Vegetation community and factors that affect the woody species composition of riparian forests growing in an urbanizing landscape along the Chao Phraya River,central Thailand

Improved knowledge of the environmental factors that affect woody composition is urgently required for species conservation in riparian zones of urbanizing landscapes. We investigated the environmental factors influencing tree abundance and regeneration in diverse forest types growing in the riparian area of an urbanizing landscape along the Chao Phraya River. We established 252 0.1-ha circular plots in remnant forest patches along 372 km of the river. Cluster analysis was applied to classify the forest types. The relationships between environmental variables and tree abundance were assessed with ordination analysis, and generalized linear models were used to assess seedling/sapling abundance. The cluster analysis revealed five forest types, including floodplain forest with three sub-forest types, swamp forest, and mangrove forest. The ordination indicated that tree abundance in the floodplain forest was positively affected by distance to the ocean and the proportion of forested area. Swamp forest was positively influenced by the proportion of urbanized area and mean rainfall. Mangrove forest was negatively related to distance to the river. Seedling/sapling abundance of the dominant species in the floodplain forests was positively affected by lowland plain topography and negatively affected by the proportion of urbanized area, whereas swamp and mangrove forest species were positively influenced by the proportion of urbanized area and estuarine topography. Mature tree density influenced seedling/sapling abundance of all forest types. Tree abundance and regeneration of the riparian landscape was prevented by the urbanized area, floodplain, estuarine topography, and mature tree densities in remnant forests. These results suggest that remnant forest patches of conserved riparian forests along the urbanized landscape of the Chao Phraya River must be protected and the factors determining their colonization must be considered to enhance restoration practices.     

Control of C,N,P distribution in soils of riparian forests

It is now well accepted that riparian forests have an important role in regulating upstream/downstream flow of matter and energy in river ecosystems. Since geomorphic processes determine the structure of channels and floodplains, we have investigated whether different geomorphic features of riparian forests had any effects on the ability of their soils to retain nutrients and organic carbon. Willow riparian forests were chosen within the annual floodplain of the Garonne River, southwest France, to represent two different geomorphic types. Erosional types of riparian forests (E-type) were characterized by sand deposition on their soils because of high current velocity which hampered fine particle deposition. Depositional types of riparian forests (D-type) were characterized by slower overflow velocity; consequently silt and clay were dominant in their soils. Soil samples were taken at the end of the vegetation growth period, coinciding with low water levels prior to annual floods. Erosion and sedimentation processes affected the distribution of total C,N, and P contents in riparian forest soils, since they were significantly correlated with soil grain size. D-type riparian forest soils act as a sink for upstream/downstream nutrients and carbon flows during floods through accumulation of total C,N and P from year to year. In contrast, E-type riparian forests act as potential nutrient sources during high water periods, since they may release from their soils large amounts of easily available C, N and P into the river. These results demonstrate that nutrients and carbon retention ability of riparian forests soils should be analyzed through their geomorphic features rather than by their vegetation composition. Even if they belong to the same vegetation succession, riparian forests should not be considered as a homogeneous buffering system for upstream/downstream flows of nutrients and organic carbon.     

Observed and modeled directional change in riparian forest composition at a cutbank edge

Lateral migrations of river meanders create transient, spatially transgressive edges where the advancing cutbank edge encroaches upon interior floodplain forest communities. This spatial movement of edge toward the forest interior should initiate directional changes in species composition within a forest plot as it is affected by a changing microclimate and hydrological regime. We found that cutbank edge and forest interior sites in an Iowa floodplain contained markedly different plant assemblages. Species commonly associated with later stages of succession dominated interior sites while cutbank edge sites favored secondary, successional species. Assuming that the cutbank edge sites once contained vegetation similar to that surveyed in the floodplain interior, the observed changes in community structure accompanying channel migration are suggestive of retrograde succession, or retrogression. To link cutbank erosional processes with retrogressional processes, we modified a computer simulation model already in use for floodplain environments. We incorporated the changing edge effects and compared model projections with the data collected from the field sites using detrended correspondence analysis. Without changes, the simulation projected a site compositionally similar to the sampled interior forest. When the changes were initiated, the simulated site progressively took on compositional characteristics similar to the riparian edge sites. Because we included only those forcing functions that would be initiated by cutbank erosion, the model supports the hypothesis that the spatially progressive edge effect results in a directional change in forest community composition analogous to retrogression. Our results demonstrate an interesting linkage between successional and fluvial-geomorphic processes and indicate that site dynamics may be controlled differently in landscapes where sites are progressively created and destroyed than where recurrent disturbances affect the same site.     

