Salinity effect on crop development and yield,analysis of salt tolerance according to several classification methods

The publication is a synthesis of previous publications on the results of a long-term lysimeter experiment. From 1989 to 1998, the experimental variables were soil salinity and soil type, from 1999 onwards, soil salinity and crop variety. The plant was studied during the whole growing period by measuring the saline stress and analyzing its effect on leaf area and dry matter development and on crop yield. Salinity affected the pre-dawn leaf water potential, stomatal conductance, evapotranspiration, leaf area and yield.The following criteria were used for crop salt tolerance classification: soil salinity, evapotranspiration deficit, water stress day index. The classification according to soil salinity distinguished the salt tolerant group of sugar beet and wheat, the moderately salt sensitive group comprising broadbean, maize, potato, soybean, sunflower and tomato, and the salt sensitive group of chickpea and lentil. The results for the salt tolerant and the moderately salt sensitive groups correspond with the classification of Maas and Hoffman, excepted for soybean.The evapotranspiration deficit criterion was used, because for certain crops the relation between yield and evapotranspiration remains the same in case of drought and salinity. This criterion, however, did not appear useful for salt tolerance classification.The water stress day index, based on the pre-dawn leaf water potential, distinguished a tolerant group, comprising sugar beet, wheat, maize, sunflower and potato, and a sensitive group, comprising tomato, soybean, broadbean, chickpea and lentil. The classification corresponds with a difference in water use efficiency. The tolerant crops show a more or less constant water use efficiency. The sensitive crops show a decrease of the water use efficiency with increasing salinity, as their yield decreases stronger than the evapotranspiration. No correlation could be found between osmotic adjustment, leaf area and yield reduction. As the flowering period is a sensitive period for grain and fruit formation and the sensitive crops are all of indeterminate flowering, their longer flowering period could be a cause of their greater sensitivity.The tolerant group according to water stress day index can be divided according to soil salinity in a salt tolerant group of sugar beet and wheat and a moderately sensitive group, comprising maize, sunflower and potato. The difference in classification can be attributed to the difference in evaporative demand during the growing period.The sensitive group according to water stress day index can be divided according to soil salinity in a moderately sensitive group, comprising tomato, soybean and broadbean, and a salt sensitive group of chickpea and lentil. The difference in classification can be attributed to the greater salt sensitivity of the symbiosis between rhizobia and grain legume in the case of chickpea and lentil.     

Chickpea water use efficiency in relation to cropping system, cultivar, soil nitrogen and Rhizobial inoculation in semiarid environments

Crops grown in semiarid rainfed conditions are prone to water stress which could be alleviated by improving cultural practices. This study determined the effect of cropping system, cultivar, soil nitrogen status and Rhizobium inoculation (Rz) on water use and water use efficiency (WUE) of chickpea (Cicer arietinum L.) in semiarid environments. The cultivars Amit, CDC Anna, CDC Frontier, and CDC Xena were grown in no-till barley, no-till wheat, and tilled-fallow systems and under various rates of N fertilizer (0, 28, 56, 84, and 112 kg N ha⁻¹) coupled with or without Rz. The study was conducted at Swift Current and Shaunavon, Saskatchewan, from 2004 to 2006. On average, chickpea used about 10 mm of water from the top 0-15 cm soil depth. In the tilled-fallow system, chickpea extracted 20% more water in the 15-30 cm depth, 70% more in the 30-60 cm depth, and 156% more in the 60-120 cm depth than when it was grown in the no-till systems. CDC Xena had WUE of 5.3 kg ha⁻¹ mm⁻¹ or 20% less than the average WUE (6.6 kg ha⁻¹ mm⁻¹) of the three other cultivars, even though these cultivars used the same amounts of water. Water use efficiency increased from 4.7 to 6.8 kg ha⁻¹ mm⁻¹ as N fertilizer rate was increased from 0 to 112 kg N ha⁻¹ when chickpea was grown in the no-till barley or wheat systems, but chickpea grown in the tilled-fallow system did not respond to changes in the fertilizer N rates averaging WUE of 6.5 kg ha⁻¹ mm⁻¹. In the absence of N fertilizer, the application of Rz increased WUE by 33% for chickpea grown in the no-till barley system, 30% in the no-till wheat system, and 9% in the tilled-fallow system. Chickpea inoculated with Rhizobium achieved a WUE value similar to the crop fertilized at 84 kg N ha⁻¹. Without the use of Rz, chickpea increased WUE in a linear fashion with increasing fertilizer N rates from 0 to 84 kg N ha⁻¹. Cropping system, cultivar, and inoculation all had greater impact on WUE than on the amount of water extracted by the crop from the soil. The improvement of cultural practices to promote general plant health along with the development of cultivars with improved crop yields will be keys for improving water use efficiency of chickpea in semiarid environments.     

