THE DIGESTIBILITY AND VOLUNTARY INTAKE OF S22 AND H.1 RYEGRASS, S170 TALL FESCUE, S48 TIMOTHY, S215 MEADOW FESCUE AND GERMINAL COCKSFOOT

Pure swards of S22 Italian ryegrass, H.1 hybrid ryegrass, S170 tall fescue, S48 timothy, S215 meadow fescue and Germinal cocksfoot were each harvested at several stages of maturity during first growth in the spring, and in the subsequent regrowths. Each batch of herbage was fed to sheep in a digestion trial under conditions of voluntary intake. Results from the first growth herbages showed the expected fall in digestibility with increasing maturity. However, the digestibility of S22, H.1, S170 and S48 began to fall well before ear emergence, whereas the digestibility of Germinal fell only slowly up to the time of ear emergence, as had been found previously with S24 and S23 ryegrass and S37 cocksfoot. The rate of fall of digestibility in H.1 and S48 (0.4 units/day) also appeared to be lower than with the perennial ryegrasses and cocksfoot (0.5 units/day). The monthly regrowths within each herbage were of similar digestibility, but the mean digestibility of regrowths differed between herbages, being highest for S215 and lowest for Germinal and S170. The grasses with highest digestibility in the early Spring, in particular S22 ryegrass, were not, however, always the most digestible in the subsequent regrowths. The numbers of sheep fed on each herbage were insufficient to show whether there were significant differences in voluntary intake between different herbage species of the same digestibility. It is suggested that an intake/digestibility relationship, based on data from more animals, should be developed for each herbage species. Only if different relationships are found can it be concluded that factors other than level of digestibility have an effect on intake.     

INTER-VARIETAL COMPETITION IN PERENNIAL RYEGRASS SWARDS

Pure cultures and mixtures of equal numbers of plants of Irish. New Zealand and S23 perennial ryegrass were grown in artificially constructed swards under field conditions. They were cut every time they reached grazing height for three successive years. Counts of plant numbers and tiller numbers per plant made each November show that S23 suppressed both plant number and tiller number per plant of Irish ryegrass and to a lesser extent of New Zealand perennial ryegrass. Its own plant number and tiller number per plant were improved in these combinations. New Zealand perennial ryegrass suppressed plant and tiller numbers of Irish perennial ryegrass.
Varying the method of cutting to match the growth of each variety resulted in improved plant survival and tiller number per unit area of Irish perennial ryegrass when cut according to the growth of S23, i.e. late in the spring. The importance of the effect of experimental technique is discussed.     

THE EFFECT OF COMPANION GRASSES ON TIMOTHY PRODUCTION IN SWARDS CUT FOR CONSERVATION

To assess the effect of companion grasses on timothy productivity, swards of timothy sown alone or with one of eight companion grasses were harvested as for silage four times per year for two years. The companion grasses were: diploid perennial ryegrasses S24 and S23; tetraploid perennial ryegrasses Reveille and Taptoe; meadow fescues S215 and S53 and cocksfoots Trifolium and S143. Averaged over the N application rates (226, 339 and 452 kg N/ha per year) the total herhage yields of timothy/companion-grass mixtures were all significantly superior to that from timothy alone in the first year. Only perennial ryegrass S24 and the cocksfoot mixtures significantly outyielded timothy alone in the second year. Winterkill hecame evident in April of the second year and pure timothy and timothy/meadow-fescue swards were least affected. Over all swards, timothy DM increased hy a mean of 19% from the first to the second year, compared with a fall of 20% in total herhage DM. The contrihution of timothy herhage to the mixtures was very low with companions other than meadow fescue. Late-heading varieties were more compatihie than early-heading varieties, and tetraploid ryegrasses were more compatihie than diploid ryegrasses. It is concluded that if the inherently valuahle characteristics of timothy are to he exploited, it should not he sovm with aggressive companion grasses.     

