertilizer when applied at a daily rate of 1 or 3 kg N/ha. At each level of N two stocking rates differing hy 20% were imposed. At the liigher N level and stocking rate, three frequencies of grazing were imposed. At the stocking rates imposed N tended to reduce the daily liveweight gain per head, but increased the total liveweight gain per ha by from 0.79 to 1.58 kg/kg N. A 20% increase in stocking rate depressed gain per head in both years. In the first year it did not improve gain/ha, hut in the second year gain/ha was increased by 12–17%. There was a tendency for performance per animal and per ha to increase as the grazing cycle was lengthened. In 1969 the highest yield in the whole season was 1880 grazing days and 1260 kg gain/ha. The overall response to N fertilizer was similar to that recorded in other reports, but it is possible that a lack of K had limited pasture growth.     

A PORTABLE-CORRAL TECHNIQUE FOR MEASURING THE EFFECT OF GRAZING INTENSITY ON YIELD, QUALITY AND INTAKE OF HERBAGE

A grazing experiment using four stocking rates of sheep, equivalent to 34, 45, 57 and 68 sheep/ha on an assumed 200-day grazing season, was conducted using a portable grazing corral technique. Plots were subjected to fotir grazing periods between mid May and early September. The effect of stocking rate on herbage yield and quality and the influence of these factors and feed intake on liveweights of the sheep were recorded. The mean daily herbage organic matter available over the trial was 9.6, 5.8, 3.3 and 23% of the total liveweight of the sheep at each of the four stocking rates. The highest grazing pressure led to the production of high-quality herbage, but also led to reduced productivity, low feed intake and liveweight losses. The most lenient grazing pressure failed to provide adequate herbage utilization. Even at the most intensive stocking pressure, only 66% of the herbage available to ground level was utilized in grazings after July. Intake results suggested that sheep of 45 kg liveweight required 1000–1200 g digestible organic matter per day to maintain body weight. Despite the higher in vitro digestibility of herbage on offer at the higher stocking rates, intake was limited through lack of herbage; a high degree of correlation existed between herbage availability and herbage intake over the four stocking rates and at all grazings. It is concluded that the portable corral technique is well suited for grazing studies and the assessment of sward response to varying stocking rates and is particularly useful where facilities for more extensive studies are limited.     

THE USE OF LIVEWEIGHT–GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

A grazing experiment is described in which the liveweight gains of small East African Zebu heifers were measured from Hyparrhenia rufa/Stylosanthes gracilis and H. rufa/Centrosema pubescens swards. The design of this trial enabled animal production to be assessed from variations in liveweight gain since stocking rate was maintained at the same level on both pastures, but the overall stocking rate was varied according to seasonal changes in herbage availability.
Production was measured both by using individual animals within a group as independent units, and by true animal replication. Gains from individual animal blocks were shown to be of value for comparing pasture species, provided that the composition of the grazing group was varied. Problems of conducting and interpreting small–scale grazing trials as means for evaluating pastures are discussed.     

THE EFFECT OF NITROGEN AND STOCKING RATE ON THE OUTPUT OF PASTURE GRAZED BY BEEF CATTLE

An experiment was conducted in two successire years to measure the effect of two levels of fertilizer N, 50 and 300 kg/ha (45 and 270 Ib/ac) on the productivity of pastures grazed by young beef cattle. Two stocking rates were imposed at the lower N level and 4 at the higher level. The responses per kg fertilizer N were approximately 1 kg liveweight gain, 20–24 Meal ME and 8–9 kg DM. Maximal yields of about 1000 kg gain/ha (890 Ib/ac) and 19,000 Meal ME/ha (7700 Mcal ME/acre) were recorded. Animal performance was similar on the low and the high N pastures. There was evidence that the chemical quality of pasture was lower on the low N pasture in the first year, but there was no difference in the second year. The numbers of dung pats per ha and the refusal of herbage due to fouling were reduced by Increasing the stocking rate.     

