THE EFFECT OF SUPPLEMENTARY BARLEY UPON GROWTH AND EFFICIENCY OF FOOD CONVERSION IN CALVES KEPT AT HIGH GRAZING INTENSITY

Three groups of Hereford × Friesian steer calves, bom in early January 1969, were strip-grazed on eqoal areas of a sward of S321 perennial ryegrass from May to September inclusive, and received kibbled barley at the rate of nil, 3/4% or 1 1/2% of liveweight per day. Adjustments were made to the number of animals per group in order to maintain the same high grazing intensity (defined as 8 cm stubble height in grazed areas) on all treatments. The animals receiving supplementary barley gained weight at a rate 17–19% higher than the controls, but there was no significant difference in growth rate between the high and low levels of supplementation. The effficiency of conversion of barley was only 5–12 kg per animal liveweight gain/100 kg barley OM. The high level of supplementation resulted in an increase in stocking rate of 36 % over that of the control group, and an increase in liveweight gain per unit area of land grazed of 63%. The additional liveweight gain due to supplements, expressed per unit area of land grazed, was relatively constant at 21–24 kg liveweight gain/ 100 kg barley OM consumed.     

THE EFFECT OF NITROGEN AND STOCKING RATE ON THE OUTPUT OF PASTURE GRAZED BY BEEF CATTLE

An experiment was conducted in two successire years to measure the effect of two levels of fertilizer N, 50 and 300 kg/ha (45 and 270 Ib/ac) on the productivity of pastures grazed by young beef cattle. Two stocking rates were imposed at the lower N level and 4 at the higher level. The responses per kg fertilizer N were approximately 1 kg liveweight gain, 20–24 Meal ME and 8–9 kg DM. Maximal yields of about 1000 kg gain/ha (890 Ib/ac) and 19,000 Meal ME/ha (7700 Mcal ME/acre) were recorded. Animal performance was similar on the low and the high N pastures. There was evidence that the chemical quality of pasture was lower on the low N pasture in the first year, but there was no difference in the second year. The numbers of dung pats per ha and the refusal of herbage due to fouling were reduced by Increasing the stocking rate.     

A PORTABLE-CORRAL TECHNIQUE FOR MEASURING THE EFFECT OF GRAZING INTENSITY ON YIELD, QUALITY AND INTAKE OF HERBAGE

A grazing experiment using four stocking rates of sheep, equivalent to 34, 45, 57 and 68 sheep/ha on an assumed 200-day grazing season, was conducted using a portable grazing corral technique. Plots were subjected to fotir grazing periods between mid May and early September. The effect of stocking rate on herbage yield and quality and the influence of these factors and feed intake on liveweights of the sheep were recorded. The mean daily herbage organic matter available over the trial was 9.6, 5.8, 3.3 and 23% of the total liveweight of the sheep at each of the four stocking rates. The highest grazing pressure led to the production of high-quality herbage, but also led to reduced productivity, low feed intake and liveweight losses. The most lenient grazing pressure failed to provide adequate herbage utilization. Even at the most intensive stocking pressure, only 66% of the herbage available to ground level was utilized in grazings after July. Intake results suggested that sheep of 45 kg liveweight required 1000–1200 g digestible organic matter per day to maintain body weight. Despite the higher in vitro digestibility of herbage on offer at the higher stocking rates, intake was limited through lack of herbage; a high degree of correlation existed between herbage availability and herbage intake over the four stocking rates and at all grazings. It is concluded that the portable corral technique is well suited for grazing studies and the assessment of sward response to varying stocking rates and is particularly useful where facilities for more extensive studies are limited.     

