The effects of allowance of deferred herbage and concentrate supplementation when offered to ewe lambs,and system of allocation when offered to ewes in late gestation,on animal performance

The effects of allowance of extended (deferred) grazed herbage (AEGH) and herbage allocation management (HAM) were studied in ewe lambs (248) and late‐gestation ewes (152), respectively, on commercial farms in south‐east Ireland in 2005–06. In Experiment 1, which consisted of four treatments, the effects of AEGH (0·75, 1·25 and 1·75 kg DM per head daily) and concentrate supplementation (0·5 kg per head daily with the 0·75 kg DM herbage allowance) on lamb performance during the extended grazing (16 December to 3 March) and subsequent grazing (4 March to 11 August) seasons were evaluated. Increasing AEGH increased herbage intake linearly (P < 0·001) and live weight (P < 0·001) at the end of extended grazing and the subsequent grazing season. In Experiment 2, single‐ and twin‐bearing ewes were allocated to either a conventional (single‐ and twin‐bearing ewes grazed separately) or leader–follower system (twin‐ and single‐bearing ewes, as leaders and followers respectively) of HAM from 30 January to 24 March. The same quantities of herbage were offered daily for each system. System of HAM affected ewe condition score at lambing but did not alter (P > 0·05) subsequent lamb birth or weaning weights. It is concluded that increasing AEGH to ewe lambs increased liveweight gain during extended grazing and resulted in heavier animals in mid August of the subsequent grazing season. For ewe lambs each 1 kg concentrate DM had the same feed value as 2·4 kg DM AEGH. Use of a leader–follower system for ewes in late pregnancy did not alter lamb birth weight or subsequent performance.     

Does post‐grazing sward height influence sward characteristics,seasonal herbage dry‐matter production and herbage quality?

Extending the grazing season through the production and utilization of high‐quality forage is a key objective in grassland‐based dairy production systems. Grazing swards to a low post‐grazing sward height (PGSH) is a strategy for improving grass utilization. A grazing experiment conducted in Ireland investigated immediate and subsequent effects of PGSH on sward production, utilization and structural characteristics. Swards were grazed to 2·7 cm (severe; S1) or 3·5 cm (moderate; M1) from 10 February to 18 April 2010 (Period 1; P1). From 19 April, each P1 paddock was halved and grazed to either 3·8 cm (S2) or 4·8 cm (M2), until 30 October (Period 2; P2). The first grazing rotation was +7 d on S1 swards compared with M1 swards (45 d), due to greater herbage utilization (+0·22). Herbage production during P1 was not affected by PGSH but a severe PGSH during this period reduced subsequent herbage production: 13·9 (S1) vs. 15·5 t dry matter (DM) ha^?1 (M1) by the end of the study. Leaf proportion was increased (+0·10) on S2 swards compared with M2 swards, but M2 swards produced 1·2 t DM ha^?1 more herbage during P2. Despite the relatively lower level of sward utilization obtained from moderate grazing in P1 (3·5 cm) and P2 (4·5–5·0 cm), such levels of PGSH increased DM production while maintaining sward quality, compared with severe grazing (2·7 cm in P1 and 3·5–4·0 cm in P2).     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 1. Effects of two annual stocking rates in combination with two sward height profiles

The consequences of controlling sward height at two levels, around 5 cm (HS) and around 3·5 cm (LS), during spring and summer by adjusting weekly the proportion of the areas grazed in response to changes in rate of herbage production and utilizing the surplus pasture areas to conserve winter fodder were studied over three complete years for Greyface ewes with their lambs at 15ha⁻¹ (SR15) and 10ha⁻¹ (SR10). The rules used to control grazed sward height resulted in acceptable sward height control in three of four treatments and, by providing supplementary feed when sward height was below target, gave rise to similar levels of individual animal performance. Total output of lamb weaned was greater for the SR15 than for the SR10 flocks (607 vs 477 kg live weight ha⁻¹; P <0·001). Amounts of winter fodder produced were less for the SR15 than for the SR10 flocks [39 vs 213kg dry matter (DM) per ewe; P < 0·001]. Significantly more supplementary feed (10·0 vs 4·4kg DM per ewe; P < 0·01) and hay (13·7 vs 4·0kg DM per ewe; P7lt;0·05) were offered around mating to SR15 flocks than to SR10 flocks. It is concluded that, provided that sward height can be controlled between 3·5 and 5·0cm during the spring and summer and that supplementation is offered when the grazed sward is below 3·5 cm, flock performance will fall within acceptable and predictable limits.     

