Spatial patterns of plant invasiveness in a riparian corridor

Analysis of landscape-scale patterns of plant invasiveness can assist in interpreting spatial patterns of plant species richness. We investigated downstream variation in plant invasiveness in the riparian corridor of the free-flowing Vindel River in northern Sweden by introducing seeds of an alien species, Helianthus annuus, in 0.25 m² plots of natural vegetation from mountain headwaters to the coast and found a significant downstream pattern with middle reaches having the highest invasiveness. We related invasiveness to species richness, both on a reach scale (200-m long stretches of riverbank encompassing the experimental plots) and on the scale of experimental plots. We found no significant correlation between plant invasiveness and species richness, neither at the reach nor at the plot scale. The number of available soil substrates shows a significant positive quadratic relationship with location along the river and substrate fineness shows a near significant negative quadratic relationship with location along the river, with middle reaches having coarser substrates. Several studies have shown that plant species richness in riparian corridors often exhibits a quadratic pattern with highest species richness in the middle reaches of a river, similar to the pattern we found for invasiveness. Although species richness per se might not be a primary factor for invasibility, the same habitat conditions as those supporting plant species richness, can help in explaining large-scale patterns of plant invasion in riparian zones.     

Scaling relations between riparian vegetation and stream order in the Whitewater River network,Kansas, USA

Riparian communities have been well-studied along individual streams, but not within the context of networks of which streams are a part. To study networks, hydrologists use Horton–Strahler ordering to assign streams to discrete categories in which increasing numerical value (ω) reflects increasing size of the stream and complexity of the network. A key use of this classification method has been to demonstrate scaling relations between hydrogeomorphic variables and order. These relations now provide a foundation to determine how ecological processes are associated with the geometry and topology of river networks. We used geographic information systems (GIS) to map and measure the stream network and riparian vegetation of the Whitewater River basin of eastern Kansas, USA. With the resulting data, we tested if (1) riparian vegetation scaled with order, and (2) riparian vegetation at confluences of two streams differed from that found along constituent streams. Most characteristics of riparian vegetation scaled with order. In confluence zones, density and diversity of riparian vegetation generally were equivalent to that of the largest constituent stream. Scaling relations between riparian vegetation and order provide a framework to quantify the role of riparian vegetation in the water balance of stream networks and a tool to predict area and distribution of riparian vegetation from network topology.     

Longitudinal- and transverse-scale environmental influences on riparian vegetation across multiple levels of ecological organization

Riparian vegetation is distinct from adjacent upland terrestrial vegetation and its distribution is affected by various environmental controls operating at the longitudinal scale (along the river) or transverse scale (perpendicular to the river). Although several studies have shown how the relative importance of transverse or longitudinal influences varies with the scale of observation, few have examined how the influences of the two scales vary with the level of ecological organization. We modeled vegetation-environment relationships at three hierarchically nested levels of ecological organization: species, plant community, and vegetation type. Our hierarchically structured analyses differentiated the spatial extent of riparian zones from adjacent upland vegetation, the distribution of plant community types within the riparian zone, and the distribution of plant species within community types. Longitudinal gradients associated with climate and elevation exerted stronger effects at the species level than at the community level. Transverse gradients related to lateral surface water flux and groundwater availability distinguished riparian and upland vegetation types, although longitudinal gradients of variation better predicted species composition within either riparian or upland communities. We concur with other studies of riparian landscape ecology that the relative predictive power of environmental controls for modeling patterns of biodiversity is confounded with the spatial extent of the study area and sampling scheme. A hierarchical approach to spatial modeling of vegetation-environment relationships will yield substantial insights on riparian landscape patterns.     