A NOTE ON THE ZERO-GRAZING OF BEEF CATTLE

Groups of 8 steers weighing 300–400 kg (660–880 Ib) were rotationally grazed on a ryegrass-dominant pasture for 115 days. The systems compared were zero grazing (Z), field grazing conducted at the same stocking rate (FC), and field grazing conducted at a stocking rate varied with the intention of giving the same liveweight gain per uiimal as zero grazing (FV). Mean daily liveweight gains were: Z, 0.98; FC, 0.78; FV, 0.90 kg/animal (2.2, 1.7 and 20 Ib/animal), and liveweight gains/unit area were in the ratio 100:78:85. Organic-matter intake, measured on four occasions, was, on average, Z, 6.54; FC, 6.18; FV, 687 kg/head daily (14.4, 13.6 and 15.1Ib). From these results it appears that a comparison of zero grazing and field grazing made at the same stocking rate is likely to underestimate the potential of field grazing for beef production from grass.     

Supplementary feeding of forage to grazing dairy cows

For three 8-week periods of the grazing season 48 spring-calving cows were continuously stocked at either a high or a low rate (average 4.9 and 4.3 cows ha⁻¹ respectively) which declined through the season. Within each stocking rate group half the cows were allowed access to hay for 45 min after morning milking; the other half received no hay.
Total dry matter (DM) intakes were increased by offering hay, and intakes of hay were greater at the high stocking rate and during prolonged periods of inclement weather. However, there were times when, because of low herbage height and adverse weather, offering hay once daily could not prevent a decline in total DM intake. Grazing time was reduced and ruminating time increased by offering hay, but the rate of biting at pasture was unaffected. Hay DM was eaten at twice the rate of intake of herbage DM.
Offering hay increased milk yield in early season and liveweight gain in late season. The benefits of offering hay were greatest for the higher yielding cows. There were no significant effects on milk composition.
Stocking rate had only small effects on herbage height, but stocking at the higher rate tended to reduce herbage DM intake and reduced live-weight gain in late season. Levels of utilized metabolizable energy from grazed herbage were high (average 106 GJ ha⁻¹) but were reduced by feeding hay and stocking at the lower rate.     

Supplementary forage for grazing sheep. 1. Effects on lactating ewes and lambs

In two experiments, ewes in weeks 1–20 of lactation received either pasture only or pasture plus ad libitum conserved forage. Herbage height was maintained at 3·3–3·4 cm on both treatments by manipulation of stocking rate in the 24 plots (twelve per treatment) used for the experiment with additional ewes and lambs. In experiment 1, silage, which was of lower metabolizable energy (ME) and crude protein (CP) contents than the grazed herbage, was consumed at an average of 375 g dry matter (DM) ewe⁻¹ d⁻¹. In experiment 2 a strawmix, containing (g kg ⁻¹ freshweight) 480 g barley straw, 300 g concentrate and 220 g molasses, again with a lower ME and CP content than the grazed herbage, was consumed at an average of 165 g DM ewe ⁻¹ d ⁻¹. In both experiments ewes not receiving forage lost weight whereas those with forage gained weight and tended to have increased body condition scores. Offering silage or strawmix did not affect lamb liveweight gain or stocking rate.     

The herbage intake and performance of set-stocked suckler cows and calves

Groups of eight Hereford × Friesian cows and their South Devon cross calves were set stocked over a 24-week grazing season at 3·23 (low), 3·21 (medium) or 4·24 (high) cows ha^-1 together with their calves. For the first 8 weeks only two-thirds of the total area was grazed. Increasing the stocking rate from low to medium reduced daily milk yield and cow and calf liveweight gains by 1·2, 0·24 and 0·29 kg d^-1 respectively, and from medium to high by 1·2, 0·24 and 0·23 kg d^-1. The main sward factor influencing faecal output and herbage intake was the quantity of organic matter on the pastures but the digestibility of the herbage selected also exerted a significant effect on the intake of cows. Major depressions in the herbage intake of cows occurred once the average sward height fell below 7 cm. Output of calf live weight was 628, 658 and 743 kg ha^-1 for the 3 stocking rates from low to high, and for cows 246, 179 and 30 kg ha^-1. It was concluded that decisions on pasture management should be taken in relation to the cow rather than the calf on set-stocked swards.     