The implications of controlling grazed sward height for the operation and productivity of upland sheep systems in the UK. 2. Effects of two annual stocking rates in combination with two levels of fertilizer nitrogen

When grazed sward surface height was controlled within the range 3·75-5·25 cm during spring and summer, the effects of two annual stocking rates of twenty (SR20) and twelve (SR12) Cheviot ewes per hectare with their lambs and two rates of nitrogen fertilizer, 100(N100) and 200(N200)kg Nha⁻¹ per year on animal performance and yield of silage from areas of pasture surplus to grazing requirements were measured. Decision rules for management of sward height resulted in good control of swards and consistent and satisfactory individual animal performance across treatments. Total output of lamb was greater for SR20 than for SR12 (699 vs 424kg live weight ha⁻¹; P < 0·001). Yield of silage was less for SR20 than for SR12 [27 vs 184 kg dry matter (DM) per ewe; P < 0·001] and less for N100 than for N200 (65 VS 146 kg DM per ewe; P < 0·01). Around the mating period, when sward height fell below 3·5cm, supplementary feed was offered. More concentrates were offered to the SR20 than to the SR12 ewes (12·3 vs 1·2kg DM per ewe; P < 0·001) and to the N100 than to the N200 ewes (8·3 vs 5·2kg DM per ewe; P < 0·01); trends in the amounts of hay offered during that period were similar.     

Effect of different grazing management systems on the herbage mass and pasture height of a Nardus stricta grassland in western Scotland, United Kingdom

The aims of this study were to examine the effect of three grazing treatments (year‐round stocking rates of 0·8 ewes ha^?1, 0·5 ewes ha^?1 and 0·5 ewes ha^?1 plus grazing cattle in summer), imposed for 4 years, on the herbage mass and surface height of a Nardus stricta‐dominated grassland in western Scotland and to obtain estimates of annual productivity of this grassland. Nardus stricta‐dominated grassland comprised proportionately 0·20 of the grazing area. Stocking rate of sheep had no significant effect on the herbage mass of the grassland in the first 2 years of the experiment, although mean summer pasture heights were significantly higher under the lower stocking rate of sheep. The pasture on the treatment with cattle grazing in summer had a significantly lower herbage mass and lower surface height than the two sheep‐only grazing treatments. Year‐to‐year variation in the herbage mass and surface height of herbage in summer was greater than the effect of treatments. Despite changes in surface height, the structural diversity of the grasslands was not increased by the treatments. The annual production of vascular plant material ranged from 417 g DM m^?2 in 1994 to 628 g DM m^?2 in 1996.     

Effect of herbage mass and allowance on sward characteristics,milk production,intake and rumen volatile fatty acid concentration

The objective of this study was to examine the effect of herbage mass and daily herbage allowance (DHA) on sward characteristics and animal performance, dry‐matter intake, rumen pH and volatile fatty acid production of unsupplemented spring‐calving dairy cows throughout the main grazing season. Sixty‐eight Holstein‐Friesian dairy cows were randomly assigned across four treatments (n = 17) in a 2 × 2 factorial design. Two swards were created with different levels of pre‐grazing herbage mass [allocated above 4 cm (>4 cm); 1700 kg DM ha^?1 (medium; M) or 2200 kg DM ha^?1 (high; H)] and two levels of DHA (>4 cm; 16 or 20 kg DM per cow d^?1). An additional eight lactating ruminally cannulated Holstein–Friesian dairy cows were randomly assigned to each treatment in a replicated 4 × 4 Latin square design. Sward and animal measurements were collected across four periods each of 1 week duration in April and May (PI) and July and August (PII). Maintaining the medium‐mass sward across the season improved the nutritive value of the sward in the latter part of the grazing season compared with high‐mass swards, thus resulting in increased animal intakes and milk production throughout PII. The higher organic matter digestibility of the medium‐compared with high‐masses during PII indicates that grazing severity and herbage mass in the spring to mid‐summer period will determine sward quality parameters in the late summer period.     