A COMPARISON OF LEAF AND TILLER GROWTH IN SEVEN PERENNIAL GRASSES AS INFLUENCED BY NITROGEN AND TEMPERATURE

Leaf and tiller development along the main shoot of cocksfoot, ryegrass, timothy, Hordeum bulbosum , meadow fescue, S170 tall fescue and an Algerian tall fescue are described. Comparisons between plants grown at two levels of applied nitrogen, and in a heated and an unheated glasshouse, from January to April, 1961, showed that timothy, cocksfoot and, to a lesser extent, H. bulbosum and ryegrass, generally developed leaves faster than the fescues. Numbers of live leaves per shoot, and numbers of actively-elongating leaves per shoot declined in a similar varietal order. Despite this, the large leaves of the fescues resulted in their having a leaf area per main shoot equal to, or exceeding that of, the other grasses for most of the experiment. In general, rate of leaf appearance, number of live leaves, number of actively-elongating leaves and leaf length were increased by a high level of N and high temperature, but high tesnperature reduced leaf width and, in some instances, tiller numbers.     

A COMPARISON OF THE REACTION OF DIFFERENT GRASS SPECIES TO FERTILIZER NITROGEN AND TO GROWTH IN ASSOCIATION WITH WHITE CLOVER

Seven species or varieties of grass, and a mixture of 3 of them, were sown in pure swards and treated with 4 levels of nitrogenous fertilizer (0,17·5, 35 and 70lb N per acre per cut). Each species, and the mixture, was also sown with white clover. The experiment was cut 4 or 5 times per year. The effect of fertilizer on the yield of each grass was compared with the effect of clover on the yield of the grass/clover swards. Mean annual yields showed an approximately linear response to N; there was a small but significant fall in response to the highest level of N. Response among the species ranged from 20 to 30 Ib of dry matter per Ib N applied for the intermediate level of fertilizer and from 14 to 23 Ib for the final increment of fertilizer.
S37 cocksfoot, S48 timothy, S24 ryegrass, and a mixture of these grasses, were high yielding and responded well to fertilizer N; Irish ryegrass and Agrostis tenuis were less productive and gave poorer responses to N. S215 meadow fescue and S23 ryegrass were intermediate in yield and response.
There were no significant differences between the annual yields of the 8 grass/clover mixtures; the yields of the grass and clover components of each mixture were inversely related. The effect of clover on the yield of the grass/clover mixtures was estimated to be equivalent to the effect of an annual application of 205 lb N per acre to Agrostis tenuis and 120 lb N to S48 timothy. The fluctuations in annual yields were greater with grass/ clover mixtures than with grass swards receiving N.
The yields of grasses when sown with clover were in similar order to their yields when sown pure; but whereas the latter tended to fall from year to year, the yields of the grass components of mixtures (except Irish ryegrass) did not.     

COMPARING THE OUTPUT FROM MIXTURES BASED ON PERENNIAL RYEGRASS AND ON MEADOW FESCUE

Herbage and livestock production from swards based on meadow fescue and on perennial ryegrass were measured over three years. The meadow fescue mixture had no practical advantages in terms of either annual output or seasonal distribution of production. This may have been due to inefficient methods of measurement or may have been peculiar to the management imposed, but it is considered that it was due to the depressing effects of timothy and cocksfoot on the meadow feseue.     

EFFECTS OF WHITE CLOVER ON THE BOTANICAL COMPOSITION OF SWARDS OF RYEGRASS, TIMOTHY AND MEADOW FESCUE AND MIXTURES OF THESE THREE

Results are presented to show the effect of white clover (Trifolium repens) and of applied N on the hotanical composition of swards sown with perennial ryegrass (Lolium perernne), timothy (Phleum pratense), meadow fescue (Festuca pratensis) and mixtures of two and three of these species. White clover had a more favourahle effect on the growth of perennial ryegrass than on either timothy or meadow fescue both in terms of yield and in terms of tiller numbers. Tiller numbers of rough-stalked meadow grass (Poa trivialis) were increased and tiller numbers of Agrostis species were reduced by application of N and by inclusion of clover. Possible reasons for these effects are considered.     