Sowing soil water content effects on chickpea (Cicer arietinum L.): Seedling emergence and early growth interaction with genotype and seed size

Soil water content at sowing is an important determinant of chickpea seed emergence and early growth. The seed emergence and seedling growth of 9 kabuli and 11 desi chickpea genotypes sown and grown under four soil moisture contents (25, 50, 75 and 100% of field capacity) was measured in a glasshouse study. Across soil moisture treatments the day of first emergence was negatively correlated to all plant growth parameters, e.g. above-ground biomass and plant height (r = −0.83 to −0.94). At 50 and 25% field capacity the onset of seed emergence was delayed, and both the number of seeds that emerged and early growth were reduced. Within the 25% soil moisture treatment, genotypes with an early emergence had a high rate of emergence (as estimated by regression against time) and produced seedlings with a high specific leaf area. Genotypic differences were observed in the number of seeds that emerged, day of first emergence, and early growth across the soil moisture treatments. Differences in final emergence were not related to chickpea type (i.e. desi or kabuli) or seed size, but the kabuli types emerged later than desi types (P < 0.001). The response of early dry matter production and leaf area to soil moisture was quadratically proportional to seed size, while the plant height response was linearly proportional to seed size. As a consequence kabuli chickpeas generated a higher degree of early vigour than desi types as soil moisture increased until seed sizes become extremely large, whereas under low soil moisture, kabuli chickpeas will have a much smaller or possibly no early vigour advantage over desi types.     

Salinity's influence on boron toxicity in broccoli: II. Impacts on boron uptake, uptake mechanisms and tissue ion relations

Limited research has been conducted on the interactive effects of salinity and boron stresses on plants despite their common occurrence in natural systems. The purpose of this research was to determine and quantify the interactive effects of salinity, salt composition and boron on broccoli (Brassica oleracea L.) performance, particularly, element accumulation, ion interactions and boron uptake processes. A greenhouse experiment was conducted using a sand tank system where salinity-B treatment solutions were supplemented with a complete nutrient solution. Chloride-dominated salinity and salinity characteristic of California's San Joaquin valley (SJV), or sulfate-dominated, were tested at ECw (electrical conductivity of the irrigation water) levels of 2, 12 and 19 dS m⁻¹. Each salinity treatment consisted of boron treatments of 0.5, 12 and 24 mg L⁻¹. Salinity, regardless of salt composition, reduced shoot boron concentration at very high boron concentration (24 mg L⁻¹). However, increased salinity increased shoot boron concentration when external boron concentration was low (0.5 mg L⁻¹). Tissue Ca, Mg, Na, K, S and Cl concentrations were also affected by salinity level, chloride or sulfate salinity composition, and in some cases by substrate boron concentration. Calcium concentrations in shoots were greater for chloride-treated salinity as compared to SJV salinity-treated plants; magnesium concentrations trended opposite and were greater in those treated with SJV salinity. Chloride and sodium shoot concentrations both increased with salinity. Shoot chloride was greater with chloride substrate salinity and shoot sodium was greater with SJV substrate salinity. Using stable isotope analysis of solutions to separate transpiration from evapotranspiration (ET), we found that boron uptake and accumulation in the shoot was not simply the product of mass flow (solution concentration × cumulative transpiration), and the vast majority of the water lost from the tank system was by transpiration (>90%) regardless of treatment. Under low substrate boron, the levels of boron in broccoli shoots could be not accounted for by simple passive uptake and transport in the transpiration stream, which suggests that some energy-dependent process was also occurring. However, under high boron treatments, broccoli plants exhibited a mechanism that restricted boron uptake, transport and accumulation in the shoot.     

Effect of saline water irrigation and manure application on the available water content, soil salinity, and growth of wheat

Good water management combined with appropriate soil management is necessary for sustainable crop production in drylands. A pot culture experiment was conducted using sand dune soil under greenhouse conditions to evaluate the response of wheat (Triticum aestivum L.) to the application of farmyard manure (FYM) or poultry manure (PM), and irrigation with water at two salinity levels (0.11 and 2.0 dS m⁻¹) and two irrigation intervals (daily and every second day). The manure was applied at a rate of 20 Mg ha⁻¹. The soil water content, measured 1 h before every irrigation, showed that soil treated with PM retained more water than that treated with FYM, while the control (no manure) contained the least water. FYM treatment resulted in 78 and 21% higher dry matter yield compared to the control and PM treatments, respectively, under daily irrigation using good-quality water. The increase was 29 and 55%, respectively, when saline water was used for daily irrigation. A similar trend was observed with the alternate day irrigation treatment; FYM gave the highest dry matter yield. The number of tillers and plant height showed that FYM was better than PM, which in turn was better than the control under irrigation with good-quality water regardless of the irrigation interval. When water of the highest salinity was used for irrigation, FYM was still always the best, but the control was now better than the PM treatment. The electrical conductivity of the soil measured at the end of the experiment was slightly higher with PM, as compared to the FYM and control treatments. A significant interaction between irrigation water quality and manure application was observed, affecting plant growth. PM aggravated the adverse affect of saline water on plant growth by increasing soil salinity.     