Changes in the riparian zone of the lower Eygues River,France, since 1830

The riparian forests along braided rivers are dynamic, frequently rejuvenated by floods and channel changes, and thus dominated by pioneer to middle stages of succession; they are sites of high biodiversity in some regions. The Lower Eygues River (drainage area 1150 km² in southeastern France) is such a braided river system with large alluvial forests dominated by Salix alba, Populus alba, and P. nigra. It was identified as a site of ecological interest by the EU under the Natura 2000 program. Such forests elsewhere in Europe have been identified as reference ecosystems. We documented the historical evolution of this alluvial forest from detailed (1:2500 scale) early 19th C parcel maps, early 20th C topographic maps, aerial photography from 1947 to 1996, and field surveys of topography and riparian vegetation in 1997–1998. Our results show that in 1830, the channel was wide, aggraded, and agricultural pressure extended literally to the channel edge. With decline in the rural population and reduced agricultural and grazing pressure in the catchment, erosion rates declined. Reduced sediment supply led to channel narrowing and incision. This channel narrowing, coupled with reduced agricultural pressure along the banks, has allowed riparian forest to colonize former active channel areas, especially within late 19th-century 20th century flood dykes. In recent decades, aggregate mining, and clearing for recreation and agriculture have fragmented the forest. Thus, the alluvial forest of the Lower Eygues is largely an artifact of changing human land-use over the past century, a context that should frame efforts for preservation and restoration.     

Human landscapes have complex trajectories: reconstructing Peruvian Amazon landscape history from 1948 to 2005

Long-term landscape history studies can probe the complexity of landscape dynamics that appear linear or determined by a single driver on shorter time scales, and may span variations of both human-initiated and naturally occurring drivers. With a variety of historical sources this study traces the history of landscape change in Amazonian communities that have existed since the early 1900’s, in a region comprising both upland and riverine ecosystems. Aerial photography from 1948, 1965 and 1977 and satellite images from 1993 to 2005 are analyzed to reconstruct spatial transformations of the study region. The reconstructed landscape history is analyzed as a result of shifts in economy, policy, local markets and river dynamics. In 1948, the upland region was used for agriculture and farms appeared to be encroaching into primary forest. However by 1965, 49% of the upland farm area had become secondary forest, as farmers left upland farms fallow and moved into the floodplain to farm crops promoted through agricultural credit programs. Between 1965 and 1977 river channel migration affected the riverine landscape, dramatic floods occurred throughout the Amazon River and many farmers migrated to the city. During the 1980’s the credit given to small farmers greatly increased, resulting in the highest density of farms in the landscape by 1993. The disappearance of these credits is reflected in reduced farming activity and increased charcoal production. The results show that agricultural activity and deforestation do not always have a simple trajectory of increment.     

Spatial scaling of core and dominant forest cover in the Upper Mississippi and Illinois River floodplains, USA