SOME EFFECTS OF GRAZING MANAGEMENT ON THE YIELD AND ITS COMPONENTS OF SOME PASTURE GRASSES

A study was made to determine the effects of grazing to a height of 1 in. when the swards reached heights of 3 and 9 in. on the dry matter production, LAI, tillering and rate of leaf production of new and old tillers in the spring–summer and autumn–winter seasons of 3 pasture species growing in association with white and red clovers. In both seasons the herbage yield under 9–1 management was higher than that under 3–1 and was significantly greater in the spring-summer season. Differences in DM production between cocksfoot, tall fescue and Ariki ryegrass failed to reach significance.
Light utilization under the 2 management systems was considered to be inefficient. In the autumn-winter period there was a linear relationship between the LAI and DM production in all treatments.
The rate of leaf production per tiller was significantly higher in cocksfoot than in ryegrass and tall fescue in both seasons.
New tillers had a significantly higher rate of leaf production than old tillers in the spring-summer period, but not in autumn-winter. The numbers of grass tillers and rooted nodes of clover were significantly higher under 3–1 than under 9–1 and were influenced by season.     

THE EFFECT OF COMPANION GRASS AND SEED RATE ON THE PRODUCTIVITY OF A TALL-FESCUE SWARD

The effects of sowing from 10 to 50 lb/acre of tall-fescue seed on plant establishment and on the annual and seasonal distribution of DM yield of tall fescue and its companion grasses were measured through two growing seasons. Increasing the seed rate increased plant numbers per unit area and decreased per cent establishment. Adding Scots timothy as a companion grass depressed both plant number and per cent establishment. S37 cocksfoot and New Zealand perennial ryegrass reduced both still further. Total annual dry-matter production was relatively unaffected by varying either the seed rate or the companion grass. The contribution of tall fescue to total production was highest when sown alone and was successively reduced by timothy, perennial ryegrass and cocksfoot. When sown alone there were only slight differences in contribution of tall fescue due to seed rate. With any companion grass the contribution from tall fescue increased with increasing seed rate. Early growth was not reduced by modifying seed rate but was reduced by all the companion grasses. It is concluded that increasing the seed rate of tall fescue above 30 lb is not justified, that Scots timothy can be sown at 3 lb/acre with tall fescue without affecting total production or early growth of the mixture, with the advantages of control of unsown species and improved palatability of the herbage.     

A COMPARISON OF BROADCAST AND WIDE-ROW SPACED GRASSES WHEN MANAGED FOR FOGGAGE PRODUCTION

A small-plot trial comparing perennial ryegrass, cocksfoot, timothy and meadow fescue sown broadcast and also in 21 inch rows was conducted at Auchincruive from 1954 to 1957. The cutting treatments were arranged so that measurement could be made of both summer and winter production.
Of the four species, cocksfoot was the most suitable for foggage production. Perennial ryegrass suffered severely from winter killing. Timothy and meadow fescue, although persistent, did not yield as much as cocksfoot.
Total dry-matter production over three years was higher from the cocksfoot rows than from the broadcast stand. For ryegrass and meadow fescue, broadcasting gave higher yields than row sowing while timothy showed no significant difference between sowing methods.
During the winter, all species showed a loss of dry matter. Losses for ryegrass, cocksfoot, timothy and meadow fescue were respectively 34%, 17%, 18% and 24%. Broadcast stands averaged 11% greater loss than the rows.     

A comparison of diploid and tetraploid perennial ryegrass and tetraploid ryegrass/white clover swards under continuous sheep stocking at controlled sward heights. 1. Sward characteristics

Two 1·0 ha plots of a late-heading diploid perennial ryegrass (var. Contender) and a late-heading tetraploid ryegrass (var. Condesa), and two 1·4 ha plots of the tetraploid with Aberystwyth S184 small-leaved white clover, were direct sown in May 1987. Over the three years 1988–90 they were continuously stocked by Mule ewes with Suffolk-cross twin lambs, from early April to the end of August, at a target sward surface height (SSH) of 4–6 cm on one set of plots (constant swards) and, on the other set, al 4–6 cm rising after June to a target 6–8 cm (rising swards). The heights were achieved by variable stocking. Fertilizer N was applied only to the grass plots at the rate of 150- 180kgN ha^-1 annually.
SSH was mainly within the target 4–6 cm, after higher initial heights at turnout in 1988and 1990. Mean heights of the constant swards (April- August) averaged 5·53, 4·43 and 5·04cm in the three years. The rising swards (July-August) increased in height over the constant swards by an average of 0·88, 0·48 and 0·55 cm, in successive years.
Clover content of the herbage mass dry matter in the grass/clover swards increased over each grazing season to average 13·0, 26·5 and 21·2% in the three years, with a high mean stolon density of 130 in m^-2 in August 1990. Ryegrass tiller densities in year 3 were 23% higher in the diploid than in the tetraploid swards, which had 43% more than the 10000 tillers m^-2 of the tetraploid ryegrass/clover swards.
It is concluded that the combination of a densely stoloniferous small-leaved clover with the open growth habit of a tetraploid ryegrass can achieve swards of high clover content under continuous sheep stocking.     