Water use and distribution profile under pulse and oilseed crops in semiarid northern high latitude areas

Oilseed and pulse crops have been increasingly used to replace conventional summer fallow and diversify cropping systems in northern high latitude areas. The knowledge of water use (WU) and its distribution profile in the soil is essential for optimizing cropping systems aimed at improving water use efficiency (WUE). This study characterized water use and distribution profile for pulse and oilseed crops compared to spring wheat (Triticum aestivum L.) in a semiarid environment. Three oilseeds [canola (Brassica napus L.), mustard (Brassica juncea L.) and flax (Linum usitatissimum L.)], three pulses [chickpea (Cicer arietinum L.), dry pea (Pisum sativum L.) and lentil (Lens culinaris Medik.)], and spring wheat were seeded in removable 100 cm deep × 15 cm diameter lysimeters placed in an Aridic Haploboroll soil, in southwest Saskatchewan in 2006 and 2007. Crops were studied under rainfed and irrigated conditions where lysimeters were removed and sampled for plant biomass and WU at various soil depths. Wheat yields were greater than pulse crop yields which were greater than oilseed yields, and WUE averaged 4.08 kg ha⁻¹ mm⁻¹ for pulse crops, 3.64 kg ha⁻¹ mm⁻¹ for oilseeds, and ranged between 5.5 and 7.0 kg ha⁻¹ mm⁻¹ for wheat. Wheat used water faster than pulse and oilseed crops with crop growth. Pulse crops extracted water mostly from the upper 60 cm soil depths, and left more water unused in the profile at maturity compared to oilseeds or wheat. Among the three pulses, lentil used the least amount of water and appeared to have a shallower rooting depth than chickpea and dry pea. Soil WU and distribution profile under canola and mustard were generally similar; both using more water than flax. Differences in WU and distribution profile were similar for crops grown under rainfall and irrigation conditions. A deep rooting crop grown after pulses may receive more benefits from water conservation in the soil profile than when grown after oilseed or wheat. Alternating pulse crops with oilseeds or wheat in a well-planned crop sequence may improve WUE for the entire cropping systems in semiarid environments.     

Yield, water-use efficiencies and root distribution of soybean, chickpea and pumpkin under different subsurface drip irrigation depths and oxygation treatments in vertisols

Most trickle irrigation in the world is surface drip yet subsurface drip irrigation (SDI) can substantially improve irrigation water use efficiency (IWUE) by minimizing evaporative loss and maximizing capture of in-season rainfall by the soil profile. However, SDI emitters are placed at depths, and in many soil types sustained wetting fronts are created that lead to hypoxia of the rhizosphere, which is detrimental to effective plant functioning. Oxygation (aerated irrigation water) can ameliorate hypoxia of SDI crops and realize the full benefit of SDI systems. Oxygation effects on yield, WUE and rooting patterns of soybean, chickpeas, and pumpkin in glasshouse and field trials with SDI at different emitter depths (5, 15, 25, and 35 cm) were evaluated. The effect of oxygation was prominent with increasing emitter depths due to the alleviation of hypoxia. The effect of oxygation on yield in the shallow-rooted crop vegetable soybean was greatest (+43%), and moderate on medium (chickpea +11%) and deep-rooted crops (pumpkin +15%). Oxygation invariably increased season-long WUE (WUE_sl) for fruit and biomass yield and instantaneous leaf transpiration rate. In general, the beneficial effects of oxygation at greater SDI depth on a heavy clay soil were mediated through greater root activity, as observed by general increase in root weight, root length density, and soil respiration in the trialed species. Our data show increased moisture content at depth with a lower soil oxygen concentration causing hypoxia. Oxygation offsets to a degree the negative effect of deep emitter placement on yield and WUE of SDI crops.     