The effect of stocking rate and calving date on grass production,utilization and nutritive value of the sward during the grazing season

A 2‐year whole‐farm system study compared the accumulation, utilization and nutritive value of grass in spring‐calving grass‐based systems differing in stocking rate (SR) and calving date (CD). Six treatments (systems) were compared over two complete grazing seasons. Stocking rates used in the study were low (2·5 cows ha^?1), medium (2·9 cows ha^?1) and high (3·3 cows ha^?1), respectively, and mean CDs were 12 February (early) and 25 February (late). Each system had its own farmlet of eighteen paddocks and one herd that remained on the same farmlet area for the duration of the study. Stocking rate had a small effect on total herbage accumulation (11 860 kg DM ha^?1 year^?1), but had no effect on total herbage utilization (11 700 kg DM ha^?1 year^?1). Milk and milk solids (MS; fat + protein) production per ha increased by 2580 and 196 kg ha^?1 as SR increased from 2·5 to 3·3 cows ha^?1. Milk production per ha and net herbage accumulation and utilization were unaffected by CD. Winter feed production was reduced as SR increased. Increased SR, associated with increased grazing severity, resulted in swards of increased leaf content and nutritive value. The results indicate that, although associated with increased milk production per ha, grazed grass utilization and improved sward nutritive value, the potential benefits of increased SR on Irish dairy farms can only be realized if the average level of herbage production and utilization is increased.     

Spring and autumn animal treading effects on pre‐grazing herbage mass and tiller density on two contrasting pasture types in Ireland

A perennial ryegrass (Lolium perenne L.)‐dominated sward on a well‐drained soil (Experiment 1) and a creeping bent (Agrostis stolonifera L.)‐dominated sward on a poorly drained soil (Experiment 2) were subjected to four treading treatments: control (C, no damage), light damage (L), moderate damage (M) or severe damage (S) to quantify the effects on herbage dry‐matter (DM) production and tiller density. In Experiment 1, treading damage was imposed in spring. In Experiment 2, one‐third of the site was damaged in autumn, one‐third in spring and one‐third in both spring and autumn. Both sites were rotationally grazed after treading treatments. Pre‐grazing herbage mass was measured eight times in Experiment 1 and seven times in Experiment 2 on each plot, and tiller density was assessed four times in each experiment. In Experiment 1, pre‐grazing herbage mass was reduced by 30% in S plots at the first harvest after damage, but cumulative pre‐grazing herbage DM production was not different between treatments (12·7 t DM ha^?1). In Experiment 2, annual cumulative pre‐grazing herbage mass was reduced by between 14 and 49%, depending on intensity of treading damage event and season when damage occurred. Tiller density was not affected by treatment in either experiment. A perennial ryegrass‐dominated sward on a well‐drained soil was resilient to heavy treading damage. A creeping bent‐dominated sward on poorly drained soil requires a more careful grazing management approach to avoid major losses in cumulative pre‐grazing herbage mass production during wet weather grazing events.     

Impact of spatial heterogeneity of plant species on herbage productivity,herbage quality and ewe and lamb performance of continuously stocked,perennial ryegrass–white clover swards

The benefits of white clover (Trifolium repens L.) in pastures are widely recognized. However, white clover is perceived as being unreliable due to its typically low content and spatial and temporal variability in mixed (grass‐legume) pastures. One solution to increase the clover proportion and quality of herbage available to grazing animals may be to spatially separate clover from grass within the same field. In a field experiment, perennial ryegrass (Lolium perenne L.) and white clover were sown as a mixture and compared with alternating strips of ryegrass and clover (at 1·5 and 3 m widths), or in adjacent monocultures (strips of 18 m width within a 36‐m‐wide field). Pastures were stocked by ewes and lambs for three 10‐month grazing periods. Over the 3 years of the experiment, spatial separation of grass and clover, compared with a grass–clover mixture, increased clover herbage production, although its proportion in the sward declined through time (0·49–0·54 vs 0·34 in the mixture in the first year, 0·28–0·33 vs 0·15 in the second year and 0·03–0·18 vs 0·01 in the third year). Total herbage production in the growing season in the spatially separated treatments decreased from 11384 kg DM ha^?1 in the first year to 8150 kg DM ha^?1 in the third year. Crude protein concentration of clover and grass components in the 18‐m adjacent monoculture treatment was greater than the mixture treatment for both clover (310 vs 280 g kg^?1 DM) and grass (200 vs 180 g kg^?1 DM). There was no clear benefit in liveweight gain beyond the first year in response to spatially separating grass and clover into monocultures within the same field.     