Different organisms respond to spatial structure in different terms and across different spatial scales. As a consequence, efforts to reverse habitat loss and fragmentation through strategic habitat restoration ought to account for the different habitat density and scale requirements of various taxonomic groups. Here, we estimated the local density of floodplain forest surrounding each of ~20 million 10-m forested pixels of the Upper Mississippi and Illinois River floodplains by using moving windows of multiple sizes (1?C100 ha). We further identified forest pixels that met two local density thresholds: ??core?? forest pixels were nested in a 100% (unfragmented) forested window and ??dominant?? forest pixels were those nested in a >60% forested window. Finally, we fit two scaling functions to declines in the proportion of forest cover meeting these criteria with increasing window length for 107 management-relevant focal areas: a power function (i.e. self-similar, fractal-like scaling) and an exponential decay function (fractal dimension depends on scale). The exponential decay function consistently explained more variation in changes to the proportion of forest meeting both the ??core?? and ??dominant?? criteria with increasing window length than did the power function, suggesting that elevation, soil type, hydrology, and human land use constrain these forest types to a limited range of scales. To examine these scales, we transformed the decay constants to measures of the distance at which the probability of forest meeting the ??core?? and ??dominant?? criteria was cut in half (S _1/2, m). S _1/2 for core forest was typically between ~55 and ~95 m depending on location along the river, indicating that core forest cover is restricted to extremely fine scales. In contrast, half of all dominant forest cover was lost at scales that were typically between ~525 and 750 m, but S _1/2 was as long as 1,800 m. S _1/2 is a simple measure that (1) condenses information derived from multi-scale analyses, (2) allows for comparisons of the amount of forest habitat available to species with different habitat density and scale requirements, and (3) can be used as an index of the spatial continuity of habitat types that do not scale fractally.     

Scale-specific land cover thresholds for conservation of stream invertebrate communities in agricultural landscapes

Context

In agricultural landscapes, riparian forests are used as a management tool to protect stream ecosystems from agricultural activities. However, the ability of managers to target stream protection actions is limited by incomplete knowledge of scale-specific effects of agriculture in riparian corridor and catchment areas.

Objectives

We evaluated scale-specific effects of agricultural cover in riparian corridor and catchment areas on stream benthic macroinvertebrate (BMI) communities to develop cover targets for agricultural landscapes.

Methods

Sixty-eight streams assigned to three experimental treatments (Forested Riparian, Agricultural Riparian, Agricultural Catchment) were sampled for BMIs. Ordination and segmented regression were used to assess impacts of agriculture on BMI communities and detect thresholds for BMI community metrics.

Results

BMI communities were not associated with catchment agricultural cover where the riparian corridor was forested, but were associated with variation in catchment agriculture where riparian forests had been converted to agriculture. Trait-based metrics showed threshold responses at greater than 70% agricultural cover in the catchment. Increasing agriculture in the riparian corridor was associated with less diverse and more tolerant BMI communities. Eight metrics exhibited threshold responses ranging from 45 to 75% agriculture in the riparian corridor.

Conclusions

Riparian forest effectively buffered streams from agricultural activity even where catchment agriculture exceeds 80%. We recommend managers prioritize protection of forested riparian corridors and that restore riparian corridors where agricultural cover is near identified thresholds be a secondary priority. Adoption of catchment management actions should be effective where the riparian corridor has been converted to agriculture.

     

Riparian influences on stream fish assemblage structure in urbanizing streams

We assessed the influence of land cover at multiple spatial extents on fish assemblage integrity, and the degree to which riparian forests can mitigate the negative effects of catchment urbanization on stream fish assemblages. Riparian cover (urban, forest, and agriculture) was determined within 30 m buffers at longitudinal distances of 200 m, 1 km, and the entire network upstream of 59 non-nested fish sampling locations. Catchment and riparian land cover within the upstream network were highly correlated, so we were unable to distinguish between those variables. Most fish assemblage variables were related to % forest and % urban land cover, with the strongest relations at the largest spatial extent of land cover (catchment), followed by riparian land cover in the 1-km and 200-m reach, respectively. For fish variables related to urban land cover in the catchment, we asked whether the influence of riparian land cover on fish assemblages was dependent on the amount of urban development in the catchment. Several fish assemblage metrics (endemic richness, endemic:cosmopolitan abundance, insectivorous cyprinid richness and abundance, and fluvial specialist richness) were all best predicted by single variable models with % urban land cover. However, endemic:cosmopolitan richness, cosmopolitan abundance, and lentic tolerant abundance were related to % forest cover in the 1-km stream reach, but only in streams that had <15% catchment urban land cover. In these cases, catchment urbanization overwhelmed the potential mitigating effects of riparian forests on stream fishes. Together, these results suggest that catchment land cover is an important driver of fish assemblages in urbanizing catchments, and riparian forests are important but not sufficient for protecting stream ecosystems from the impacts of high levels of urbanization.     