THE EFFECT OF NITROGEN, STOCKING RATE AND FREQUENCY OF GRAZING BY BEEF CATTLE ON THE OUTPUT OF PASTURE

Two successive grazing experiments were conducted over 12 weeks on perennial ryegrass pastures with 50 and 44 young cattle to study the effect of N fertilizer when applied at a daily rate of 1 or 3 kg N/ha. At each level of N two stocking rates differing hy 20% were imposed. At the liigher N level and stocking rate, three frequencies of grazing were imposed. At the stocking rates imposed N tended to reduce the daily liveweight gain per head, but increased the total liveweight gain per ha by from 0.79 to 1.58 kg/kg N. A 20% increase in stocking rate depressed gain per head in both years. In the first year it did not improve gain/ha, hut in the second year gain/ha was increased by 12–17%. There was a tendency for performance per animal and per ha to increase as the grazing cycle was lengthened. In 1969 the highest yield in the whole season was 1880 grazing days and 1260 kg gain/ha. The overall response to N fertilizer was similar to that recorded in other reports, but it is possible that a lack of K had limited pasture growth.     

A FURTHER INVESTIGATION OF THE EFFECT OF NITROGEN AND STOCKING RATE ON THE PRODUCTIVITY OF PASTURE FOR BEEF CATTLE

The liveweight gain was measured of beef cattle on pastures stocked at 5.9 to 11.8 cattle/ha and receiving 50, 182 or 329 kg N/ha in a 12-week season. N levels and stocking rates were so adjusted that similar yields of herbage per animal were expected. Results showed that similar liveweight gains per animal, averaging 1.03 kg/day, occurred at all N levels. Liveweight gain/ha ranged from 547 to 1037 kg in 12 weeks and the corresponding quantities of metabolizable energy utilized were 9890 and 17590 Meal. The results confinn earlier work (4, 7).     

THE EFFECT OF STOCKING RATE ON ANIMAL PRODUCTION FROM CONTINUOUS AND ROTATIONAL GRAZING SYSTEMS

Liveweight and wool production data are presented from a grazing management experiment in which continuous stocking was compared with a four-paddock rotational system at three stocking rates on a sown pasture. Significant differences were recorded in liveweight and wool production between stocking rates. The overall effect of management was non-significant but its interaction with stocking rate and year were both significant. At the intermediate stocking rate (8 wethers/ac, 20 wethers/ha) increases in both liveweight and wool production were recorded for the rotational management system; in this treatment there was also less need for supplementary feeding. Greater wool production was recorded from the continuously grazed treatments at both the high and the low stocking rates but there was little effect on liveweight or the need for supplementary feeding.     

THE EFFECT OF NITROGEN, STOCKING RATE AND GRAZING METHOD ON THE OUTPUT OF PASTURE GRAZED BY BEEF CATTLE

An experiment was conducted in 1971 and 1972 to study the effects of two levels of fertilizer N (50 and 504 kg/ha) on the productivity of pastures grazed by young beef cattle. Two stocking rates were imposed at the lower N level and three at the higher. In addition to rotational grazing, set-stocking and an integrated grazing-conservation system were included. The responses/kg N were approximately 1 kg live-weight gain, 19–24 Meal ME and 8–9 kg DM. Maximum yields of 1200–1300 kg live-weight gain/ha and 25,000–27,000 Meal ME/ha were recorded. Productivity of set-stocked pasture was similar to rotational grazing at the high N level, but lower at low N. The integrated system yielded comparable results to high-N rotational grazing. Performances and herbage intakes per animal reflected the stocking rates imposed.     

