杨树无性系生长和材性的遗传变异及多性状选择

以７年生２５个杨树无性系为材料研究了生长和材质性状的遗传变异及性状之间的相关．６个性状纤维长、纤维宽、长宽比、木材密度、树高、胸径在无性系间均存在极显著的差异。生长和材质性状受中等更强度的遗传控制．６个性状的重复力分别为８１．９％，７３．３％，７０．５２％。６４．３％．６８．８％，７０．５４％．胸径和树高之间、纤维长度与纤维长宽比、胸径之间、木材密度与胸径之间都呈现出较高的遗传正相关．宜接选择效果优于间接选择，指数选择优于直接选择。由４性状树高、胸径、纤维长度与基本密度配合的选择指数对本实验群体比较合适。利用选择指数评选出４个生长和材质兼优的优良无性系。     

Study on Inheritance and Variation of Wood Fiber Length of Populus tomentosa Clones

本文对４个地点的毛白杨无性系进行了木材纤维长度遗传变异的研究．结果表明木材纤维长度在地点和无性系水平上均有显著的差异．木材纤维长度的无性系重复力估计为０．７９，表明这一材性性状受到较强的遗传控制，可以通过遗传手段得到改良．本文还研究了木材纤维长度在株内的遗传变异．木材纤维长度以及其它性状间的相关分析发现木材纤维长度和树木生长性状（包括材积、树高、胸径和树干通直度）之间均呈显著的正相关关系，但在木材纤维长度和木材基本密度之间存在显著的负相关关系．研究结论认为根据不同地点生长速率与材性的关系，可以进行以生长性状或木材材性性状为主的选择，也可以进行两方面的联合选择．     

杉木无性系生长和木材密度的遗传变异及选择

研究了８年生杉木无性系试验林的树高、胸径、材积和木材密度的遗传变异规律。结果表明：试验中各性状在无性系间存在显著差异,遗传变异系数树高为９．０％,胸径为１３．７％,材积为３０．０％,而木材密度只有６．７％,各性状均有较高的重复力,木材密度和胸径的重复力在９０．０％以上。树高、胸径、材积３个生长性状间有较高的遗传正相关,而胸径、材 与木材密度间表现出中度的遗传负相关,树高与木材密度间则表现出弱度的遗     

毛白杨无性系木材基本密度遗传变异研究(英文)

对河北、河南、山东、甘肃的４块毛白杨无性系测定林中约１００个无性系进行了木材基本密度的测定，对木材基本密度在地点和无性系间的遗传变异进行了研究．结果表明：毛白杨无性系的木材基本密度平均值为０．４３６３ｇ／ｃｍ３，木材基本密度在地点和无性系间均存在极显著差异．木材基本密度的无性系重复力为０．８２～０．９１，属于受遗传控制较强的材性性状．对生长和材性的相关研究表明，木材基本密度与生长性状（包括胸径、树高和材积）一般存在显著的负相关，但不同地点上表现为不同的正或负的相关关系     

杉木无性系生长和木材密度的遗传变异及选择

研究了８年生杉木无性系试验林的树高、胸径、材积和木材密度的遗传变异规律。结果表明：试验中各性状在无性系间存在显著差异，遗传变异系数树高为９０％，胸径为１３７％，材积为３００％，而木材密度只有６７％，各性状均有较高的重复力，木材密度和胸径的重复力在９００％以上。树高、胸径、材积３个生长性状间有较高的遗传正相关，而胸径、材积与木材密度间表现出中度的遗传负相关，树高与木材密度间则表现出弱度的遗传负相关。采用简化的指数选择法评选出４个优良的无性系，材积现实增益２５５２％，遗传增益２１０５％，用木材密度与ＣＫ相近。     

毛白杨无性系木材性状与生长性状的相关分析

对24个20年生毛白杨无性系（含对照）的生长性状、木材基本密度和纤维形态进行了测定分析,并分析了木材性状与生长性状的相关性。结果表明：毛白杨无性系各生长性状差异均达显著和极显著水平,为选择优良无性系提供了丰富的变异来源。毛白杨无性系木材材质指标满足造纸要求,适合作纤维原料,通过对毛白杨无性系生长性状与木材性状之间的相关分析,发现毛白杨木材基本密度与胸径、材积存在一定的相关,与树高存在微弱的相关,总的来看木材基本密度与生长性状相关均不显著;而纤维长度与胸径、材积存在弱负相关,与树高存在微弱的相关。研究中也发现生长快的无性系也可以有较大的木材基本密度。     