Control of C,N,P distribution in soils of riparian forests

It is now well accepted that riparian forests have an important role in regulating upstream/downstream flow of matter and energy in river ecosystems. Since geomorphic processes determine the structure of channels and floodplains, we have investigated whether different geomorphic features of riparian forests had any effects on the ability of their soils to retain nutrients and organic carbon. Willow riparian forests were chosen within the annual floodplain of the Garonne River, southwest France, to represent two different geomorphic types. Erosional types of riparian forests (E-type) were characterized by sand deposition on their soils because of high current velocity which hampered fine particle deposition. Depositional types of riparian forests (D-type) were characterized by slower overflow velocity; consequently silt and clay were dominant in their soils. Soil samples were taken at the end of the vegetation growth period, coinciding with low water levels prior to annual floods. Erosion and sedimentation processes affected the distribution of total C,N, and P contents in riparian forest soils, since they were significantly correlated with soil grain size. D-type riparian forest soils act as a sink for upstream/downstream nutrients and carbon flows during floods through accumulation of total C,N and P from year to year. In contrast, E-type riparian forests act as potential nutrient sources during high water periods, since they may release from their soils large amounts of easily available C, N and P into the river. These results demonstrate that nutrients and carbon retention ability of riparian forests soils should be analyzed through their geomorphic features rather than by their vegetation composition. Even if they belong to the same vegetation succession, riparian forests should not be considered as a homogeneous buffering system for upstream/downstream flows of nutrients and organic carbon.     

The formation of natural levees as a disturbance process significant to the conservation of riverine pastures

Disturbances and patch dynamics are inherent to many ecosystems of the world, especially in the riparian zone. This paper describes the influence of natural levee overbank deposition on riverine grasslands along the meandering River Dinkel (The Netherlands). Here, the rare vegetation type Diantho-Armerietum, characterised by Dianthus deltoides, Thymus pulegioides, Pimpinella saxifraga and Galium verum, has been identified as important to nature conservation. Diantho-Armerietum shows a strong preference for dry, nutrient poor, sandy and relatively young soils, with an elevation approximately 30–50 cm above bankfull discharge level, corresponding to a flooding frequency of 2–3 times per year. The lower zones are strongly influenced by nutrient-rich water, whereas the higher zones are vulnerable to soil acidification. In the intermediate zone, soil development may be reset due to the supply of calcium, adsorbed to recently deposited levee sands. Since deposition rates will decrease with increasing levee heights, new levees are regularly needed to stop the decline of this floriferous vegetation type. The formation of new natural levees is favoured by the occurrence of meander cutoffs, causing a cyclic succession of landforms along the river. Therefore a conservation strategy for this vegetation type needs to aim at the rehabilitation of the natural levee disturbance process, in conjunction with encouraging the meandering of the river.     

Vegetation community and factors that affect the woody species composition of riparian forests growing in an urbanizing landscape along the Chao Phraya River,central Thailand

Improved knowledge of the environmental factors that affect woody composition is urgently required for species conservation in riparian zones of urbanizing landscapes. We investigated the environmental factors influencing tree abundance and regeneration in diverse forest types growing in the riparian area of an urbanizing landscape along the Chao Phraya River. We established 252 0.1-ha circular plots in remnant forest patches along 372 km of the river. Cluster analysis was applied to classify the forest types. The relationships between environmental variables and tree abundance were assessed with ordination analysis, and generalized linear models were used to assess seedling/sapling abundance. The cluster analysis revealed five forest types, including floodplain forest with three sub-forest types, swamp forest, and mangrove forest. The ordination indicated that tree abundance in the floodplain forest was positively affected by distance to the ocean and the proportion of forested area. Swamp forest was positively influenced by the proportion of urbanized area and mean rainfall. Mangrove forest was negatively related to distance to the river. Seedling/sapling abundance of the dominant species in the floodplain forests was positively affected by lowland plain topography and negatively affected by the proportion of urbanized area, whereas swamp and mangrove forest species were positively influenced by the proportion of urbanized area and estuarine topography. Mature tree density influenced seedling/sapling abundance of all forest types. Tree abundance and regeneration of the riparian landscape was prevented by the urbanized area, floodplain, estuarine topography, and mature tree densities in remnant forests. These results suggest that remnant forest patches of conserved riparian forests along the urbanized landscape of the Chao Phraya River must be protected and the factors determining their colonization must be considered to enhance restoration practices.     

Spatio-temporal variations in sediment nutrient levels: the River Adour

The retention capacity of the flood zone of the River Adour (SWFrance) has been estimated. This zone consists of ca. 1 km²of riparian vegetative strips (rvs) and ca. 16 km² offloodplain (Barthes). A novel method of sediment collection hasbeen used in both sectors to determine the quantities of sedimenttogether with concentrations of organic nitrogen and carbon. Thestudy indicates that the vegetative strips accumulate largequantities of sediment with total nitrogen and total carbonconcentrations of ca. 4 mg/g dry matter and ca. 30 mg/g drymatter respectively. These concentrations were found to vary asa function of topography and vegetation. The floodplain receivesless sediment but the observed concentrations of nitrogen andcarbon are more variable and found in the range of 1–9 mg/gdry matter and 10–82 mg/g dry matter respectively. Thehighest levels of C and N are generally found in wooded areas ofthe floodplain.     