Continuous and rotational grazing of dairy cows – the interactions of grazing system with level of milk yield, sward height and concentrate level

An experiment was conducted to test the hypothesis that for cows with high levels of milk yield, rotational grazing produces higher milk yields than continuous grazing. The comparison of grazing systems was made at two levels of milk yield (initially 20·3 and 32·5 kg d^?1), and interactions with sward height and concentrate level were also examined. The study used 48 multiparous Holstein Friesian cows over a period of 62 d. Mean milk yield, its persistency and composition, live weight, body condition score and liveweight gain were not significantly affected by grazing system at either level of milk yield. There were no significant interactions between grazing system and sward height or concentrate level for any milk production measurement. Mean estimated herbage and total dry matter (DM) intake (P < 0·01), grazing time (P < 0·05) and ruminating time (P < 0·01) were significantly greater on the continuous grazing system. The cows in the higher milk yield group and those grazed at the higher sward height had a significantly (P < 0·05) higher estimated daily herbage DM intake and rate of herbage intake on the continuous grazing system than those on the rotational grazing system. There was no evidence to support the hypothesis that rotational grazing systems support higher levels of milk production than continuous grazing for cows of high milk yield. The shorter grazing time on the rotational grazing system indicated that cows may anticipate the timing of the daily movement of the electric fence, and this reduces their time spent grazing residual herbage.     

Grazing cycle length and silage supplementation: effects on the performance of lactating ewes with twin lambs

Masham ewes, stocked at 20 ewes ha⁻¹, reared twins at pasture in 1979 and 1980 and were rotationally grazed around six paddocks. There were three lengths of grazing cycle (18, 30 or 42 d) and the ewes at pasture were either offered silage ad libitum (S) for 8 weeks of lactation or they were not supplemented with silage (N). By feeding silage, S ewes and lambs moved into greater herbage masses (3735 u. 3390 kg organic matter (OM) ha⁻¹), but there was little effect on net herbage accumulation (10.5 v. 10.0 t OM ha⁻¹) and herbage intakes per ewe were similar (2.31 v . 2.27 kg OM d⁻¹) over the 7-month grazing season. During the period of silage supplementation, total intake was 19% higher for S than N ewes, lamb growth rates were slightly higher (280 v . 273 g d⁻¹) and the ewes lost less weight (−78 v −96 g d⁻¹).
For 18-, 30- and 42-d grazing cycles respectively the herbage masses before grazing were 2825, 3620 and 4240 kg OM ha⁻¹; net herbage accumulations were 9.8, 10.4 and 10.6 t OM ha⁻¹; mean daily herbage intakes by each unit (ewe+lambs) were 2.22, 2.35 and 2.30 kg OM and lamb growth rates from birth to weaning at 20 weeks were 205, 204 and 204 g d⁻¹.
Frequent grazing of relatively light herbage masses or infrequent defoliation of heavier herbage masses made little difference to sward or animal performance. It is concluded that, at this stocking rate, different grazing cycles of 18, 30 or 42 d do not affect the performance of lactating ewes.     