毛白杨多功能新品种选育研究

观测北京、河北、河南３地点的１３年生毛白杨无性系试验林,无性系间的树高、胸径、木材密度、纤维长度等性状的差异均极显著;无性系的遗传稳定性、生长适应性、主干通直系数、树体形态系数、抗病虫指数等性状存在一定差别。材积生长量无性系×地点交互作用极显著,无性系间生长型不同（“广适型”、“高产型”、“低产型”）。无性系的材积生长量、木材密度、主干通直系数、树体系数和抗病虫害指数的重复率分别为０．５８０、０．５３６、０．３９０、０．４１１和０．５００,性状经济权重分别为０．３５、０．２５、０．１０、０．１０和０．２０,建立多功能型无性系的选择指数,其综合育种增益比对照品种≥１５％为中选优良无性系新品种。为北京、河北、河南分别选育出８个、７个和６个优良无性系,平均综合增益分别提高２３．９％、２４．９％和２１．０％;３个地点都中选的３７、３５１、９８３２和１０１２等优良无性系的平均综合育种增益提高２５．１％。所选无性系皆为雄性,显著提高了城乡绿化效应。     

群改杨无性系材性性状遗传分析

对22个群改杨无性系的材性和生长性状进行综合评定,分别采用六项指标（纤维长度、壁腔比、纤维长宽比、密度、树高、胸径）评分法和三项指标（纤维性状、木材密度、材积）评分法,探讨了无性系评定方法。     

江西省优质速生杉木无性系选育研究

在种源、林分和个体选择的基础上,选育出一批杉木无性系,通过13 a的测定表明,各无性系间胸径、树高、单株材积、无性系木材基本密度和纤维长度差异极显著,且具有较大的广义遗传力.根据杉木无性系培育目标,选择出一批生长、材性兼优的杉木无性系以供生产上应用.12年生入选杉木无性系平均胸径、树高和单株材积、木材基本密度和纤维长分别为16.8 cm、11.0 m、0.133 1 m3、0.319 3 g·cm-3和331 7 μm,遗传增益分别为27.99%、8.09%、94.29%、6.72%和4.53%.     

杨树无性系生长与材性遗传变异及综合选择研究

以6年生18个杨树无性系为材料，研究其生长与材性性状的遗传变异及性状问的相关性，结果表明：除纤维含量、纤维长度外，树高、胸径、材积、纤维宽度、长宽比、木材基本密度6性状在无性系间均存在显著差异，各性状重复力在0．6109～0．9053之间，生长性状受高度遗传控制，纤维宽度、长宽比、木材基本密度受中等至强度遗传控制；生长与材性性状遗传具有相对独立性，就杨树纸浆材而言，材性改良应以生长性状的改良为前提；利用灰色关联度分析法评选出XL-90、XL-77、XL-101、XL-75、XL-92共5个综合性状优异的美洲黑杨杂交新无性系，其中XL-90、XL-77表现尤为突出，与I-69比较，其单株材积生长量分别提高60．71％、46．45％，单株纤维产量分别提高51．43％、44．13％。     

杨树新无性系木材性质的变异研究

The wood basic density, cellulose content and fiber form were investigated for all sample trees at breast height (1.3m) in seven poplar clones, and at 0 (butt), 5.6, 9.6, 13.6, 17.6, 19.6 and 21.6 m for clone Nanlin-95 and Nanlin-895, respectively,for providing information on variation patterns of wood density, fiber characteristics and holocellulose content within trees and among clones. The results showed that significant variations about wood density, cellulose content, fiber diameter and the ratio of fiber length to diameter existed among poplar clones examined. Variance analysis indicated that there were significant differences in wood basic density, fiber length, fiber diameter and cellulose content among the growth rings, which had an increasing tendency along the direction from pith to bark. The significant differences also existed in wood basic density, fiber length and fiber diameter at different tree height. The mean wood basic density had a general increase trend with increasing height of trees and the lowest was found at the base, while fiber length and fiber diameter had a general decline pattern with increasing height of trees and the biggest value was observed at the height of 5.6 m. Regression analysis indicated that the relationship between examined wood properties and growth ring number (cambial age), and the relationship between examined wood properties and tree height can be described by polynomial functions.     

Genetic control of fiber properties and growth in triploid hybrid clones of Populus tomentosa

To better understand the genetic control of fiber properties (fiber length [FL], fiber width [FW] and coarseness [C]) and growth traits (tree height [H], diameter at breast height [DBH] and stem volume [SV]) in triploid hybrid clones of Populus tomentosa carr., genetic relationships among selected fiber properties with growth traits were examined in five-year-old clonal field trials located in Yanzhou, Gaotang and Xiangfen, China. In total, 180 trees from 10 clones were sampled from the three sites. The site had a moderate effect on FW, C and tree growth, and a highly significant (P<0.001) effect on FL and fiber length/width (FL/W). Clonal effects were also significant (P<0.05) for all studied traits except for the DBH and SV. Clone×site interactions were significant for all studied traits (except for FL). The estimated repeatability of clonal mean for FL (0.91) was higher than for FW (0.88), C (0.77), FL/W (0.74), H (0.62), DBH (0.61) and SV (0.55). Intersite genetic correlation estimates indicated that fiber properties were more stable than growth traits. Positively insignificant genetic correlation estimates between FL and growth trait and negatively insignificant genetic correlation estimates between C and growth traits were observed. This suggests that in the triploid hybrid poplar, clones selection for growth traits will not result in a significant increase or reduction in the fiber properties.     