The effect of surface height of swards continuously stocked with cattle on herbage production and clover content over four years

The long-term influence of sward height from April to July (Phase 1) and from July to early October (Phase 2) on total herbage and white clover production was measured over four years (1988–1991) as herbage accumulation. A subsidiary experiment to determine the influence of leaf area index (LAI) on gross canopy photosynthesis was conducted to aid interpretation of growth from herbage accumulation data. Target sward heights in 0·5 ha plots in two blocks were 5,7 or 9 cm in Phase 1 and 7 or 9 cm in Phase 2, although mean actual heights per phase were slightly higher. Net herbage accumulation (NHA) was measured within mobile exclosure areas over successive two-week intervals. Gross photosynthesis was measured in circular turves removed from the trial area representing a range of LAIs with an at least reasonable clover content. Despite wide differences in mean sward height and herbage mass, NHA and net clover accumulation for a given phase were not generally affected by treatments. Positive effects of grazing at 5 cm in Phase 1 on NHA and clover accumulation later in the year, and of grazing at 7 cm in Phase 2 on NHA in the following spring were sometimes apparent. Gross canopy photosynthesis (g CO₂ m^?2 h^?1) at 1500 μE m^?2 s^?1 and 18–21°C was linearly related to LAI described by 1·003 + 1·165 LAI over the LAI range 0·7 to 4·5. Total herbage and clover growth, interpreted from NHA by a previously described model, was predicted to be marginally lower in shorter swards. Similarity in NHA and clover accumulation between treatments was considered to be because of lower senescene and decomposition, and a higher proportion of new tissue being assigned to lamina growth, despite lower LAI and gross photosynthesis in the shorter swards. It was concluded that stocking intensity in swards continuously stocked with cattle did not have a strong influence on net total herbage and clover growth.     

Variations in herbage mass and digestibility, and the grazing behaviour and herbage intake of adult cattle and weaned calves

Two experiments are described in which groups of lactating cows, non-lactating cattle and weaned calves grazed a sequence of swards varying in maturity and herbage mass under strip grazing management at a daily herbage allowance of 60 g dry matter per kg live weight. Lactating cows ate 43% and 76% more herbage than non-lactating cattle of similar weight in the two experiments but herbage OM intakes per unit live weight by the calves and lactating cows were similar.
Variations in diet digestibility and herbage intake in the lactating and non-lactating cattle with changes in sward conditions were similar. In the first experiment the calves were experienced grazers; the variation in the digestibility of the herbage selected was less in the calves than the adult cattle, but the variation in herbage intake was greater. The calves in the second experiment were younger, and they were inexperienced grazers; their ability to increase herbage intake in response to changing sward conditions was poorer than that of adult cattle, though variations in diet digestibility were similar in all classes.
The practical implications of these results are discussed briefly.     

The effect of cutting for conservation on a grazed perennial ryegrass-white clover pasture

Continuous stocking with sheep at high stocking rates may reduce the content of white clover (Trifolium repens) in mixed grass-clover swards. The present experiment was carried out to investigate the effects on sward production and composition of resting a perennial ryegrass (Lolium perenne)- white clover sward from grazing and taking a cut for conservation. Swards were set-stocked with 25 and 45 yearling wethers ha^?1 either throughout a grazing season, or on swards that were rested for a 6-week period and then cut in early, mid- or late season. In an additional treatment swards were cut only and not grazed. Net herbage accumulation was higher at the lower of the two stocking rates and was marginally increased by the inclusion of a rest period at the high but not the low stocking rate. Clover content was higher at the lower stocking rate and was increased by the inclusion of a rest period by 30% at 45 sheep ha^?1and by 11% at 25 sheep ha^?1 The effect was most marked at the end of the rest period before cutting. When rested from grazing the tiller density of ryegrass decreased although tiller length increased, and clover stolon length, petiole length and leaflet diameter increased though leaf and node number per unit length of stolon decreased; the reverse applied when the sward was returned to grazing after cutting. At the high stocking rate, rest periods in mid-season or later maintained the greatest clover content and marginally increased total net herbage accumulation. At the low stocking rate the timing of the rest period had no significant effect on total net herbage accumulation or on clover content. These results show that the combination of grazing and cutting is of benefit where the stocking rate is high enough to threaten clover survival and limit sheep performance. However, at such a stocking rate, feed reserves are at a minimum throughout the grazing season and so opportunities for resting the sward are probably low.     