Influence of solar radiation on the productivity and nutritive value of herbage of cool-season species of an understorey sward in a mature conifer woodland

Silvopastoral systems in the Appalachian region of the USA could increase the carrying capacity of livestock and contribute to a reliable supply of high‐quality herbage. In 2000, 2001 and 2002, the influence of solar radiation [0·20, 0·50 or 0·80 of maximum solar radiation (MSR); treatments 20‐, 50‐ and 80‐MSR respectively] on the productivity and nutritive value of a mixture of sown grasses and legumes established under a mature stand of conifers was investigated. Yields of dry matter (DM), crude protein (CP), total non‐structural carbohydrates (TNC) and total digestible nutrients (TDN) were greater for the 80‐MSR treatment except in 2000 when DM yield did not differ. As a proportion of the sward, introduced species (Dactylis glomerata L., Trifolium repens L., and Lolium perenne L.) increased over time for the MSR‐80 treatment, corresponding with a decrease in the proportion of bare area and of non‐introduced species. CP concentration of herbage was 207 g kg^?1 DM or greater across treatments and years with higher concentrations on the 20‐ and 50‐MSR treatments. Herbage from the 80‐MSR treatment had a greater concentration of TNC than that of the 20‐ and 50‐MSR treatments. Estimated concentration of TDN was similar for all treatments in 2000 and greater for the 80‐MSR treatment than the other two treatments in 2001 and 2002. High CP concentrations in herbage, as a result of appropriate thinning of trees in an Appalachian silvopastoral systems, could be utilized as a protein supplement to herbage with low CP and higher fibre concentrations.     

Production performance and meat quality of grazing lambs finished on red clover, lucerne or perennial ryegrass swards

The combined benefits of a high crude protein concentration, and possible protein protection and growth‐promoting properties, make forage legumes potentially attractive as a natural means of increasing liveweight gain and time to slaughter of lambs in lamb finishing systems. An experiment was conducted to compare the production performance and meat quality of grazing lambs finished on red clover (Trifolium pratense), lucerne (Medicago sativa) or perennial ryegrass (Lolium perenne) swards. Replicate (n = 2) swards of red clover, lucerne and perennial ryegrass were rotationally grazed by ten ram lambs and ten ewe lambs from weaning until selection for slaughter at UK fat class 3L. Lambs grazing the red clover sward had a significantly higher liveweight gain and required significantly fewer days to slaughter than lambs grazing the lucerne sward (305 g d^?1 vs. 243 g d^?1; 38 d vs. 50 d), which in turn had a higher liveweight gain and required fewer days to slaughter than lambs grazing the perennial ryegrass sward (184 g d^?1; 66 d). Lambs grazing the red clover and lucerne swards had significantly higher herbage intakes than those grazing the perennial ryegrass sward (2·06, 1·72 and 1·16 kg DM d^?1 respectively), but in vivo digestibility of herbage was similar. Lambs grazing the red clover and lucerne swards also had significantly higher serum urea concentrations than those grazing ryegrass (12·5, 11·1 and 6·2 mmol L^?1 respectively). Killing‐out percentage was significantly higher for lambs grazing the red clover sward than for lambs grazing the perennial ryegrass sward (48% vs. 46%). There were no significant effects of finishing system on meat flavour, but meat from lambs finished on the lucerne sward was oxidatively less stable than that from lambs finished on the perennial ryegrass sward. Grazing the forage legume swards significantly increased the proportion of linoleic and linolenic acid in muscle tissue, and therefore the proportion of unsaturated to saturated fatty acids (0·19, 0·16 and 0·12 for the red clover, lucerne and perennial ryegrass swards respectively). However, the n?6/n?3 ratio was significantly lower for the muscle of lambs grazing the perennial ryegrass sward compared with those grazing the forage legume swards (1·13, 1·08 and 0·98 for the red clover, lucerne and perennial ryegrass swards respectively). The results indicate that by grazing lambs on forage legume swards it is possible to increase individual lamb performance without compromising meat quality.     