Riparian habitat changes across the continental United States (1972–2003) and potential implications for sustaining ecosystem services

Riparian ecosystems are important elements in landscapes that often provide a disproportionately wide range of ecosystem services and conservation benefits. Their protection and restoration have been one of the top environmental management priorities across the US over the last several years. Despite the level of concern, visibility and management effort, little is known about trends in riparian habitats. Moreover, little is known about whether or not cumulative efforts to restore and protect riparian zones and floodplains are affecting the rates of riparian habitat change nationwide. To address these issues, we analyzed riparian land cover change between the early 1970s and the late 1990s/early 2000s using existing spatial data on hydrography and land cover. This included an analysis of land cover changes within 180 m riparian buffer zones, and at catchment scales, for 42,363 catchments across 63 ecoregions of the continental US. The total amount of forest and natural land cover (forests, shrublands, wetlands) in riparian buffers declined by 0.7 and 0.9%, respectively across the entire study period. Gains in grassland/shrubland accounted for the 0.2% lower percentage of total natural land cover loss relative to forests. Conversely, urban and developed land cover (urban, agriculture, and mechanically disturbed lands) increased by more than 1.3% within riparian buffers across the entire study period. Despite these changes, we documented an opposite trend of increasing proportions of natural and forest land cover in riparian buffers versus the catchment scale. We surmise that this trend might reflect a combination of natural recovery and cumulative efforts to protect riparian ecosystems across the US. However, existing models limit our ability to assess the impacts of these changes on specific ecosystem services. We discuss the implications of changes observed in this study on the sustainability of ecosystem services. We also recommend opportunties for future riparian change assessments.     

Forest fragmentation and regrowth in an institutional mosaic of community, government and private ownership in Nepal

This study analyzes forest change in an area of Nepal that signifies a delicate balance between sustaining the needs and livelihood of a sizable human population dependent on forest products, and an effort to protect important wildlife and other natural resources. The study area, a portion of the Chitwan valley district of Nepal, represents what may be becoming a common institutional mosaic in many countries of the world who have a population reliant on forest products for their livelihood: (1) a national park; (2) a designated park buffer involving participatory forest management programs; (3) scattered patches of designated community forest; and (4) large areas of adjacent landscape made up of mostly private landholdings under agricultural practices. Utilizing Landsat images from 1989 and 2000, we analyze land cover change in each of these management zones using landscape ecology metrics and quantifying proportional distributions of land cover categories. Our results show significant differences in terms of land cover dynamics and landscape spatial pattern between these land ownership classes. These findings indicate that community-based institutions (participatory management programs in the park buffer and the designated community forests) are capable of halting or even reversing trends in deforestation and forest fragmentation.     

Floodplain ecosystem response to climate variability and land-cover and land-use change in Lower Missouri River basin

This contribution aims at characterizing the extreme responses of Lower Missouri River basin ecosystems to land use modification and climate change over a 30-year temporal extent, using long term Landsat data archives spanning from 1975 to 2010. The inter-annual coefficient of variation (CoV) of normalized difference vegetation index was used as a measure of vegetation dynamics to address ecological consequences associated with climate change and the impact of land-cover/land-use change. The slope of a linear regression of inter-annual CoV over the entire time span was used as a sustainability indicator to assess the trend of vegetation dynamics from 1975 to 2010. Deduced vegetation dynamics were then associated with precipitation patterns, land surface temperature, and the impact of levees on alluvial hydrologic partitioning and river channelization reflecting the links between society and natural systems. The results show, a higher inter-annual accumulated vegetation index, and lower inter-annual CoV distributed over the uplands remaining virtually stable over the time frame investigated; relatively low vegetation index with larger CoV was observed over lowlands, indicating that climate change was not the only factor affecting ecosystem alterations in the Missouri River floodplain. We cautiously conclude that river channelization, suburbanization and agricultural activities were the possible potential driving forces behind vegetation cover alteration and habitat fragmentation on the Lower Missouri River floodplain.     