Defoliation and productivity of a phalaris-subterranean clover sward, and the influence of grazing experience on sheep intake

Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha⁻¹) and high (22 sheep ha⁻¹) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.
There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward < lightly grazed sward < heavily grazed sward < repeatedly cut sward.
When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.     

Out-of-season management of grass/clover swards to manipulate clover content

Two experiments are described in which the effect of grazing or defoliating mixed swards at different times over winter and spring on clover content and development was investigated. In the first experiment swards were grazed with sheep (to about 3 cm) for a short period in (a) November, (b) November, January and March, (c) March or (d) not at all, in three consecutive years. All swards were grazed intermittently during the grazing season with cattle and cut for silage once each year. Each plot received either 0 or 50 kg N ha^?1 in March. The effect of N fertilizer was to reduce clover content in each summer and clover growing point density in the third year. In two of the three years, treatments involving grazing in March had lower subsequent net annual herbage accumulation compared with the other two treatments and higher clover content in summer of the third year. Reduction in growing point density in all plots during the grazing season was associated with cattle grazing when conditions were wet, suggesting that stolon burial was implicated. Grazing with sheep in November, January and March resulted in significantly more visible (when counted in situ) clover growing points in April in year 2 and more total growing points (counted after dissection of turves) in the third year than the November grazed and ungrazed treatments which had, on occasions, higher grass tiller density. In a microplot experiment, high herbage mass standing over winter was associated with lower potential photosynthesis per unit clover lamina area and lower growing point density in March. Cutting herbage in March to 2-3 cm resulted in higher clover content and higher growing point number per unit stolon length. The latter was significantly correlated with total irradiance and red: far red at the canopy base. Potential photosynthesis of clover was not affected by cutting in March. It is concluded that growing point density can be increased by grazing or cutting during winter or spring. However, in order for these new stolons to contribute to clover yield during the summer, they have to be maintained until then by ensuring that competition from grass is minimized by keeping the sward short in winter and spring and avoiding the burial of stolons during grazing.     

THE EFFECT OF HIGH NITROGEN LEVELS AND STOCKING RATES ON MILK OUTPUT FROM PASTURE

An experiment using 40 lactating dairy cows was conducted for 3 successive years to examine the effects of fertilizer N applied at 400 and 700 kg/ha (358 and 627 lb/ac) at stocking rates of 4.94 and 7.41 cows/ba (2 and 3 cows/ac). The animals were grazed in groups of 10 per treatment, using a fixed rotational grazing system, for 22 weeks in each year. Milk ontput/cow and/ha was significantly affected by stocking rate, the mean milk outputs being 2499 and 2218 kg/cow (5498 and 4880 lb/ cow) and 12313 and 16396 kg/ha (11032 and 14691 lb/ac) at the lower and higher stocking rates, respectively. Milk yield/cow at the lower stocking rate was not affected by N level. At tbe higher stocking rate, increasing the amount of N resulted in a 7.4% increase in milk output/cow and/ha. This response to N declined from 1.7% in the first year of the trial to 2.4% in the third year. The response in milk yield to N appeared to he greatest during the first part of the grazing season. Milk composition, liveweight change, and herbage availahilities and intakes were also recorded.     