Effect of salinity on yield and nitrogen uptake of four grain legumes and on biological nitrogen contribution from the soil

Broadbean, chickpea, lentil and soybean were grown in a tank experiment and irrigated with waters of three different levels of salinity. The nitrogen uptake of the crop was determined from the yield of aereal biomass and grain, and the corresponding nitrogen contents. The biological contribution of the soil from nitrogen fixation and transformation of organic nitrogen was calculated as difference between the plant uptake and the amount supplied by fertilizer and irrigation minus the loss by drainage.Soil salinity affected crop yield, crop total nitrogen uptake and the nitrogen contribution of the soil. The latter decreased in % of plant uptake at increasing salinity and also decreased stronger than the plant uptake, pointing to a salinity effect on the mineral nitrogen production by biological activity in the soil through nitrogen fixation and transformation of organic nitrogen. A salinity effect on nitrogen fixation could explain, at least partly, the salt sensitivity of grain legumes.     

Assessing the response of chickpea (Cicer aeritinum L.) yield to irrigation water on two soils in Punjab (India): A simulation analysis using the CROPMAN model

Shrinking water resources in northwest India calls for diversification from a rice–wheat cropping system to low-water-requiring crops and development of water-efficient technologies in Punjab state. Chickpea, because of its lower water demand (evapotranspiration) and irrigation requirement has been identified as a suitable alternate crop to wheat. Simulations, averaged over 18 years, using the CROPMAN model indicated that the yield of chickpea on coarse- to medium-textured soils was higher in a rice–chickpea cropping system compared with maize–chickpea and mung–chickpea systems because of increased availability of water. Yield response of chickpea to irrigation depended upon soil texture, the timings and number of irrigations. The optimum yield (2 t ha⁻¹) on coarse- to medium-textured soils after rice can be obtained with one heavy pre-plant and two post-plant irrigations, i.e., one in mid-February and one in mid-March synchronizing irrigations with flowering and grain development stages. Grain yield with irrigation water followed a quadratic function and linear with evapotranspiration. Water use efficiency and evapotranspiration was curvilinear. Grain yield was significantly sensitive to water stress during the pod setting to grain development period irrespective of soil texture.     

Yield and quality of two tomato (Solanum lycopersicum L.) cultivars as influenced by drip and furrow irrigation using waters having high residual sodium carbonate

Performance of tomato when irrigated with sodic waters particularly under drip irrigation is not well known. A field experiment was conducted for 3 years to study the response of tomato crop to sodic water irrigation on a sandy loam soil. Irrigation waters having 0, 5 and 10 mmol_c L⁻¹ residual sodium carbonate (RSC) were applied through drip and furrow irrigation to two tomato cultivars, Edkawi (a salt tolerant cultivar) and Punjab Chhuhara (PC). High RSC of irrigation water significantly increased soil pH, ECe and exchangeable sodium percentage progressively; the increases were higher in furrow compared to drip irrigation. Effect of high RSC on increasing bulk density and decreasing infiltration rate of soil was also pronounced in furrow-irrigated plots. Higher soil moisture and lower salinity near the plant was maintained under drip irrigation than under furrow irrigation. Performance of the two cultivars was significantly different; pooled over 2002–03 and 2003–04 seasons, PC yielded 38.8 and 30.0 Mg ha⁻¹ and Edkawi yielded 31.8 and 22.9 Mg ha⁻¹ under drip and furrow irrigation, respectively. At RSC10, cultivar PC produced 38 and 46% higher fruit yield than cultivar Edkawi under drip and furrow irrigation, respectively. Reduction in fruit yield at higher RSC was due to lower fruit weight under drip irrigation and due to reduced fruit number as well as fruit weight under furrow irrigation. Decrease in fruit weight was more pronounced in cultivar Edkawi than in cultivar PC. Increase in RSC lowered quality of the fruits except the ascorbic acid content. High RSC under drip irrigation, in general, had lesser deteriorating effect on the fruit quality particularly for cultivar PC than under furrow irrigation. For obtaining high tomato yield and better-quality fruits using high RSC sodic waters, drip irrigation should be preferred over furrow irrigation. Better performance of local cultivar PC compared to Edkawi at medium and high RSC suggests that cultivars categorized as tolerant to salinity should be evaluated in the sodic environment particularly when irrigated with high RSC sodic waters.     