From past patterns to future potential: using historical ecology to inform river restoration on an intermittent California river

Context

Effective river restoration requires understanding a system’s potential to support desired functions. This can be challenging to discern in the modern landscape, where natural complexity and heterogeneity are often heavily suppressed or modified. Historical analysis is therefore a valuable tool to provide the long-term perspective on riverine patterns, processes, and ecosystem change needed to set appropriate environmental management goals and strategies.

Objective

In this study, we reconstructed historical (early 1800s) riparian conditions, river corridor extent, and dry-season flow on the lower Santa Clara River in southern California, with the goal of using this enhanced understanding to inform restoration and management activities.

Method

Hundreds of cartographic, textual, and visual accounts were integrated into a GIS database of historical river characteristics.

Results

We found that the river was characterized by an extremely broad river corridor and a diverse mosaic of riparian communities that varied by reach, from extensive (>100 ha) willow-cottonwood forests to xeric scrublands. Reach-scale ecological heterogeneity was linked to local variations in dry-season water availability, which was in turn underpinned by regional geophysical controls on groundwater and surface flow.

Conclusions

Although human actions have greatly impacted the river’s extent, baseflow hydrology, and riparian habitats, many ecological attributes persist in more limited form, in large part facilitated by these fundamental hydrogeological controls. By drawing on a heretofore untapped dataset of spatially explicit and long-term environmental data, these findings improve our understanding of the river’s historical and current conditions and allow the derivation of reach-differentiated restoration and management opportunities that take advantage of local potential.

     

Floodplain ecosystem response to climate variability and land-cover and land-use change in Lower Missouri River basin

This contribution aims at characterizing the extreme responses of Lower Missouri River basin ecosystems to land use modification and climate change over a 30-year temporal extent, using long term Landsat data archives spanning from 1975 to 2010. The inter-annual coefficient of variation (CoV) of normalized difference vegetation index was used as a measure of vegetation dynamics to address ecological consequences associated with climate change and the impact of land-cover/land-use change. The slope of a linear regression of inter-annual CoV over the entire time span was used as a sustainability indicator to assess the trend of vegetation dynamics from 1975 to 2010. Deduced vegetation dynamics were then associated with precipitation patterns, land surface temperature, and the impact of levees on alluvial hydrologic partitioning and river channelization reflecting the links between society and natural systems. The results show, a higher inter-annual accumulated vegetation index, and lower inter-annual CoV distributed over the uplands remaining virtually stable over the time frame investigated; relatively low vegetation index with larger CoV was observed over lowlands, indicating that climate change was not the only factor affecting ecosystem alterations in the Missouri River floodplain. We cautiously conclude that river channelization, suburbanization and agricultural activities were the possible potential driving forces behind vegetation cover alteration and habitat fragmentation on the Lower Missouri River floodplain.     

Patch-level based vegetation change and environmental drivers in Tarim River drainage area of West China

Information on vegetation-related land cover change and the principle drivers is critical for environmental management and assessment of desertification processes in arid environments. In this study, we investigated patch-level based changes in vegetation and other major land cover types in lower Tarim River drainage area in Xinjiang, West China, and examined the impacts of environmental factors on those changes. Patterns of land cover change were analyzed for the time sequence of 1987–1999–2004 based on satellite-derived land classification maps, and their relationships with environmental factors were determined using Redundancy Analysis (RDA). Environmental variables used in the analysis included altitude, slope, aspect, patch shape index (fractal dimension), patch area, distance to water body, distance to settlements, and distance to main roads. We found that during the study period, 26% of the land experienced cover changes, much of which were the types from the natural riparian and upland vegetation to other land covers. The natural riparian and upland vegetation patches were transformed mostly to desert and some to farmlands, indicating expanding desertification processes of the region. A significant fraction of the natural riparian and upland vegetation experienced a phase of alkalinity before becoming desert, suggesting that drought is not the exclusive environmental driver of desertification in the study area. Overall, only a small proportion of the variance in vegetation-related land cover change is explainable by environmental variables included in this study, especially during 1987–1999, indicating that patch-level based vegetation change in this region is partly attributable to environmental perturbations. The apparent transformation from the natural riparian and upland vegetation to desert indicates an on-going process of desertification in the region.     