Historical landscape ecology of an urbanized California valley: wetlands and woodlands in the Santa Clara Valley

Historical records provide information to land managers and landscape ecologists attempting to understand current trajectories in altered landscapes. In this study, we synthesized a heterogeneous array of historical sources to reconstruct historical land cover in California’s Santa Clara Valley (a.k.a. “Silicon Valley”). To increase and assess accuracy, we used the triangulation of overlapping, independent data sources and the application of certainty level standards. The region has been subject to extensive urbanization, so we also evaluated the applicability of historical landscape reconstructions to the altered landscape. We found evidence for five major land cover types prior to significant Euro–American modification. Valley freshwater marsh, wet meadow, alkali meadow, willow grove, and valley oak savanna have all experienced extreme decline (85–100%) since Euro–American settlement. However, comparison of historical land cover patterns to contemporary land use suggested several new strategies for environmental recovery, despite the limitations of surrounding urbanization. We also observed a temporal shift in riparian habitat along the mainstem of Coyote Creek, from a relatively open mixture of riparian scrub, sycamore woodland, and unvegetated gravel bars to dense riparian forest, likely resulting from stream flow regulation. By identifying former land cover patterns we provide a basis for evaluating local landscape change and setting restoration targets, including the identification of residual features and under-recognized land cover types. These findings suggest that reliable historical landscape reconstructions can be developed in the absence of standardized historical data sources and can be of value even in highly modified regions.     

Irrigation induced change in vegetation and evapotranspiration in the Central Valley of California

Landscape changes in the Central Valley of California, USA, have been dramatic over the past 100 years. Irrigated agriculture has replaced natural communities of California prairie, riparian forest, tule marsh, valley oak savannah, and San Joaquin saltbrush. This paper addresses the implication of vegetation change on evapotranspiration as a consequence of these changes. It was found that an increase in irrigated agriculture and a 60% reduction in the aerial extent of native vegetation has not produced significant changes in the moisture transfer to the atmosphere. The apparent reason for this result is that irrigated agriculture has substituted one actively transpiring surface for another and, therefore, has not significantly altered the transpiration flux of the landscape.     

Distribution of woodland amphibians along a forest fragmentation gradient

Understanding how changes in land-use affect the distribution and abundance of organisms is an increasingly important question in landscape ecology. Amphibians may be especially prone to local extinction resulting from human-caused transformation and fragmentation of their habitats owing to the spatially and temporally dynamic nature of their populations. In this study, distributions of five species of woodland amphibians with differing life histories were surveyed along a 10 km, spatially continuous gradient of forest fragmentation in southern Connecticut, U.S.A. Redback salamanders (Plethodon cinereus) and northern spring peepers (Pseudacris c. crucifer) occupied available habitat along the gradient's length. Wood frogs (Rana sylvatica) and spotted salamanders (Ambystoma maculatum) were absent from portions of the gradient where forest cover was reduced to below about 30%. Red-spotted newts (Notophthalmus v. viridescens) did not persist below a forest cover threshold of about 50%. Correlations between species' biological traits and their fragmentation tolerance imply that low density, population variability, and high mobility coupled with restricted habitat needs predispose woodland amphibians to local extinction caused by habitat fragmentation. These patterns are in contrast to the widely held notion that populations of the best dispersers are those most tolerant of habitat fragmentation.     

Road Density and Landscape Pattern in Relation to Housing Density,and Ownership,Land Cover,and Soils

Roads are conspicuous components of landscapes and play a substantial role in defining landscape pattern. Previous studies have demonstrated the link between roads and their effects on ecological processes and landscape patterns. Less understood is the placement of roads, and hence the patterns imposed by roads on the landscape in relation to factors describing land use, land cover, and environmental heterogeneity. Our hypothesis was that variation in road density and landscape patterns created by roads can be explained in relation to variables describing land use, land cover, and environmental factors. We examined both road density and landscape patterns created by roads in relation to suitability of soil substrate as road subgrade, land cover, lake area and perimeter, land ownership, and housing density across 19 predominantly forested counties in northern Wisconsin, USA. Generalized least squares regression models showed that housing density and soils with excellent suitability for road subgrade were positively related to road density while wetland area was negatively related. These relationships were consistent across models for different road types. Landscape indices showed greater fragmentation by roads in areas with higher housing density, and agriculture, grassland, and coniferous forest area, but less fragmentation with higher deciduous forest, mixed forest, wetland, and lake area. These relationships provide insight into the complex relationships among social, institutional, and environmental factors that influence where roads occur on the landscape. Our results are important for understanding the impacts of roads on ecosystems and planning for their protection in the face of continued development.     