Herbage intake and liveweight gain of bulls and steers continuously stocked at fixed sward heights during autumn and spring

Two experiments were conducted to examine the relationship between sward surface height, herbage intake and liveweight gain in beef cattle grazed on pasture. In Experiment 1, two 'animal types' (18 Charolais × Angus steers and 18 Friesian bulls) were continuously grazed for 22 days during the late autumn on replicated swards maintained at sward surface heights of 6, 10 and 15 cm. Herbage intakes, assessed from the faecal concentration of chromium delivered from an intraruminal controlled release capsule and the in vitro digestibility of hand-plucked herbage samples, were curvilinearly related to sward height (r = 0·76, p <0·0·01). Average liveweight gains were 0·02, 0·61 and 1·31 kg d^-1 ( P <0·05) and increased linearly ( r = 0·84, P <0·001) with sward height. The maintenance organic matter intakes of the steers and bulls, with initial mean (± s.e.) live weights of 225 ± 15 kg and 172 ± 15 kg respectively, were estimated to be 3·6 and 3·5 kg d^-1 respectively. In Experiment 2 (spring) 36 cattle, including 35 of those used in Experiment 1, were reallocated to sward heights of 5, 10 and 15 cm using the same design as for Experiment 1. Average liveweight gains were 0·94, 1·57 and 1·68kg d^-1 ( p 0·05) and were curvilinearly related to sward height ( r = 0·093, p <0·05). Maintenance intakes could not be reliably extimated for the cattle in Experiment 2 because few animals had liveweight gains close to zero. These trials confirm that liveweight gain in continuously grazed finishing steers and bulls increases with sward surface height to maximum of 8–10cm with spring ryegrass/white clover pastures while, in autumn, swards of 12–15cm height are required to achieve maximum performance.     

The effect of grazed sward height and stocking rate on animal performance and output from beef cow systems

During three consecutive summers, forty spring-calving beef cows and their calves grazed perennial ryegrass-dominant swards receiving 250 kg N ha^?1 at one of two annual stocking rates [2.0 (SR 2.0) or 2.5 (SR 2.5) cows ha^?1] and one of two sward heights [4–5 (LS) or 7–8 (HS) cm] in a 2 × 2-factorial experiment, replicated twice. Sward heights were maintained from turn-out in spring by weekly adjustment of the area grazed and herbage was cut for silage in June and again in August from the areas not grazed. After the second cut of silage there was no control of sward height. Calves were weaned in early October and cows removed from pasture and housed when sward heights fell to 4 cm in autumn. Cows were fed in groups a variable but measured quantity of silage during winter to achieve a body condition score of 2.0–2.25 at turn-out the following spring. During the period of sward height control the cows on the HS treatment gained more live weight than those on the LS treatment (0.841 vs. 0.496 kg day^?1; P<0.01) as did the calves (1.167 vs. 1.105; P<0.05). but the stocking rate treatment had no effect. From the time of second-cut silage to the time of weaning and housing respectively, calves and cows gained more live weight on the SR 2.0 treatment because sward heights were higher. Reproductive performance of cows was not affected by treatment. The quantity of silage produced and consumed per cow was not affected by sward height treatment, but the SR 2.0 treatment produced significantly (P<0.001) more silage (1559 kg dry matter per cow) than the SR 2.5 treatment (833 kg dry matter per cow) and had higher winter silage requirements (1249 vs. 1153 kg dry matter per cow: P<0.05). The overall mean stocking rate at which winter fodder production and requirements would be in balance was calculated as 2.25 cows ha^?1 but values were 1.86, 2.60 and 2.28 in each of the three years of the experiment. The results showed that it was possible to control sward height in temperate beef cow systems by adjusting the area available for grazing. Body condition score can be used as a means of determining the feeding levels required to manipulate body condition of cows over winter to achieve prescribed levels of body condition. The experimental approach allows the identification of the stocking rate at which self-sufficiency in winter fodder can be achieved and the year-to-year variation associated with that stocking rate. This approach could be generalized if information on herbage growth rate were available, either from direct measurement or from predictive models.     