THE USE OF PERENNIAL RYEGRASS SOWN AT 1 LB. PER ACRE IN A STANDARD SEEDS MIXTURE

An attempt was made in a survey-type trial to combine perennial ryegrass with cocksfoot, meadow fescue and timothy together in the same sward, without the association being dominated by the ryegrass. It is concluded from this work that such non-dominant ryegrass swards can be obtained by limiting the seeding rate of the ryegrass to 1 lb. per acre in a seeds mixture sown at the rate of about 20 lb. per acre.     

Sward growth under cutting and continuous stocking managements: sward canopy structure, tiller density and leaf turnover

The change in structure of continuously grazed versus infrequently cut swards of perennial ryegrass ( Lolium perenne L), cv. S23, was investigated during their first full harvest year. Measurements were made from early May until late September. The intensity of stocking by sheep in the grazed sward was adjusted in an attempt to maintain a high level of radiation interception and the cut sward was harvested at approximately monthly intervals.
The herbage mass, lamina area index and radiation interception of the cut sward varied in a cyclic pattern between harvests but in the grazed sward these parameters showed considerably less variation, although they all increased early in the season and then declined later. The proportion of dead material above ground increased throughout the season in both sward types but was more marked in the grazed sward.
There were major differences between the grazed and cut swards in the number of tillers per unit ground area; the difference became more marked throughout the season and by September the tiller densities in the grazed and cut swards were 3·20⁴ m^-2 and 6·20³ m^-2 respectively. Divergence in tiller density was associated with differences in specific stem weight and leaf area per tiller.
Rates of appearance and death of leaves on tillers in the grazed sward were determined. During May, leaf appearance exceeded leaf death but this was reversed in June. During the rest of the season as a new leaf appeared on a tiller so the oldest leaf died.     

A COMPARISON OF THE REACTION OF DIFFERENT GRASS SPECIES TO FERTILIZER NITROGEN AND TO GROWTH IN ASSOCIATION WITH WHITE CLOVER. II. YIELD OF NITROGEN

Seven species or varieties of grass, and a mixture of 3 of them, were sown in pure swards, treated with 4 levels of nitro-chalk (0, 17·5, 35, and 70 Ib N/ac/cut) and cut 4 or 5 times each year. Each species and the mixture were also sown with white clover, and the effect of fertilizer N on the yield of N in each grass was compared with the effect of clover on the yield of N harvested from the grass/clover swards.
The regression line for response in yield of N with increasing levels of fertilizer N showed slight, but significant, upward curvature. The grasses differed in their uptake of N from the soil, S37 cocksfoot and S48 timothy showing relatively high uptakes, and the ability of each grass to take up fertilizer N was usually related to its uptake of N from the soil. Differences in the yield of dry matter between the species, at a similar level of N, are discussed, and it was concluded that perennial ryegrasses were most efficient and Agrostis tenuis was least efficient in using the N taken up in the production of DM.
In general, there were no significant differences between the yields of N of the grass/clover mixtures; the N yields of the grass components were significantly different and tended to be inversely related to the N yields of clover.
Grasses which gave high yields of N with fertilizer were also high yielding when grown in association with clover. Pure grass swards required more than 200 Ib fertilizer N/ac/yr in order to yield the same amount of N as the grass/clover swards. The amount of N estimated to have been derived by grass from clover (indirect effect of clover) increased each year; it was highest with S37 cocksfoot and lowest with Irish perennial ryegrass, averaging 46 and 23 Ib N/ac/yr, respectively.     