Grassland ecotopes of the upper Meuse as references for habitats and biodiversity restoration: A synthesis

The river valley of the French upper Meuse and its floodplain, constitutes a relatively natural ecosystem which still contains many endangered species of high conservation value. For example, several birds (Crex crex, Numenius arquata) as well as plant species (Gratiola officinalis, Inula britannica, Teucrium scordium, Ranunculus lingua and Mentha pulegium) which have declined seriously in France in recent times are found in the upper Meuse floodplain. Phytosociological studies and water level measurements have shown that the floristic diversity is mainly influenced by hydrological fluctuations and agricultural practices. The plant communities are structured along a topographical gradient in the high water bed reflecting the duration of floods and the ground water table depth. Agricultural practices have influenced the vegetation changes by selecting species adapted to particular management practices (e.g., fertiliser use, grazing, cutting regime). The data collected in this study from the upper Meuse as enabled 13 grassland and wetland ecotopes to be defined which are correlated with different environmental factors. Fertiliser use, grazing and reduction in the frequency of the cutting lead to a lower species richness because they encourage competitive species. However, it is also demonstrated, that maximum biodiversity is not always synonymous with high conservation value because some impoverished ecosystems, e.g., sedges and tall forb formations, may contain endangered plant and bird species. Knowledge of the boundaries between the different plant communities enables likely changes in floristic composition after modification of one or more site factors to be forecasted. Such factors include, water table depth and flood frequency, cutting regime, fertiliser use and grazing pressure. Thus, the definition of these ecotopes, corresponding to correlations between water regime, agricultural practice and vegetation composition, could lead to the establishment of guidelines for water and agricultural managements that could be involved in restoration projects.     

Effects of changing landscape pattern and U. S. G. S. land cover data variability on ecoregion discrimination across a forest-agriculture gradient

We examined the use of coarse resolution land cover data (USGS LUDA) to accurately discriminate ecoregions and landscape-scale features important to biodiversity monitoring and management. We used land cover composition and landscape indices, correlation and principal components analysis, and comparison with finer-grained Landsat TM data, to assess how well LUDA data discriminate changing patterns across an agriculture-forest gradient in Minnesota, U.S.A. We found LUDA data to be most accurate at general class levels of agriculture and forest dominance (Anderson Level I), but in consistent and limited in ecotonal areas of the gradient and within forested portions of the study region at finer classes (Anderson Level II). We expected LUDA to over-represent major (matrix) cover types and under-represent minor types, but this was not consistent with all classes. 1) Land cover types respond individualistically across the gradient, changing landscape grain as well as their spatial distribution and abundance. 2) Agriculture is not over-represented where it is the dominant land cover type, but forest is over-represented where it is dominant. 3) Individual forest types are under-represented in an open land matrix. 4) Within forested areas, mixed deciduous-coniferous forest is over-represented by several orders of magnitude and the separate conifer and hardwood types under-represented. Across gradual, transitional agriculture-forest areas, LUDA cover class dominance changes abruptly in a stair-step fashion. In general, rare cover types that are discrete, such as forest in agriculture or wetlands or water in forest, are more accurately represented than cover classes having lower contrast with the matrix. Northward across the gradient, important changes in the proportions of conifer and deciduous forest mixtures occur at scales not discriminated by LUDA data. Results suggest that finer-grained data are needed to map within-state ecoregions and discriminate important landscape characteristics. LUDA data, or similar coarse resolution data sources, should be used with caution and the biases fully understood before being applied in regional landscape management. This revised version was published online in July 2006 with corrections to the Cover Date.     