Defoliation and productivity of a phalaris-subterranean clover sward, and the influence of grazing experience on sheep intake

Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha⁻¹) and high (22 sheep ha⁻¹) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.
There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward < lightly grazed sward < heavily grazed sward < repeatedly cut sward.
When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.     

Variations in the surface characteristics of the sward and the short-term rate of herbage intake by calves and lambs

The results are reported of an examination of detailed measurements on the ingestive behaviour of calves and lambs in response to variations in the surface characteristics of swards under strip-grazing and continuous stocking managements. It is shown that intake per bite and the short-term rate of herbage intake were both sensitive to the height of the surface horizon above ground level but, at least under strip-grazing management, not to variations in herbage density in the grazed horizon. The rate of biting was less sensitive to variations in sward conditions, particularly under strip-grazing.
Both intake per bite and rate of intake were more sensitive to variations in grazing height under strip-grazing, where sward changes were rapid, than under continuous stocking, where they were slow. Under continuous stocking, ingestive behaviour was more sensitive to changes in sward conditions in lambs than in calves.     

EFFECT OF HIGH FERTILIZER NITROGEN AND STOCKING RATES ON LIVEWEIGHT GAIN PER ANIMAL AND PER HECTARE

The effect of increasing annual fertilizer N application rate from 400 to 800 kg/ha (357 to 714 lb N/ac) and stocking rate from 5.0 to 7.5 animals per forage ha (2.02 to 3.04 animals per forage ac) on herbage availability, digestible OM intake and live weight gain was investigated in a grazing experiment repeated in 3 consecutive years. A 21×1 day rotational paddock grazing system was used in which the grazing area was increased in the ratio of 1:1.5:3 on two predetermined occasions. Increasing the fertilizer N application rate increased herbage availability by 12% but failed to increase live weight gain. Increasing stocking rate decreased herbage availability per ha and per animal, and also live weight gain per animal, but increased live weight gain per forage ha by 17.6%.     

THE ENERGY RETENTION OF LAMBS GRAZING S37 COCKSFOOT, S23 RYEGRASS AND S24 RYEGRASS AT LOW GRAZING PRESSURE

A grazing experiment to measure the feeding values of 4-week-old regrowths of S37 cocksfoot, S23 ryegrass and S24 ryegrass is described. A low grazing pressure was maintained on the pastures to ensure that animal production was not limited by herbage availability. Each sward was grazed by a group of 12 lambs under "worm-free" conditions. Increases in empty body weight, dry body weight and energy were measured by the comparative slaughter technique.
The liveweight gain of the cocksfoot-fed lambs was the same as the ryegrass-fed groups during the first half of the experiment but was lower in August and September. The final liveweight gains on S23 and S24 ryegrass were 14% higher than on S37 cocksfoot, but in terms of energy retention S23 ryegrass was 18% and S24 7% better than S37 cocksfoot. Digestibility measurements of herbage cut from the plots being grazed showed a positive correlation with energy retention by the lambs. Energy retention was negatively correlated with the proportion of acetic acid in the rumen adds, a fraction which was itself correlated with the soluble carbohydrate content of the herbage, but not with the digestibility of the herbage.
When interpreting animal-production results, intake, digestibility and rumen volatilc-fativ-acid proportions, should be taken into account. Because of the inadequacy of present methods of estimating the herbage intake of grazing animals, measurement of body-energy gains is not justified and empty-liveweight gains are sufficiently precise to measure relative energy retentions at pasture.     