Ameliorative effects of inoculation with the plant growth-promoting rhizobacterium Pseudomonas sp. DW1 on growth of eggplant (Solanum melongena L.) seedlings under salt stress

The objective of this work was to evaluate the effect of inoculation with the plant growth-promoting rhizobacterium Pseudomonas sp. DW1 on eggplant (Solanum melongena L.) growth, mineral uptake and activities of the antioxidant enzymes including superoxide dismutase (SOD), peroxidase (POD) and catalase (CAT) of plant leaves under salinity stress. The study was conducted in pot experiments using eggplant (S. melongena L., cv. Yinjia) and a coastal soil. The NaCl concentration of the coastal soil was 0.57 g (kg soil)⁻¹. Four NaCl levels were tested: 0.57, 1.0, 2.0, and 3.0 g NaCl (kg soil)⁻¹, by adding NaCl to soil, respectively. Pseudomonas-inoculated seeds had an increase in the germination percentage over its non-inoculated seeds under salinity. Salinity negatively affected growth of eggplant; however, plants inoculated with Pseudomonas sp. DW1 grew to a significantly greater extent than plants that were not treated with this bacterium. Salinity significantly decreased K⁺ concentration, increased Na⁺ concentration, and did not significantly decrease Ca²⁺ content in shoots of eggplants. Inoculating with Pseudomonas sp. DW1 increased shoot Ca²⁺ of eggplant compared to the non-inoculating eggplant plants under salinity. Inoculating treatments with Pseudomonas sp. DW1 had no effect on shoot Na⁺ concentration in 0.57 and 1 g (kg soil)⁻¹ NaCl, but there were significant decreases in inoculated treatments than in non-inoculated ones at 2 and 3 g (kg soil)⁻¹ NaCl. Salinity decreased SOD activities and increased POD activities, and inoculated Pseudomonas sp. DW1 had an increase effect on SOD activity in the leaves of eggplants. Alteration of mineral uptake and increase in the antioxidant enzyme activities may be two mechanisms for the alleviation of salt stress. Based on the results of the experiment reported herein, the use of the plant growth-promoting rhizobacterium treatment may provide a means of facilitating plant growth under salt stress.     

The effect of saline irrigation water on “Shamouti” orange trees

Summary An irrigation experiment with water of different salinities (2.8, 7.6 and 12.7 mol Cl m^–3) was carried out from 1982 to 1988 in a mature Shamouti orange grove in the coastal plain of Israel. Seasonal accumulation of salts in the soil solution of the root zone (EC of more than 4.0 dS m^–1 at the end of the irrigation season) was almost totally leached during the winter. The average annual rainfall of 550 mm reduced EC values below 1.0 dS m^–1. Tree growth, as measured by the increase in cross sectional area of main branches, was retarded by saline irrigation water (123, 107 and 99 cm² growth per tree during six years for the 2.8, 7.6 and 12.7 mol Cl m^–3 treatments, respectively). Potassium fertilization (360 kg K₂O ha^–1) increased yield at all salinity levels during the last three years of the experiment, mainly by increasing fruit size. Saline irrigation water slightly increased sucrose and C1 concentrations in the fruit juice. Salinity decreased transpiration, increased soil water potential before irrigation and decreased leaf water potential. However, the changes in leaf water potential were small. Leaf Cl and Na concentrations increased gradually during the experimental period, but did not reach toxic levels up to the end of the experiment (4.4 g Cl kg^–1 dry matter in the high salt treatment vs. 1.7 in the control). Relatively more leaf shedding occurred in the salinized trees as compared to the control. The sour orange root-stock apparently provided an effective barrier to NaCl uptake; therefore, the main effect of salinity was probably osmotic in nature. No interactions were found between N or K fertilization and salinity. Additional N fertilization (160 kg N ha^–1 over and above the 200 kg in the control) did not reduce Cl absorption nor did it affect yield or fruit quality. Additional K had no effect on Na absorption but yield and fruit size were increased at all salinity levels. No significant differences were obtained between partial and complete soil surface wetting (30% and 90% of the total soil area resp.) with the same amounts of irrigation water. The effect of salinity on yield over the six years of the experiment was relatively small and occurred only after some years. But, in the last three years salinity significantly reduced average yields to 74.6, 67.1, and 64.2 Mg ha^–1 for the three levels of salinity, respectively.These results suggest that saline waters of up to 13 mol Cl m^–3 primarily influence the tree water uptake and growth response of Shamouti orange trees, whereas yield was only slightly reduced during six years.     