Mapping and analyzing landscape patterns

Landscapes were mapped as clusters of 2 or 3 land cover^** types, based on their pattern within the clusters and tendency for a single type to dominate. These landscapes, called Landscape Pattern Types (LPTs), were combined with other earth surface feature data in a Geographic Information System (GIS) to test their utility as analysis units. Road segment density increased significantly as residential and urbanized land cover components increased from absent, to present as patch, to present as matrix (i.e., the dominant land cover type). Stream segment density was significantly lower in LPTs with an urbanized or residential matrix than in LPTs with either a forest or agriculture matrix, suggesting an inverse relationship between stream network density and the prevalence of human development other than agriculture in the landscape. The ratio of average forest patch size to total forest in the LPT unit decreased as agriculture replaced forest, then increased as residential and urban components dominated. Wetland fractal dimension increased as agriculture and residential land cover components of LPTs increased. Comparison of LPT and LUDA land cover area statistics in ecoregions suggested that land cover data alone does not provide information as to its spatial arrangement.     

Landscape ecology: a framework for integrating pattern and process in river corridors

Investigations of European floodplain rivers demonstrate how landscape ecology can provide an effective framework to integrate pattern and process in river corridors, to examine environmental dynamics and interactive pathways between landscape elements, and to develop viable strategies for river conservation. The highly complex and dynamic nature of intact river corridors is particularly amenable to a landscape ecology perspective. Analysis of spatial patterns has provided considerable insight into environmental heterogeneity across river corridors and is an essential prelude to examining dynamic interactions. For example, data from aerial photographs, digitized maps and year-round field measurements in a glacial flood plain, enabled us to distinguish six channel types, based on the correspondence between connectivity and physicochemical attributes. Spatial data were also used to analyze longitudinal changes in landscape elements along the course of a morphologically-intact riverine corridor, providing insight into the structural complexity that must have characterized many Alpine rivers in the pristine state. Landscape indices were employed to investigate seasonal dynamics in a glacial flood plain of the Swiss Alps which exhibits a predictable expansion/contraction cycle, with corresponding shifts in flow paths (surface and subsurface) and water sources (snowmelt, englacial, subglacial, alluvial aquifer, hillslope aquifer). Surface connectivity exhibited an unexpected biphasic relationship with total channel length, whereas riverscape diversity progressively increased along the entire range of channel length. Reconstituting the functional integrity that characterizes intact river corridors should perhaps be the major goal of river conservation initiatives. Although understanding functional processes at the landscape scale is essential in this regard, few data are available. In the Alluvial Zone National Park on the Austrian Danube, three phases of hydrological connectivity were identified (disconnection, seepage connection and surface connection) that corresponded to the predominance of three functional processes (biotic interactions, primary production and particulate transport) within the river corridor.     

Agriculture intensification and forest fragmentation in the St. Lawrence valley,Québec,Canada

Quantifying remaining forest cover and understanding how thefragmentation process operates with respect to the various land-use practicesare important steps when working to preserve the biodiversity associated withwoodlots in agricultural landscapes. We used LANDSAT satellite imagery, soiltypes, and boundaries of regional county municipalities (RCM) as the samplingunit of a 6 million-ha territory located in southern Québec (Canada), to provide a picture of the forest situation in the St. Lawrence Valley.We assessed the effect of human population densities and various types ofagricultural production on the fragmentation process. On average, 45% of thetotal land area of RCMs is forested. However, in 8 of the 59 RCMs studied 20%orless of the total area is still forest habitat. As agricultural use of landincreased, the density of woodlots also increased but their average sizedecreased. An overall fragmentation effect seems to occur where less than 50%ofthe territory is forested, as it is the case for 31 of the 59 studied RCMs.Fragmentation increased along a gradient from traditional dairyagriculture to more intensive cash crop agriculture. Finally, we foundthat the forest discontinuity index, mean woodlot area, and woodlot densitywerethe best indicators of the ongoing forest fragmentation process, but overallhuman population density is the most useful predictive variable.This revised version was published online in May 2005 with corrections to the Cover Date.