Effects of field drainage on the growth of herbage and the liveweight gain of grazing beef cattle

A long-term field grazing experiment was begun in 1982 to examine the impact of efficient field drainage on herbage and animal production from swards on an impermeable clay loam in the south-west of England. Drained and undrained lysimeter plots (each of 1 ha) were established on the existing permanent sward and received annual applications of fertilizer N of 200 or 400 kg ha⁻¹. Similar plots were initially ploughed and reseeded with Lolium perenne (cv. Melle), and received fertilizer N at an annual rate of 400 kg ha⁻¹. All plots were continuously stocked by beef cattle and stock numbers were adjusted to maintain a constant sward height and to avoid poaching damage. Results for the first 5 years show that the benefits from drainage were modest and, for beef production, unlikely to pay for its costs over the shorter term. The main benefit was in spring when herbage dry matter yield was 11% greater on the drained plots, but with no significant interaction with fertilizer N level or sward type. This benefit was reduced to 3% on an annual basis, due to the effect of the larger soil water deficits sustained by the drained swards in mid-season. Drainage increased the annual liveweight gain per ha by the grazing cattle by 11%. Possible mechanisms accounting for these effects are discussed in relation to the influences of seasonal patterns of weather.     

Turnover of grass laminae and white clover leaves in mixed swards continuously grazed with steers at a high-and low-N fertilizer level

Turnover rates of grass laminae and clover leaf tissue were estimated over a range of intervals within three periods each year in the second to fourth years (1983-85) of a trial involving swards continuously grazed by steers and receiving either 60 kg N ha-1 in spring (60N) or 360 kg N ha^?1 throughout the year (360N). Within the 60N swards initial stocking rates at turnout were low (60N LS) at 7-2 steers ha^?1 and high (60N HS) at 90 steers ha^?1 in 1983, and in 1984 and 1985 corresponding rates were 10-8 and 13-5 ha^?1. The 360N swards were initially stocked at turnout at 96 (360N LS) and 120 (360N HS) steers ha^?1. Stocking rates were reduced by 33% in midsummer except for 60N in 1984 and 1985 when they were reduced by 50%. Meaned over 3 years, 360N HS had lower herbage mass than 60N LS. Tiller density in 360N was almost 50% higher than in 60N and clover growing point density was only one quarter that of 60N with the 60N LS having lower clover densities than 60N HS in 1985. Generally, leaf extension rate per tiller was higher in 360N than 60N and, when significant, 60N LS had higher senescence rates per tiller than 360N HS. Rate of increase in new clover lamina tissue per stolon was not affected by treatments, whereas in 1983 LS had higher senescence rates of clover laminae than HS. Petiole growth per stolon was higher in LS than HS in 1983 and 1984, the mean over these years for 360N HS being 77% that of 60N LS. Petiole senescence per stolon was lower in 360N HS than 60N LS only in 1983. When comparing 60N HS and 360N LS (representing similar levels of grazing intensity, having similar herbage mass) the gross growth of leaf material in the former was 75% of the latter, in contrast to 57% for net growth. Clover contributed 18% to the estimated growth of leaves compared to a mean of 7% in herbage mass. Taking inflorescence and pseudostem into account in 1984 and 1985,60N HS had 7% clover in standing herbage and 14% in net growth. Therefore, the contribution of clover to growth is considerably higher than its presence in herbage mass would suggest in continuously grazed swards. It is concluded that low-N swards, owing to their lower tiller density and slower grass leaf extension rate, will be less efficiently grazed than swards at higher N levels at a given herbage mass, but the presence of clover will partly offset that disadvantage.     