Effects of saline water irrigation on soil salinity,Pecan tree growth and nut production

Summary Irrigated cultivation of pecans (Carya illinoensis K.) has increased dramatically in the Southwestern USA, yet their tolerance to salinity remains largely unknown. The first part of this study was conducted to assess if stunted tree growth reported in clayey soils is related to salinity, and the second part was to evaluate changes in soil salinity and the performance of 11 year old Western trees irrigated with water of 1.1 dSm^–1 and 4.3 dSm^–1 for 4 years. The first study, conducted at a commercial orchard (49 ha) in the El Paso valley (TX), showed a highly significant correlation between tree trunk size and salinity of the saturation extract (EC_e) with r=–0.89. Soil salinity above which trunk size decreased in excess of the standard error was 2.0 dSm^–1 in EC_e from 0–30 cm depth, and 3.0 dSm^–1 in 0 to 60 cm depth with corresponding Na concentrations of 14 and 21 mmol l^–1. Excessive accumulation of salts and Na was found only in silty clay and silty clay loam soils. The second study, conducted at a small experimental field (1 ha), indicated that irrigation with waters of 1.1 and 4.3 dSm^–1 increased EC_e of the top 60 cm profile from 1.5 to 2.2 and 4.2 dSm^–1 and Na concentration in the saturation extract to 17 and 33 mmol l^–1, respectively. The leaching fractions were estimated at 13 and 37% when irrigated with waters of 1.1 and 4.3 dSm^–1, respectively. Tree growth progressively slowed in the saline plots irrigated with water of 4.3 dSm^–1, and became minimal during the 4th year. The cumulative shoot length over the 4 year period was reduced by 24% and trunk diameter by 18% in the saline plots relative to nonsaline plots. Irrigation with the saline water also reduced nut yields by 32%, nut size by 15% and leaflet area by 26% on the 4 year average, indicating that pecans are only moderately tolerant to salinity. The concentration of Na, Cl and Zn in the middle leaflet pair did not differ significantly between the two treatments. Soil salinity provided a more reliable measure for assessing salinity hazard than leaf analysis. However, soil salinity was found to be highly spatially variable following a normal distribution within a soil type. This high variability needs to be recognized in soil sampling as well as managing irrigation.Contribution from Texas Agricultural Experimental Station, Texas A & M University System. This program was supported in part by a grant from the Binational Agricultural Research and Development (BARD) fund     

Potassium-salinity interactions in irrigated corn

Summary Potassium uptake by plants can be affected by high salinity and the Na concentration in the soil solution. There is abundant evidence that Na and the Na/Ca ratio affects K uptake and accumulation within plant cells and organs and that salt tolerance is correlated with selectivity for K uptake over Na. This provides the basis for hypothesis which exists in the literature and was examined in this study, that K application can reduce salinity damage to plants. The main objectives of this study were to: (i) study the effects of salinity and K fertilization interactions on corn yield and nutrient uptake; (ii) test the possibility that salinity damage can be reduced by elevating K fertilization rate; and (iii) study K dynamics in soil as a function of the salinity of the irrigation water, in soils with high and low indigenous potassium. The response of corn (Zea mays (L.) cv. Jubilee) to K fertilization under saline and non-saline conditions was studied by growing corn in two soil types in a pot experiment. Rates of K application to a 3 kg pot were: 0, 15 and 30 mmol K to the Gilat soil and 7.5, 15 and 30 mmol K to the Nordiya soil as KCl. The desired quantity of K was applied in one dose after seedling emergence. The salinity levels of the irrigation water were 4, 20 and 40 mmol charge 1^–1. The irrigation was applied at least every second day and in excess to avoid water stress and to ensure drainage. Increased salinity in the irrigation water significantly decreased yield in both soils. Potassium significantly increased yield at all salinity levels only in the sandy soil which had a low natural level of K, but there was no difference in the relative yield decrease with salinity increase between the lowest and highest K application rates. Potassium fertilization did not eliminate the deleterious effects of salinity on corn yield despite its beneficial effect of increasing K content and reducing the NaK ratio in plant tissue. Potassium uptake by plants was the major factor in K dynamic processes. Potassium adsorption, release and fixation were secondary factors while leaching was an insignificant factor in overall K balance under cropping conditions.     

Effect of frequency of sodic and saline-sodic irrigations and gypsum on the buildup of sodium in soil and crop yields

The effect of three frequencies of irrigation with sodic (high residual alkalinity) and saline-sodic (high residual alkalinity and high NaCl concentration) waters in presence and absence of gypsum application on soil properties and crop yields were investigated under millet (fodder) — wheat — maize (fodder) rotation in a field experiment carried out for 6 years (1986–1992) on a well drained sandy loam Typic Ustochrept soil. Irrespective of the irrigation intervals, sustained use of sodic and salinesodic waters increased pH, electrical conductivity and ESP of the soil and hence significantly decreased crop yields. Application of gypsum decreased ESP and significantly improved crop yields. The beneficial effect of gypsum was lower under saline-sodic irrigation. There were no significant beneficial effects of increasing the frequency of sodic and saline-sodic irrigation, both in presence and absence of applied gypsum, on the yields of wheat and millet (f) crops grown during winter and monsoon seasons, respectively. But decrease in irrigation interval significantly improved yields of maize (f) grown during the hot dry summer period. Frequency of irrigation did not appreciably alter the effectiveness of applied gypsum in wheat and millet (f) but in maize (f), the gypsum treatment was more effective under more frequent irrigation.     