日本落叶松纸浆材优良家系多性状联合选择

以12年生日本落叶松自由授粉家系子代测定林为研究对象,对生长、干形和材性性状进行相关分析、通径分析及间接选择研究,利用综合选择指数法开展了日本落叶松纸浆材优良家系选择,并为湖北等亚热带高海拔山区筛选出一批生长、干形和材性兼优的家系,最后提出了日本落叶松纸浆材选育指标及优良家系利用途径。研究表明:生长性状与纤维长度之间存在显著正相关关系,与壁腔比之间存在显著负相关关系,与木材密度、1%NaOH抽出物、综纤维素含量间相关不显著;木材密度与主枝长之间,纤维长度与主枝长、主枝粗和新枝粗之间,晚材壁腔比与主枝粗、主枝长、皮厚、冠幅、新枝数之间,1%NaOH抽出物与主枝长之间存在显著正相关关系。各控制性状对单木材积直接作用的大小依次为:胸径、树高、主枝长、壁腔比、主枝粗、皮厚、冠幅、晚材纤维长和早材纤维长,其中胸径和树高2个性状对单株材积性状的直接作用占材积总变异的63%,虽然冠幅、壁腔比等性状对单木材积的直接控制作用很小,但这些性状通过胸径对单木材积具有很大的间接遗传控制作用,由这9个性状组成的控制系统可以说明单木材积变异的98.66%。     

马尾松无性系种子园半同胞子代变异分析和家系选择

以福建省漳平五一林场马尾松无性系种子园自由授粉子代半同胞家系为研究对象,分析其树高、胸径、材积、冠幅、通直度、高径比、冠径比、木材基本密度、管胞长度和宽度、管胞长宽比等性状,发现家系间除管胞长宽比性状差异显著,其余性状均达差异极显著,暗示着马尾松种子园半同胞家系间存在较为丰富的变异,具选育潜力。8年生林分内各性状的家系遗传力以树高性状为最高(0·691),其次是木材基本密度(0·640)。各性状间的相关性分析结果表明,生长量性状的改良可间接改善通直度,且生长量性状与木材基本密度存在显著正相关关系,木材基本密度与管胞长度和宽度不相关。采用10%入选率,发现5年生林分和8年生林分入选家系一致,材积遗传增益分别达19·74%和19·23%,且5年生林分和8年生林分生长量性状的遗传相关极显著,认为马尾松制浆造纸材短轮伐期的初选年龄可确定为5年生;根据性状遗传力的相对大小和典型相关分析的结果,认为马尾松家系选择可适当注重树高生长量指标。以树高和材积为选择指标,按照10%入选率,兼顾材性和种子园内无性系的开花结实情况,筛选出制浆造纸材短轮伐期优良家系12个。     

长白落叶松纸浆材优良家系多性状联合选择研究

采用胸径木芯与伐倒木取样方法,对24年生长白落叶松优树子代测定林14个处理的生长性状、木芯材质性状、解析木材质性状及其制浆造纸特性进行了研究,结果表明:生长性状、木芯基本密度、木芯管胞性状均存在丰富的变异;家系间生长性状差异极显著;木芯基本密度、木芯管胞性状差异显著,家系遗传力在0.56 0.80之间;解析木基本密度、早材微纤丝角、管胞长度、早材壁腔比、木质素、阿拉伯糖含量、抗张强度、撕裂度等差异极显著,家系遗传力在0.64 0.86之间;晚材率、管胞长宽比、综纤维素含量等差异显著,家系遗传力在0.51 0.61,家系水平的材性改良潜力较大。生长性状与木芯管胞长度呈极显著正相关,与木芯基本密度、管胞长宽比相关不显著;木芯基本密度、管胞长度、管胞宽度以及管胞长宽比与解析木相应性状呈极显著正相关关系,且回归模型较为理想,可以利用胸径木芯各性状值预测其单株值,间接选择和评价纸浆材优良家系。通过生长性状、木芯基本密度和木芯管胞长宽比综合分析选出166、169为优良家系,根据解析木生长、材质以及纸张物理性状选出的优良家系与其结果一致。优良家系材积、木芯基本密度和木芯管胞长宽比的遗传增益(超过对照)分别为48.34%(38.80%)、14.01%(3.63%)和19.89%(6.42%)。     