Effect of early season sward management on sward quality and lamb liveweight gain during autumn

In two experiments (1990 and 1991) perennial ryegrass (cv. Talbot) pasture was subjected to six different management treatments by adjusting stock numbers to achieve set sward heights for set periods between turn-out in early April and weaning in early July. The main objective of the experiments was to measure the effect of differences in sward management up to July on subsequent sward type, and the effect of sward type on the performance of weaned lambs between mid-July and early October when all swards were grazed at a sward height of 7 cm. In Experiment 1 swards were grazed at 7 cm initially and then reduced to 3 cm at various dates and maintained at 3 cm until early July. In Experiment 2 swards were grazed initially at 7 cm, and this was reduced to 3 cm and then allowed to return to 7 cm at various dates up to early July. Grazing at a sward height of 7 cm up to weaning, in early July, gave mean twin lamb growth rates of 310 g d^?1, while at a tower sward surface height of 3 cm lamb growth rate was reduced to 206 g d^?1 The density of seed heads produced from July was strongly influenced by earlier management. Grazing at 7 cm up to July gave a high density of seed heads (213m^?2), and this reduced (126 m^?2) by grazing to 3 cm. Seed heads were almost eliminated (17 m^?2) by grazing at 7 cm until late June followed by grazing down to 3 cm in early July. Reducing sward surface height from 7 cm to 3 cm at an earlier date tended to increase the development of seed heads from early July. Where swards were grazed from 7 cm to 3 cm and then allowed to return to 7 cm, the later in the pre-weaning period this took place the greater the reduction in seed head development. Greater seed head density was associated with a significant decrease in the nitrogen content of the herbage on offer in Experiment 2, but an increase in modified acid detergent (MAD) fibre content was not significant. Despite the differences in the sward type on offer, the effect on weaned lamb growth rate was small. Lamb liveweight gain during autumn was significantly reduced in Experiment 2 following earlier grazing at 7 cm compared with 3 cm, but liveweight gain differences in Experiment 1 were not significant.     

THE USE OF LIVEWEIGHT–GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

A grazing experiment is described in which the liveweight gains of small East African Zebu heifers were measured from Hyparrhenia rufa/Stylosanthes gracilis and H. rufa/Centrosema pubescens swards. The design of this trial enabled animal production to be assessed from variations in liveweight gain since stocking rate was maintained at the same level on both pastures, but the overall stocking rate was varied according to seasonal changes in herbage availability.
Production was measured both by using individual animals within a group as independent units, and by true animal replication. Gains from individual animal blocks were shown to be of value for comparing pasture species, provided that the composition of the grazing group was varied. Problems of conducting and interpreting small–scale grazing trials as means for evaluating pastures are discussed.     

Comparison of herbage production under continuous stocking and intermittent grazing

The possibility of increasing the herbage utilized over a grazing season was investigated in a study comparing continuously stocked steady-state swards maintained at optimum height (3.5 cm) with intermittently grazed swards. The intermittent systems were designed (a) to allow periodic increase in leaf area and hence growth rate, (b)to ensure that the accumulated herbage was eaten before it senesced, and (c) lo retain high tiller density by alternating periods of herbage accumulation with periods of continuous stocking. Two treatments (no animals or animal numbers reduced to half those on the 3.5 cm steady-state treatment) were used during the 17-18-d periods of herbage accumulation. Grazing down was completed in 3–4 d, after which two treatments (14 d or 28 d) were used for the intervening periods of continuous stocking when sward height was maintained at 3.5 cm. Herbage production was estimated using the tissue turnover technique, with tiller population densities and rates of growth, senescence and net production per tiller measured at frequent intervals. Intermittent grazing treatments where animals were removed during herbage accumulation resulted in changes in tiller size and number, and in growth rates, but not senescence rates, per tiller such that short-term deviations in the net rate of herbage production occurred compared with the continuously stocked control. The periods of advantage during phases of herbage accumulation were counterbalanced by those of disadvantage during the subsequent steady-state phases. Where animal numbers were reduced during herbage accumulation, sward conditions differed little from those of the continuously stocked control, implying that intake per individual animal was increased. It was concluded that intermittent grazing systems offered no advantage over simpler continuous stocking systems, provided that a flexible approach to conservation was incorporated to allow control of sward conditions on the grazed area.