Stocking rate effects on liveweight gain of ewes and their twin lambs when grazing subterranean clover–perennial grass pastures

Two grazing experiments were conducted on non‐irrigated tall fescue–subterranean clover and cocksfoot–subterranean pastures subject to summer‐dry conditions in Canterbury, New Zealand, to measure the effect of low (8·3–10 ewes and their twin lambs ha^?1) vs. high (13·9–20 ewes and their twin lambs ha^?1) stocking rates (SR) on lamb and ewe liveweight gain in spring. In tall fescue–subterranean clover pasture, lambs grew faster at low (374 g per head d^?1) than high (307 g per head d^?1) SR, but total liveweight gain per ha was greater at high (12·3 kg ha^?1 d^?1) than low (7·5 kg ha^?1 d^?1) SR. In successive years in spring in cocksfoot–subterranean clover pastures, lambs grew faster at low (327, 385 g per head d^?1) than high (253, 285 g per head d^?1) SR but total liveweight gain per ha was greater at high (7·26, 7·91 kg ha^?1) than low (5·43, 6·38 kg ha^?1) SR. These studies indicate that in summer‐dry areas, subterranean clover‐based pastures will support high twin lamb growth rates in spring, with lower SR leading to higher lamb growth rates and more lambs reaching slaughter weights before the onset of dry conditions.     

Defoliation pattern, foraging behaviour and diet selection by lactating dairy cows in response to sward height and herbage allowance of a ryegrass-dominated pasture

The effects of sward surface height (SSH) and daily herbage allowance (HA) on the defoliation pattern and grazing mechanics of early lactation dairy cows grazing on irrigated perennial ryegrass–white clover pasture were studied. The hypothesis tested was that SSH and HA affect intake and diet selection through their effects on the pattern of defoliation which is influenced by the resistance to prehension bites. Factorial combinations of two initial SSH (14 and 28 cm) and two daily HA (35 and 70 kg DM cow^?1 d^?1) were examined in a replicated experiment. The peak longitudinal tensile force required to break the sward portion encompassed in a 100 cm² area [bite fracture force (BFF₁₀₀)] was measured as an index of the resistance to prehension. The volume of herbage defoliated and herbage intake increased with SSH (P < 0·05) and HA (P < 0·01). Corresponding to an increase in HA from 35 to 70 kg DM cow^?1 d^?1, there was a proportional increase in the total defoliation area (TDA) and intake by 0·24 and 0·55 in the short sward compared with 0·16 and 0·32 in the tall sward respectively. The results of this experiment suggest that a consistent spatial pattern of reduction of the canopy exists during defoliation by cows and that the volume of sward canopy defoliated is the major variable affecting herbage intake. The BFF increased down the sward profile at a rate that was higher (P < 0·05) for the taller sward than for the shorter sward. It is proposed that a relatively lower resistance to prehension in the short sward compared with the tall sward explains the greater proportionate increase in TDA and intake corresponding to an increase in HA. The rate at which BFF₁₀₀ increases down the sward profile is suggested as a sward physical variable that can influence the defoliation process. The estimated time and energy costs of prehension bites are discussed in the context that larger bites are handled more efficiently than smaller bites.     

Effect of breed and sward height on sheep performance and production per hectare during the spring and autumn in Northern Spain

Abstract Effects of breed and sward surface height on ewe liveweight and body condition changes and on lamb liveweight gains during grazing in the spring and autumn were studied. The output per hectare in each season was also calculated. A total of 112 Gallega ewes (35·6 kg live weight) and 204 Latxa ewes (47·5 kg live weight) with their lambs were used during the spring (March–June) to study the effect of five target sward heights (3·0, 4·5, 5·5, 6·5 and 8·0 cm) and the interaction with breed. In autumn (October–December) 155 Gallega and 126 Latxa ewes were used to study the effect of three target sward heights (4·0, 5·5 and 7·0 cm) and the interaction with breed. Each study was replicated twice. A quadratic relationship was found between sward height and ewe liveweight and body condition changes and lamb liveweight gain during the spring grazing season, with the maximum individual performance being achieved at around a sward height of 6·0 cm. There were significant breed × sward height treatment interactions for ewe liveweight and body condition changes, and lamb liveweight gains. Higher liveweight gains were achieved by Latxa ewes and lambs in swards taller than 6·5 cm but they also sustained higher liveweight losses in swards shorter than 4·5 cm. Nevertheless, a higher output (liveweight gains of ewes + lambs) per hectare for a given weaning date or age was achieved by the Gallega ewes. In autumn, the relationship between liveweight change and sward height was linear, with a breed–sward height interaction similar to that observed in the spring. Sward surface heights of 4·0–4·5 cm and 4·5–5·0 cm were required to maintain live weight and body condition in spring and autumn respectively. Lamb and ewe liveweight gains per hectare decreased considerably in swards taller than 6·5 cm.     