A COMPARISON BETWEEN SPRING AND AUTUMN PASTURE FOR BEEF CATTLE AT EQUAL GRAZING PRESSURES

Two comparisons between spring and autumn pasture for beef cattle were made. Animals used in all comparisons were of similar breed and weight and were subjected to the same feeding regime for 6–8 weeks before turn-out. The pastures were grazed on an equal grazing pressure basis between season comparisons. The pastures received similar rates of fertilizer N between seasons and had similar lengths of rest period for regrowth. Intakes of digestible OM were greater per unit of metabolic liveweight in spring than in autumn. Daily liveweight gains in spring were high, being 1·09 and 1·37 kg (2·4 lb and 3·0 lb), but were lower from autumn pasture at 0·98 and 0·71 kg (2·2 lb and 1·6 lb). Weather was implicated as a factor affecting daily gain from autumn pasture. Greater herbage yields in spring supported 42 and 204 more grazing days per ha which together with the greater gains per animal supported 80–120% more liveweight gain/ha.     

Comparison of herbage production under continuous stocking and intermittent grazing

The possibility of increasing the herbage utilized over a grazing season was investigated in a study comparing continuously stocked steady-state swards maintained at optimum height (3.5 cm) with intermittently grazed swards. The intermittent systems were designed (a) to allow periodic increase in leaf area and hence growth rate, (b)to ensure that the accumulated herbage was eaten before it senesced, and (c) lo retain high tiller density by alternating periods of herbage accumulation with periods of continuous stocking. Two treatments (no animals or animal numbers reduced to half those on the 3.5 cm steady-state treatment) were used during the 17-18-d periods of herbage accumulation. Grazing down was completed in 3–4 d, after which two treatments (14 d or 28 d) were used for the intervening periods of continuous stocking when sward height was maintained at 3.5 cm. Herbage production was estimated using the tissue turnover technique, with tiller population densities and rates of growth, senescence and net production per tiller measured at frequent intervals. Intermittent grazing treatments where animals were removed during herbage accumulation resulted in changes in tiller size and number, and in growth rates, but not senescence rates, per tiller such that short-term deviations in the net rate of herbage production occurred compared with the continuously stocked control. The periods of advantage during phases of herbage accumulation were counterbalanced by those of disadvantage during the subsequent steady-state phases. Where animal numbers were reduced during herbage accumulation, sward conditions differed little from those of the continuously stocked control, implying that intake per individual animal was increased. It was concluded that intermittent grazing systems offered no advantage over simpler continuous stocking systems, provided that a flexible approach to conservation was incorporated to allow control of sward conditions on the grazed area.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK: 6. The effect of reducing stocking rate and application of fertilizer nitrogen in Wales

Abstract The implications for the agricultural productivity of the UK upland sheep systems of reducing nitrogen fertilizer application and lowering stocking rates on perennial ryegrass/white clover swards were studied over 4 years at a site in Wales. The system involved grazing ewes and lambs from birth to weaning on swards maintained at a constant height with surplus herbage made into silage, thereafter ewes and weaned lambs grazed on separate areas until the onset of winter with adjustments to the size of the areas grazed and utilizing surplus pasture areas for silage. Four stocking rates [SR 18, 15, 12 and 9 ewes ha^?1 on the total area (grazed and ensiled)] and two levels of annual nitrogen fertilizer application (N 200 and 50 kg ha^?1) were studied in five treatments (N200/SR18, N200/SR15, N50/SR15, N50/SR12 and N50/SR9). Average white clover content was negatively correlated with the level of annual nitrogen fertilizer application. White clover content of the swards was maintained over the duration of the experiment with an increasing proportion of clover in the swards receiving 50 kg N ha^?1. Control of sward height and the contribution from white clover resulted in similar levels of lamb liveweight gain from birth to weaning in all treatments but fewer lambs reached the slaughter live weight by September at the higher stocking rates and with the lower level of fertilizer application. Three of the five treatments provided adequate winter fodder as silage (N200/SR15, N50/SR12 and N50/SR9). Because of the failure to make adequate winter fodder and the failure of white clover to fully compensate for reduction in nitrogen fertilizer application, it is concluded that nitrogen fertilizer can only be reduced on upland sheep pastures if accompanied by reduced stocking rates.