Effects of recreational use on restored urban floodplain vegetation in urban areas

Urban river restorations focus on restoring aquatic and riparian habitats, increasing flood protection, and enhancing recreational potential. The increased recreational value such newly created urban green spaces is a key benefit of these measures as urban riparian areas are highly valued for recreation. However, high recreational pressure may contribute to the loss of natural vegetation and of biodiversity in restored riparian sites. This study investigates the impact of different recreational intensities and use types on the vegetation structure and vegetation quality by documenting direct (foot-traffic, breaking of branches, stems and roots) and indirect damages (litter and excrements). The major results are fourfold. First, while the proportion of some vegetation types can be correlated to the recreational intensity, neither recreation intensity nor recreation types enhanced the colonization success of invasive species. However, monitoring data showed that human-induced disturbances such as hydro-morphological changes favor alien plant establishment. Second, the study suggests a tipping point for pioneer vegetation at around a density of one user per 10 m river stretch. Already at lower user densities, pressure from trampling can slow down vegetation development. Third, the results indicate that users prefer urban greening and gravel bar elements rather than natural vegetation. Finally, while intensity of direct damages on the vegetation are weakly correlated with the user density, indirect damages increase with the user density. This study concluded that the identification of user hotspots would be helpful in developing a resilient restoration design, which in addition to information about the sensitive vegetation types in relation to recreational users and nature friendly recreational behavior could decrease vegetation damages. In particular, younger recreational users should be targeted by environmental protection campaigns.     

Effects of forest regeneration on songbird movements in a managed forest landscape of Alberta,Canada

Recent studies have shown that barrier effects exist even in relatively vagile species such as forest songbirds. The objectives of this study were to determine whether a 560 × 100 m riparian buffer strip of mature forest was used as a movement corridor by forest songbirds and, if so, to what extent corridor effects persisted as woody vegetation regenerated in the adjacent clearcut. Over a 4-yr period, juvenile movement rates decreased in the riparian buffer strip and increased in the regenerating clearcut. Adult movement rates increased in the riparian buffer strip in the first year after logging, then gradually decreased, while still increasing in the regenerating clearcut. However, both juvenile and adult movement rates were higher in the buffer strip than in an undisturbed control site. Results suggest that most adults we captured held territories in the vicinity of the net lanes,and that most of the juveniles captured were dispersing away from their natal territory. Four years after harvest, juvenile movement rates were higher in the regenerating clearcut than in the riparian buffer strip, but several species had not yet been captured or detected in the regeneration. Our results suggest that the use of the riparian buffer strip as a movement corridor decreased with forest regeneration for both adults and juveniles. However, the buffer strip still acted as a movement corridor for the following species: Philadelphia and Red-eyed Vireos, Red-breasted Nuthatch, and Ovenbird. This revised version was published online in July 2006 with corrections to the Cover Date.     

Scale-specific land cover thresholds for conservation of stream invertebrate communities in agricultural landscapes

Context

In agricultural landscapes, riparian forests are used as a management tool to protect stream ecosystems from agricultural activities. However, the ability of managers to target stream protection actions is limited by incomplete knowledge of scale-specific effects of agriculture in riparian corridor and catchment areas.

Objectives

We evaluated scale-specific effects of agricultural cover in riparian corridor and catchment areas on stream benthic macroinvertebrate (BMI) communities to develop cover targets for agricultural landscapes.

Methods

Sixty-eight streams assigned to three experimental treatments (Forested Riparian, Agricultural Riparian, Agricultural Catchment) were sampled for BMIs. Ordination and segmented regression were used to assess impacts of agriculture on BMI communities and detect thresholds for BMI community metrics.

Results

BMI communities were not associated with catchment agricultural cover where the riparian corridor was forested, but were associated with variation in catchment agriculture where riparian forests had been converted to agriculture. Trait-based metrics showed threshold responses at greater than 70% agricultural cover in the catchment. Increasing agriculture in the riparian corridor was associated with less diverse and more tolerant BMI communities. Eight metrics exhibited threshold responses ranging from 45 to 75% agriculture in the riparian corridor.

Conclusions

Riparian forest effectively buffered streams from agricultural activity even where catchment agriculture exceeds 80%. We recommend managers prioritize protection of forested riparian corridors and that restore riparian corridors where agricultural cover is near identified thresholds be a secondary priority. Adoption of catchment management actions should be effective where the riparian corridor has been converted to agriculture.