The value of timothy (Phleum pratense L.) in ryegrass — timothy mixtures managed to simulate intensive grazing

The performance of timothy in mixtures with perennial ryegrass was assessed under a simulated intensive grazing management over two harvest years in 1974–75. Three seed rates of S23 perennial ryegrass were factorially combined with three rates of Scots timothy and compared with pure stands of each grass. All were sown with Huia white clover. When cut six times at monthly intervals and with an annual N input of 350 kg ha^?1, there were no significant differences in total DM production in either year. The 2-year mean DM yield for the nine mixtures and six pure swards was 9·77 t ha^?1 (range 9·34–10·16). Compared with the pure ryegrass swards, in both years the ryegrass-timothy mixtures produced earlier spring growth but were significantly lower yielding at the second cut. Over the first five cuts the proptortion of timothy in the three mixtures with 22·4 kg ha^?1 ryegrass seed averaged 26% in the first year and 37% in the second. Corresponding calculated mean DM yields of timothy were 2·75 and 3·00 t ha^?1. It is concluded that an early timothy variety is capable of competing with a late-heading perennial ryegrass in frequently cut swards managed to simulate intensive grazing. The strong development of timothy in the dry summer of 1975 suggests that in mixtures of late perennial ryegrass varieties, an early variety of timothy should be beneficial for its spring growth in grazed swards.     

THE EFFECT OF METHOD OF SOWING ON THE ESTABLISHMENT OF GRASS SEED CROPS

Methods used by growers to establish seed crops of perennial ryegrass, cocksfoot, timothy and meadow fescue have been examined. While satisfactory establishments were obtained by sowing perennial ryegrass broadcast, cocksfoot, timothy and meadow fescue should be sown in rows, especially if they are to be sown under a cover crop.     

Effects of continuous sheep stocking and strategic rest periods on the sward characteristics of binary perennial grass/white clover associations

Five binary perennial grass/white clover (Trifolium repens, cv. Menna) mixtures were evaluated over a 3-year period under continuous sheep stocking together with the imposition of a rest period for either an early or a late conservation cut; the experiment with plot sizes of 0·16 ha was replicated three times. The grass species and cultivars used were Merlinda tetraploid and Magella diploid perennial ryegrass (Lolium perenne), Prairial cocksfoot (Dactylis glomerata), Rossa meadow fescue (Festuca pratensis) and Goliath timothy (Phleum pratense). The greatest total lengths of white clover stolon developed in the meadow fescue (171·6 m m^?2) and timothy (151·9 m m^?2) associations compared with those in tetraploid perennial ryegrass (98·6 m m^?2), diploid perennial ryegrass (91·9 m m^?2) and cocksfoot (74·6 m m^?2) (s.e.d. 16·4, P < 0·001). On average, the proportion of white clover stolon that was buried was between 0·86 and 0·89 and this was more abundant in late than early season. Whereas timothy persisted, the persistence of meadow fescue was low under any of the managements tested and this was markedly reduced by the third grazing season. In the diploid perennial ryegrass sward, a late June to early August rest period for conservation enhanced white clover stolon length. An early April to late May rest period greatly reduced total white clover stolon length in both diploid perennial ryegrass and tetraploid perennial ryegrass associations (diploid perennial ryegrass-unrested 89 m m^?2, early rest 56·1 m m^?2, late rest 130·7 m m^?2; tetraploid perennial ryegrass - unrested 125·1 m m^?2, early rest 71 m m^?2, late rest 99·7 m m^?2; s.e.d. 19·19, P < 0·001). The numbers of white clover stolon growing points per unit stolon length were greatest when the sward was rested during late June to early August ?55·9 m^?1 stolon length compared with 45·7 m^?1 for an April to late May rest and 46 m^?1 in the absence of a rest (s.e.d. 2·59, P < 0·001). Likewise, the percentage of stolon above ground was greatest with the late June to early August rest ?15·78% compared with 10·61% for the April to late May rest and 7·69% for no rest (s.e.d. 1·569, P < 0·001). The complementary percentages of buried stolon indicate the important role this fraction has and the need to study stolon behaviour in grazing studies generally. It is concluded that, in relation to perennial ryegrass as a companion grass, meadow fescue and timothy allow better white clover development and cocksfoot less. However, other attributes have to be considered, for example the poor persistence of meadow fescue and the slower regrowth of timothy, both of which allow the invasion of weed grasses, or the lower acceptability of cocksfoot to livestock. The timing of the rest period before the conservation cut can influence white clover development considerably, but the effects differed with different companion grasses.     