Estimating plant root water uptake using a neural network approach

Water uptake by plant roots is an important process in the hydrological cycle, not only for plant growth but also for the role it plays in shaping microbial community and bringing in physical and biochemical changes to soils. The ability of roots to extract water is determined by combined soil and plant characteristics, and how to model it has been of interest for many years. Most macroscopic models for water uptake operate at soil profile scale under the assumption that the uptake rate depends on root density and soil moisture. Whilst proved appropriate, these models need spatio-temporal root density distributions, which is tedious to measure in situ and prone to uncertainty because of the complexity of root architecture hidden in the opaque soils. As a result, developing alternative methods that do not explicitly need the root density to estimate the root water uptake is practically useful but has not yet been addressed. This paper presents and tests such an approach. The method is based on a neural network model, estimating the water uptake using different types of data that are easy to measure in the field. Sunflower grown in a sandy loam subjected to water stress and salinity was taken as a demonstrating example. The inputs to the neural network model included soil moisture, electrical conductivity of the soil solution, height and diameter of plant shoot, potential evapotranspiration, atmospheric humidity and air temperature. The outputs were the root water uptake rate at different depths in the soil profile. To train and test the model, the root water uptake rate was directly measured based on mass balance and Darcy's law assessed from the measured soil moisture content and soil water matric potential in profiles from the soil surface to a depth of 100 cm. The ‘measured’ root water uptake agreed well with that predicted by the neural network model. The successful performance of the model provides an alternative and more practical way to estimate the root water uptake at field scale.     

Chloride accumulation and partitioning in Barley as affected by differential root and foliar salt absorption under saline sprinkler irrigation

Barley is a crop that has been classified as tolerant to soil salinity, but under sprinkler irrigation with saline water it can readily absorb salts through its leaves and develop injury. Experiments using a triple-line-source sprinkler system were conducted on barley between 1989 and 1991 to determine: (1) the specific effects of foliage wetting on the mass of different shoot components; (2) the relative contribution of root and foliar absorption processes to foliar Cl accumulation; and (3) the extent by which these processes affect Cl partitioning in the shoot at the end of the season. Some plants were covered with plastic during the irrigation process to prevent foliar wetting while others remained uncovered. Salinity affected the partitioning of dry matter in the shoots regardless of whether plants were covered during the irrigation process. The organs associated with reproduction, e.g., heads and peduncles, comprised a larger fraction of the total shoot biomass under high salinity than under low salinity, indicating that plants under salinity stress were able to redistribute their dry matter to favor reproductive growth. The Cl concentration of the young leaves sampled from uncovered plants was linearly related (i.e., r ²>0.71) to the Cl concentration of the irrigation water. Equivalent leaves from covered plants also contained a substantial amount of Cl but concentrations were weakly correlated (i.e., r ²<-0.41) with the concentration of Cl in the irrigation water. At low salinity, there were no differences in leaf Cl concentrations between covered and uncovered treatments. In young leaves, differences between these treatments progressively increased with increasing salinity, indicating that the relative contribution of Cl in the leaf from foliar absorbed salts increased with increasing Cl in the irrigation water. Only in the youngest leaves sampled at the end of the season from plants grown at high salinity was the Cl concentration in uncovered plants (foliar plus root-absorbed Cl) found to be more than twice that in covered plants (only root-absorbed Cl) indicating that most of the Cl in young leaves originated from foliar absorption. In addition, only in the youngest leaves (e.g., flag leaves) was the slope of the relationship between leaf-Cl concentration and Cl concentration of the sprinkling water of uncovered plants more than twice that of covered plants, also indicating that foliar-Cl absorption was more substantial than root-Cl absorption. At high salinity, the difference in leaf Cl concentration between covered and uncovered plants was maximum in the youngest leaf (flag leaf), but differences became progressively smaller with increasing leaf age until ultimately concentrations of chloride in leaves older than the flag leaf-2 were highest in covered plants. In older tissue, it was difficult to distinguish which process, foliar or root absorption, was most responsible for leaf-Cl accumulation. These processes may not be entirely independent of one another and much of the Cl in the oldest leaves of uncovered plants could have been derived from foliar sources during the first month of sprinkling, reaching maximal levels, and thereby restricting root-absorbed Cl. Furthermore, since these leaves at the end of the season are more injured and drier than those from covered plants, late-season sprinkler irrigations may have been responsible for leaching some of the Cl out of these necrotic leaves.     