     

Riparian habitat changes across the continental United States (1972–2003) and potential implications for sustaining ecosystem services

Riparian ecosystems are important elements in landscapes that often provide a disproportionately wide range of ecosystem services and conservation benefits. Their protection and restoration have been one of the top environmental management priorities across the US over the last several years. Despite the level of concern, visibility and management effort, little is known about trends in riparian habitats. Moreover, little is known about whether or not cumulative efforts to restore and protect riparian zones and floodplains are affecting the rates of riparian habitat change nationwide. To address these issues, we analyzed riparian land cover change between the early 1970s and the late 1990s/early 2000s using existing spatial data on hydrography and land cover. This included an analysis of land cover changes within 180 m riparian buffer zones, and at catchment scales, for 42,363 catchments across 63 ecoregions of the continental US. The total amount of forest and natural land cover (forests, shrublands, wetlands) in riparian buffers declined by 0.7 and 0.9%, respectively across the entire study period. Gains in grassland/shrubland accounted for the 0.2% lower percentage of total natural land cover loss relative to forests. Conversely, urban and developed land cover (urban, agriculture, and mechanically disturbed lands) increased by more than 1.3% within riparian buffers across the entire study period. Despite these changes, we documented an opposite trend of increasing proportions of natural and forest land cover in riparian buffers versus the catchment scale. We surmise that this trend might reflect a combination of natural recovery and cumulative efforts to protect riparian ecosystems across the US. However, existing models limit our ability to assess the impacts of these changes on specific ecosystem services. We discuss the implications of changes observed in this study on the sustainability of ecosystem services. We also recommend opportunties for future riparian change assessments.     

富阳市洞桥镇生态滨水堤岸景观设计研究

在城镇水系设计中,滨水堤岸植物的合理选择及河岸原生植物的保护,是城镇环境森林化、生态化的重要途径。洞桥镇作为一个自然资源保护良好的山区城镇,如何开展对滨水堤岸植物的选择和规划设计,将对该城镇水系河道产生一定的影响。针对洞桥镇河岸水系现状,提出了几种堤岸植物的种植形式,并对堤岸景观设计提出具体工程方法。     

Regional patterns of riparian characteristics in the interior Columbia River basin, Northwestern USA: applications for restoration planning

Recent declines in anadromous Pacific salmonids (Oncorhynchus spp.) have been attributed, in part, to degradation of freshwater habitat. Because riparian areas directly affect instream habitat, assessing riparian characteristics is essential for predicting salmon habitat quality and for prioritizing restoration projects. We quantified land use modification of anadromous fish-bearing streams in the interior Columbia River basin at multiple resolutions. We identified riparian areas in several land use and land cover classes using remotely sensed data. We then interpreted aerial photographs at random locations within each class to quantify riparian modifications at a local (stream reach) scale. Riparian areas in agricultural and urban areas were significantly narrower (~30 m, median) than those in forested or shrub/grass areas (~70 m). The largest proportion of modified riparian areas occurred in low-gradient streams with floodplains in semi-arid ecoregions. Riparian vegetation in these areas is unlikely to provide adequate in-stream functions, making these areas a natural starting point for restoration prioritization. We investigated how existing riparian restoration projects were spatially related to riparian land use and found that restoration effort varied among subwatersheds. Effective strategies for restoring high quality salmon habitat will be watershed-specific and must restore natural watershed processes. By using a hierarchical analysis to identify regional strategies, restoration or conservation activity can be focused in specific basins and thereby increase the likelihood that efforts will significantly improve habitat conditions for listed salmonids.     

Long-term changes in willow spatial distribution on the elk winter range of Rocky Mountain National Park (USA)