Early prediction of basic density, shrinking, presence of growth stress, and dynamic elastic modulus based on the morphological tree parameters of Tectona grandis

Significant efforts have been made to improve teak; however, evaluation in juvenile step is limited. The objective of this study was to conduct an early assessment of the wood properties of 4-year-old Tectona grandis. Samples of 36 clones were collected to determine their morphological tree parameters [diameter a breast height (DBH), diameter of the second log, tree height, and log quality]. Presence of growth stress, heartwood percentage, shrinkage (radial, tangential, and volumetric), basic density, and ultrasound velocity (USV) were determined for standing trees, logs, green lumber, and dry lumber. The results indicate that DBH and USV in standing trees can be used to predict elastic module (ED), mainly ED of the standing tree and dry lumber, as well as the possible presence of growth stress. Additionally, growth stress can be predicted by UVS in standing tree. Tangential and volumetric shrinkage were not predicted by tree morphology, but radial shrinkage was predicted by diameter and UVS was not affected by any shrinkage. Basic density was predicted by DBH and UVS measured in log.     

Variation analysis and fine family selection on half-sib progenies from clonal seed orchard of Pinus massoniana

Variation of traits that include height, diameter at breast height (DBH), stem volume, crown width, straightness, wood basic density, tracheid length and width of open-pollinated progenies from clonal seed orchard of Pinus massoniana was analyzed. Differences in traits such as growth, tree form and wood quality among families was significant, which indicated that the improvement potential was notable. Heritability of height and wood basic density reached 0.691 and 0.640, respectively. According to a correlation analysis between traits, stem straightness could be improved indirectly when growth traits are improved; growth traits were significantly correlated with wood basic density; and wood basic density was not significantly correlated with length and width of the tracheid. Under the selection ratio of 10%, genetic gains were 19.74% and 19.32% respectively, and selected families from the progeny test stands at the age of five years and eight years were the same. Genetic correlation of height, DBH and stem volume was also significant at the level of 0.01. These show that selection of the short rotation of P. massoniana for pulp and paper use would be efficient for five-year-old trees. According to the comparison of the heritability of all the traits and taking into account the result of canonical correlation analysis, height could be considered as the most important index for selection. Given wood quality of all the families, and flowering and fruit settings of the clones in a seed orchard, twelve fine families were selected at 10% selection ratio of height growth and stem volume. Translated from Scientia Silvae Sinicae, 2005, 41(6): 43–49 [译自: 林业科学]     

枫香优树14年生子代遗传变异及选择

在福建省洋口国有林场,对来自安徽省南部黄山地区的53个枫香家系进行子代测定,研究家系的遗传变异规律,开展优良家系及单株选择.结果表明:子代试验林14年生时,树高、胸径和单株材积平均值分别为8.05m、6.30cm和0.014 93 m3,生长性状的家系遗传力为0.437～0.576,受中强度的遗传控制,单株遗传力为D.113～0.195,受较弱的遗传控制;木材材性的家系遗传力为0.507～ 0.591,单株遗传力为0.360～0.500,均受中强度的遗传控制.以生长量为主要指标,选出速生优良家系9个,其树高、胸径和单株材积平均值分别为8.85 m、7.22 cm和0.021 49 m3,遗传增益分别为2.86％、5.47％和13.4％;从木材基本密度大于0.500 g·m-3的家系中选出7个生长材性兼优家系,树高、胸径、单株材积和木材基本密度的平均值分别为8.34m、6.79 cm、0.018 58 m3和0.518 g·m-3.所选出的速生优良家系和生长材性兼优家系可作为枫香育种材料在生产上推广应用.     

Within-stem variation in wood properties of triploid hybrids of Populus tomentosa at three clonal trials

To obtain a better understanding of variability in wood properties along the stem in triploid hybrids of Populus tomentosa L., basic wood density (BWD) and fiber traits were measured at four sample heights for each tree sampled from three clonal trials. The BWD showed a similar trend at each site in that density was initially decreased then increased with increasing height position. For fiber traits, the fiber length (FL) decreased linearly with the height of the trunk, whereas fiber width and coarseness changed relatively slightly with increasing height position. In addition, the vertical wood properties distribution patterns were different from clones so that some clones showed more markedly change in height positions than others. The joint analysis indicated significant height position effect for BWD and fiber traits, in accordance with site and clonal effect. Significant site × height interaction was found on wood properties, whereas the significant clone × height interaction was only obtained at fiber width. Moreover, variation in the relationships between FL and BWD or other fiber traits showed no consistent pattern with increasing height position.