Water use and water-use efficiency of wheat and barley in relation to seeding dates,levels of irrigation and nitrogen fertilization

Response of timely and late seeded wheat (Triticum aestivum L.) and barley (Hordeum vulgare L.) to three levels of irrigation and four rates of nitrogen was investigated under pre-seeding irrigation. Water extraction pattern and water use of these crops varied markedly. Barley outyielded wheat by 27 and 9%, but used 9 and 21 mm less water, when one and two irrigations were given at critical stages, respectively. These results indicate the possibilities of considerable saving of water (100 mm) for barley without any decrease in grain yield and increased water-use efficiency (WUE) of wheat and barley by irrigating at critical stages as compared to irrigation at 75% depletion of available soil water (ASW). In general WUE decreased with increase in irrigation frequency and delay in seeding.Nitrogen fertilization with marginal additional water use (4–9%) increased yield linearly (45–98%) and thus increased WUE of wheat and barley. This additional water was extracted from below 60 cm depth at tensions ? 1.5 MPa and particularly in maximum growth and reproductive stages. These results suggest that barley should be preferred to wheat under medium to severe water stress and late seeding conditions.     

Response of alfalfa (<Emphasis Type="Italic">Medicago sativa</Emphasis> L.) to diurnal and nocturnal saline sprinkler irrigations. I: total dry matter and hay quality

Little information is available on the quantitative effects on crops of saline sprinkler irrigations and the presumable beneficial effects of nocturnal versus diurnal irrigations. We measured crude protein content, carbon isotope discrimination and total dry matter (TDM) of alfalfa (Medicago sativa L.) subject to diurnal and nocturnal saline sprinkler irrigations. The work was carried out in Zaragoza (Spain) during the 2004–2006 growing seasons with a triple line source sprinkler system using synthetic saline waters dominated by NaCl with an irrigation water EC ranging from 0.5 to 5.6 dS m⁻¹. The quality of alfalfa hay assessed through its crude protein concentration was not significantly affected by salinity. Carbon isotope discrimination, an indicator of the effect of osmotic stress on plant water status, tended to decrease with increases in salinity. Based on a piecewise linear response model, alfalfa grown under saline sprinkler irrigation was shown to be more tolerant (threshold soil salinity, EC_e = 3.5 dS m⁻¹) than in previous experiments under surface irrigation (threshold EC_e = 2.0 dS m⁻¹) at relatively low salinity values, but became more sensitive at higher salinity values as shown by the higher absolute slope (13.4%) for sprinkler as compared to surface irrigation (7.3%). No significant differences in TDM were found between diurnal and nocturnal saline sprinkler irrigations. The recommended practice of irrigating at night for sprinkler irrigation using saline water is therefore not supported by our results in alfalfa grown under semiarid conditions.     

Yield and fibre quality of cotton cultivars as affected by the build-up of sodium in the soils with sustained sodic irrigations under semi-arid conditions

Cotton is commonly grown in many arid and semi-arid regions of the world having sodic ground waters. A field plot study was, therefore, conducted for 2 years to study the effect of sustained sodic irrigation on yield and fibre quality of two hirustum cotton cultivars (F-505 and F-846) and one arborium cultivar (LD-327). The exchangeable sodium percentage (ESP) of 0–30 cm soil under sustained canal water (CW) irrigation treatment was 3.5 whereas long-term irrigations (for more than 10 years) with sodic waters having residual sodium carbonate (RSC) of 5, 10 and 15 me l⁻¹ resulted in ESP bulid-up of 16.4, 39.6 and 56.2, respectively. These sodic waters were used for irrigation in the respective plots, for both years of the study. High ESP of the soil decreased the growth (in terms of plant height) and yield of all the three cotton cultivars. The rate of decline in plant height at 50, 80 and 140 days of sowing (DAS) was maximum in F-505 and minimum in F-846. Compared with CW treatment, relative seed-cotton yield under 16.4, 39.6 and 56.2 ESP levels obtained with respective sodic water treatments for 2 years were 99, 70 and 69%, respectively in F-846, 101, 46 and 29%, respectively in F-505 and 98, 67 and 49%, respectively in LD-327. Similar trends were observed in case of boll number per square metre and boll weight. The cultivar F-846 produced heavier bolls than the other two cultivars under ESP levels of 56.2 obtained under RSC₁₅ treatment which helped this cultivar to perform relatively better. The harmful effect of sodic waters on fibre quality (2.5% span length, micronaire value and bundle strength) were also not observed in the case of F-846. A slight deterioration in fibre quality was, however, observed in cultivars F-505 and LD-327 at an ESP of 56.2 in the soil.