We determined changes in willow (Salixspp.) cover in two valleys of the eastern slope of Rocky Mountain National Park,Colorado, USA, and related these changes to suspected causative factors. Changes in vegetation were inferred from digital maps generated from aerial photo-interpretation and field surveys conducted with a global positioning system. The decrease in riparian shrub cover was approximately 20% in both valleys over the period between 1937/46 and 1996, while the decline in tall willow (> 2 m tall) cover was estimated to be approximately 55%in both valleys. Suppressed willows (< 1.5 m tall) were predominantly located in areas affected by flooding and in areas where major river reductions were observed. Both valleys had sites that were being colonized by willows in wet meadows, and open areas created by flood disturbance. The potential causes of willow decline are many. Willow decline was associated with simplification of river spatial pattern, i.e., less complex branching and channelization, and a large flood disturbance. The causes of the reduction in river meanders were not determined, but are likely related to a decline in beavers, an increase in elk, and, possibly climate change. An increase in elk placed increased browsing pressure on willow during the period of the willow decline. Other factors such as climate changes and human activities could have also contributed to the willow decline. The persistence of these riparian ecosystems depends in large part on biotic interactions, particularly between willow, beaver, and elk. This revised version was published online in July 2006 with corrections to the Cover Date.     

Historical influence of man on the riparian dynamics of a fluvial landscape

Man's influence, over the last three centuries, has gradually influenced the dynamics of forest cover along the valley of the Garonne, a seventh order river in Southern France. The vegetation cover of the floodplain depends on topographical levels which govern the frequency and duration of submergence during flooding. Along the valley, forest patches vary from a continuous ribbon of riparian wood along the river to a mosaic of groves towards the upland terraces. In the floodplain, the forest dynamics are influenced by floods, appear to be reversible, and are subject to dominant allogenic processes. On the contrary, forest dynamics on the terraces, which are not influenced by floods, are irreversible and subjected to dominant autogenic processes. Since the end of the 17th century, the structure of riparian woods has been modified by navigation and agriculture leading to a fragmentation of forest cover in the floodplain. Modern agriculture and urbanization have accentuated these tendencies by modifying the hydrologic regime of the river. These historical changes result in a fragmentation of forest cover and a substitution of species in the riparian zone, the forest dynamics being still reversible in the floodplain.     

Using riparian Zone scaling to optimize buffer placement and effectiveness

Context

Riparian buffers reduce subsurface nutrient losses to streams but there is a clear need to identify and prioritize locations for riparian buffer placement to optimize buffer performance. Scaling relations can be used to extrapolate hydrologic behavior within river networks and across catchments.

Objectives

We combined field and laboratory measurements of soils and groundwater quality collected at five riparian monitoring sites of different stream-order scales with landscape analysis to accomplish the following objectives: (1) evaluate the degree to which riparian zone patterns and processes are scaled in a pre-Wisconsin glacial landscape; and (2) use the scaling information to identify optimal placement of riparian buffers in the landform region for nutrient reduction benefits.

Results

Results indicated that there is proportional scaling of riparian zones within the region in terms of sediment texture, groundwater geochemistry and, to a lesser extent, in groundwater nutrient concentrations.

Conclusions

Placement of riparian buffers should be a priority along low order streams (< 3rd order) to best utilize the scaling characteristics of regional riparian zones, although buffering 2nd and 3rd streams may be the most cost effective locations.

     

The contribution of lateral aquatic habitats to insect diversity along river corridors in the Alps

River floodplains are among the most threatened ecosystems globally. Understanding the mechanisms that create and maintain biodiversity and ecosystem functioning of floodplains, is therefore a prerequisite for developing scientifically-sound management and conservation strategies. We quantified the spatial distribution of lateral aquatic habitats (i.e. tributaries, ponds, backwaters) and the associated insect assemblages (Ephemeroptera, Plecoptera, Trichoptera) along three Alpine river corridors (Tagliamento, Thur, and Rhône). Our objective was to assess the relative contribution of lateral habitats to river corridor diversity, and to identify the scale that contributed the most to regional (Alpine scale) species diversity. The number of lateral habitats decreased by 72 % from the near-natural Tagliamento (101) to the Thur river (42) and the regulated Rhône river (28). More than 50 % of the total species richness along each river was restricted to lateral habitats, which also exhibited higher taxon turnover rates (Whittaker’s beta diversity) than the associated main channel. Hierarchical diversity partitioning revealed that beta diversity among corridors was higher than expected, and accounted for 48 % of regional richness, partly reflecting biogeographical differences among catchments. However, diversity partitioning, excluding catchment-specific effects, showed that beta diversity among habitat types contributed significantly to regional richness (79 %). The present study is among the first to quantify the distribution and biodiversity of floodplain habitats along entire river continua. Our results demonstrated that biodiversity would be best preserved by protecting multiple catchments, and that lateral floodplain habitats contribute disproportionally to species richness at the river corridor and regional scale.