The effect of herbage allowance on intake and performance of ewes suckling twin lambs

The effect of herbage allowance on the herbage intake and performance of ewes and their twin lambs at pasture was investigated. Daily herbage allowances of 40,80.120 and 160 g organic matter (OM) kg^-1 ewe live weight, based on herbage mass measured to ground level, were offered during the first 12 weeks of lactation.
The sheep were grazed rotationally around four paddocks of a perennial ryegrass pasture for 7-day periods and herbage mass, extended tiller length and digestibility of the herbage consumed by the animals were estimated. Herbage intake by the ewes was estimated during weeks two to twelve and live weights were recorded weekly: during the last two weeks of the experiment grazing behaviour of one ewe on each treatment was recorded continuously.
Herbage intakes by the ewes (164, 1–81, 2–42 and 268 ± 0153 kg d^-1) and live weight gains of the lambs (202, 245, 274 and 300 ± 7–3 g d^-1) increased with increasing herbage allowances. Herbage intakes by the ewes and growth rates of their lambs increased up to a herbage allowance which was over five times the amount of herbage eaten by the ewes.
As the animals reduced herbage mass and sward height, biting rates by the ewes during grazing increased by 4 (± 0·08) bites min^-1 cm^-1 and masticating rates decreased. Mastications reached a maximum of 90 (± 3·5) min^-1 at a sward surface height of 9 cm.     

A comparison of diploid and tetraploid perennial ryegrass and tetraploid ryegrass/white clover swards under continuous sheep stocking at controlled sward heights. 3. Sward characteristics and animal output, years 4–8

Results for years 4–8 of a long-term grazing experiment on swards of a diploid perennial ryegrass (Lolium perenne), var. Contender (D swards), a tetraploid ryegrass, var. Condesa (T swards) and Condesa with S184 white clover (Trifolium repens) (TC swards), direct sown in May 1987, are presented. The swards were continuously stocked with sheep from 1988 to 1990, as previously reported, and for a further 5 years, 1991–95, at a target sward surface height (SSH) of 4–6 cm. Control of sward height was successfully achieved by variable stocking, except in 1993 when paddocks were set stocked and the resulting mean SSH was 9·3 cm. Grass swards received on average 160 kg N ha^?1 year^?1; grass/clover swards were mainly not fertilized with N with the exception that they were given 30 kg N ha^?1 as a remedial mid-summer application during a period of low herbage mass on offer in 1994 and 1995. Mean white clover content of the swards fell from 18·2% of herbage dry-matter (DM) in 1992 to 8·5% in 1993, whereas stolon lengths fell from 120 to 58 m m^?2. A return to lower sward heights in 1994–95 resulted in an increase in white clover content to 12·8% by the final sampling in August 1995. Perennial ryegrass content of the grass swards remained high throughout (mean 96·7% in 1995). Perennial ryegrass tiller densities recorded in August 1991, 1993 and 1994 showed consistently significant (P < 0·001) sward differences (3-year mean 16 600, 13 700 and 10 100 perennial ryegrass tillers m^?2 for the D, T and TC swards). In 1994, the year after lax grazing, a low perennial ryegrass tiller density (9100 m^?2) and low white clover content (mean 4·3%) in the TC swards resulted in a much lower herbage bulk density than in the grass swards (April–July means 72, 94 and 44 kg OM ha^?1 cm^?1 for the D, T and TC swards). There was a consistent 40 g d^?1 increase in lamb liveweight gain on the TC swards over the T swards, except in 1994. In that year there was a reduction in lamb liveweight gain of 33 g d^?1 on the TC swards and a significant increase in ewe liveweight loss (117 g d^?1) associated with low herbage bulk density despite optimal sward height. Lamb output (kg liveweight ha^?1) on TC swards reflected white clover content, falling from a similar output to that produced from grass given 160 kg N ha^?1, at 18% white clover DM content, down to 60% of grass + N swards with around 5% clover. A 6% greater output from the T than the D swards was achieved mainly through higher stocking rate. The experiment demonstrated a rapid, loss in white clover under lax grazing, and showed that the relationship between performance and sward height is also dependent on herbage density. High lamb output from a grass/clover sward was only achieved when the clover content was maintained at 15–20% of the herbage DM.     