Herbage productivity of brome grass (Bromus carinatus)

Herbage production and quality of swards of brome grass (Bromus carinatus. Hook and Am) were compared with other commonly sown grasses at two sites in Scotland. At Ayr, the comparison was with perennial ryegrass ( Lolium perenne L.), cocksfoot ( Dactylis glomerata L.) and timothy ( Phleum pratense L.) under 6-cut (experiment 1) and 4-cut (experiment 2) regimes over 3 years with 360 kg ha⁻¹ fertilizer N applied annually in each experiment. At Edinburgh, brome grass was compared over 3 years with perennial ryegrass and cocksfoot under a 7-cut system given 3(X)-35O kg N ha⁻¹ year⁻¹ (experiment 3) and with perennial ryegrass, cocksfoot and Italian ryegrass ( Lolium multiflorum Lam.) under a 4-cut system given 250–325 kg N ha⁻¹ annually (experiment 4).
Over the 3 years, brome grass gave 1.18, 6.19 and 1.3% less dry matter (DM) production than the other grasses in experiments 1, 2 and 3 respectively; in experiment 4, it was 1 % less productive than Italian ryegrass but 1.2% more productive than the other grasses. The organic matter digestibility (OMD) of brome grass was lower than that of perennial ryegrass but higher than timothy at Ayr, similar to perennial and Italian ryegrasses at Edinburgh but markedly superior to cocksfoot at both sites. N concentrations in brome grass were higher than in the ryegrasses but lower than in cocksfoot. Mineral composition data showed brome grass to be high in P and K, low in Ca and Mg and very low in Na compared with corresponding concentrations in the other grasses.
The variable performance of this brome grass species ( B. carinatus ) against commonly used grasses in the reported experiments, together with similar evidence from the literature, leads to the conclusion that it is unlikely to be suitable for widespread use in the UK; nevertheless, it has shown some promise in drought-prone situations.     

Effect of dairy slurry application rate and forage type on production,soil nutrient status and nitrogen‐use efficiency

Seven forage types (diploid and tetraploid perennial ryegrass, Italian ryegrass and hybrid ryegrass, a low‐input mixture of perennial ryegrass, cocksfoot, timothy and meadow fescue, a mixture of perennial ryegrass and white clover, and monoculture of red clover) were sown in late July 2004. Each received one of four rates of dairy cattle slurry in three annual applications by trailing shoe, which supplied average nitrogen (N) inputs of 0·0, 114·9, 204·8 and 301·2 kg N ha^?1 annum^?1. Treatments were cut either three or four times annually over four years. Average dry‐matter yield (DM) response to slurry N was 15·6 kg DM kg^?1 N. Lowest recovery of slurry N was in the second application each year (after first cut). The data suggest that slurry applied to Italian ryegrass, and also to swards containing legumes on soils with high phosphorus content, will produce a lower DM response to slurry N and result in a lower slurry N recovery than on swards of perennial ryegrass or cocksfoot‐dominant low‐input mixtures. Apparent recovery of slurry N was low at the second cut, especially when first‐cut yields had been high. To maximize slurry N recovery, application to regrowths with potentially slow rates of growth or high legume content should be avoided.     

THE EFFECT OF TIME OF CUTTING ON EAR PRODUCTION AND SEED YIELD IN S.48 TIMOTHY

After uniform defoliation in February, spaced plants of S.48 timothy were cut on one of 12 fortnightly dates, starting at the end of March.
Analysis of the herbage removed at each cut showed an increase in plant and unit tiller weight throughout the season, but a decline in the number of tillers, especially before the beginning of shooting.
The date of ear emergence was not affected by cutting up to 5 May and only slightly delayed in plants cut on 19 May. Plants treated subsequently flowered much later but after a fairly constant interval following defoliation.
Cutting on and after 7 April depressed fertile tiller formation with increasing severity, until only vegetative tillers appeared in the recovery growth following treatment at the beginning of August.
Seed yield per plant declined with the decrease in fertile tiller numbers, but after the early cuts some compensation by production of slightly more seed per ear was recorded.
The results are discussed in terms of the physiological transition of the plant towards flowering.