Extensification of grassland use in the Welsh uplands: sheep performance in years 1–6

An experiment was established in 1991 on a 25-year-old perennial ryegrass/bent ( Lolium perenne L. /Agrostis capillaris L.)-dominated pasture in Wales to study the effects of reducing nutrient inputs to previously fertilized upland pasture. The effects of the removal of applications of (1) N (denoted by CaPK) (2) N, P and K (Ca) and (3) N, P, K, and Ca (Nil) were compared with a treatment which received applications of all four nutrients (CaPKN) over a 6-year period (1991–96) in a randomized block design replicated three times. The experiment was managed under a continuous variable stocking regime (ewes and lambs until weaning and ewes thereafter) maintaining a sward surface height of 4·0 cm throughout the grazing season. Although individual liveweight gain of the lambs was unaffected by the treatments, there was a significant reduction ( P  < 0·05) in total lamb liveweight gain, ewe stocking rate and length of grazing season as a result of the withdrawal of nutrients. Over the 6 years total lamb liveweight gain was reduced by 17%, 32% and 45% and ewe stocking rate by 21%, 36% and 49% on treatments CaPK, Ca and Nil, respectively, compared with treatment CaPKN. The effect of withdrawing nutrient inputs on ewe stocking rate was progressive and by 1996 the Nil input treatment displayed a 63% reduction compared with the CaPKN treatment and this was also coupled with a 21-day reduction in length of the grazing season. During the post-weaning period, ewes from the Nil input treatment recorded a liveweight loss in 1995 and only a modest liveweight gain in 1996. This coupled with significantly lower body condition scores ( P  < 0·01) of these ewes in the autumn indicated that the Nil input treatment could lead to reductions in reproductive performance.     

Pattern of herbage intake rate and bite dimensions of rotationally grazed dairy cows as sward height declines

To allow improved prediction of daily herbage intake of dairy cows in rotational grazing systems, intake behaviour was assessed throughout the day in 24‐h paddocks. Herbage intake in 16 lactating Holstein–Friesian cows was assessed using the short‐term (1‐h) weight gain method at four predetermined natural meal times throughout the day (early morning, T1; late morning, T2; mid‐afternoon, T3; and early evening, T4). The study comprised two 4‐day experiments, each with a cross‐over design of four blocks. In both experiments, cows grazed a 24‐h paddock daily, and the effect of the immediately previous grazing experience on intake behaviour was investigated throughout the day, taking account of daily fluctuations in the short‐term physiological condition of the cows. Experiment 1 was carried out to investigate overall grazing behaviour during meals as a sward is progressively depleted during the day, with intake being assessed within the paddock and, hence, on a depleted sward. Experiment 2 similarly investigated the effect of sward depletion and physiological condition throughout the day on intake, but cows were removed to fresh, undefoliated swards during intake measurement periods; thus, intake rate was not influenced by differences in sward condition. Intake behaviour from both experiments was compared to establish the effect on herbage intake of changes in sward state and non‐sward factors. In Experiment 1, sward surface height, available herbage mass, proportion of leaf and green leaf mass declined as the day progressed. Bite mass declined with sward depletion, and mean intake rate was 1·64 kg dry matter (DM) h^–1, which was significantly lower at T3 (P < 0·01) than during other meals. In Experiment 2, plot sward conditions did not change throughout the day, and intake behaviour also remained constant, with a mean intake rate of 2·11 kg DM h^–1. Mean bite depth as a proportion of pregrazing extended tiller height was constant throughout the day (mean 0·32). The results show that, although cows grazed throughout the day on progressively depleted swards, indicative of rotationally grazed paddocks (Experiment 1), bite mass declined linearly and intake behaviour was variable. However, where intake was assessed on high‐quality, undefoliated swards (Experiment 2), intake behaviour was similar regardless of the time of day and the immediately previous experience. There was some indication of an interaction between the effects of the sward and the physiological condition of the animal on herbage intake.