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1.
Summary The effectiveness of yield components, harvest index and morphological characteristics as selection criteria among four field pea (Pisum sativum L.) genotypes was examined. Genotypes were grown at a wide range of plant populations (9 to 400 plants m-2) to maximise environmental diversity.Both biological and seed yields approximately doubled from 9 to 100 plants m-2. This response flattened from 100 to 400 plants m-2. Differences among the genotypes were found only at 225 and 400 plants m-2. Analysis of the yield components highlighted the plasticity and large genotype by environment interactions of field peas. The numbers of pods per plant and peas per pod were maximised when each genotype was grown as spaced plants, but the low plant numbers meant seed yields per unit area were at their lowest.Genotypic differences for plant harvest index (PHI) were also only found at 225 and 400 plants m-2. Broad sense heritability estimates indicated that direct selection for PHI was not feasible. The inference from the yield component and PHI results was that alternative selection criteria such as physiological or morphological characteristics may be necessary for improved yield potential. Classification of each genotype indicated that low seedling vigour may be a positive attribute for crop plants of semi-leafless and conventionally leafed field peas. Selection based on any single plant attribute is unlikely to lead to dramatic improvements in the yield potential of field peas. Selection should be based on plant characteristics rather than on differences in yield components.  相似文献   

2.
Genetic variance, heritability, and expected response from selection arc useful in devising alternative methods and criteria of: selection. The objectives of this study were to estimate these for seed yield and its components from 200 F2: populations involving 80 cultivars and lines of mostly small-seeded dry bush bean (Phaseolus vulgaris L.) of habits growth I, II, and III of Middle-American origin. All cultivars and lines were crossed in eight sets of ten parents each in a Design II mating system. The F2 populations, without parents, were evaluated in the field in a replicates-in-sets design at two locations in Colombia in 1983. Estimates of additive genetic variance were significant for yield, pods/m2, seeds/pod, and seed weight. Interaction with environments was also significant. Values for nonadditive genetic variance were not significant for either yield or yield components. The estimates of narrow sense heritability, based on the F2 population mean and unbiased by genotype x environment interaction, were 0.21 ± 0.13 for yield. 20 ± 0.13 for pods/m2, 0.57 ± 0.13 for seeds/pod, and 0.74 ± 0.15 for seed weight. The expected direct response from selection of the top 20 % of F2 populations for yield per se would result in a 4.30 % increase in yield with a correlated response of 0.21 % in seed weight. In contrast, the expected gain from direct selection for seed weight would result in a 11.76 % increase in seed weight with a, correlated gain of 0.28 % for yield. Direct selection for pods/m2 would decrease yield, seeds/pod and seed weight, while direct selection for seeds/pod would reduce pods/m2 and seed weight but increase seed yield by 0.37 %. Data on yield from replicated trials in the early segregating generations could be utilized for identification and selection of promising crosses and families or lines with crosses for dry bean yield improvement.  相似文献   

3.
M. A. Faluyi 《Euphytica》1990,50(3):197-201
Summary Three maturity groups of soybeans (Glycine max L.) were used to investigate the relationship between dry-matter accumulation (DMA) and grain yield (GY), and the prospects for selection of high seed yielding strains among the existing soybean cultivars in a tropical environment. The positive and significant association between DMA and GY (r=0.888***) indicated that selection for high DMA could give gains in GY. However, the higher harvest index (HI=37.5%) for the low seed yielding early maturing genotypes than the more vegetatively endowed and higher seed producing late maturity group, is an indication that excessive DMA could be disadvantageous. Total seed yield per land area for the three maturity groups of soybeans showed that the genotypes with high harvest index and low seed yield could be as good as those ones with high seed and dry-matter yields with low harvest index. The high coefficient of variation which ranged between 18.1 and 59.8% and the heritability estimations which also ranged from 34.4 to 82.2% are indicative of the presence of substantial genetic diversity and there are good prospects for the improvement of the crop through selection.  相似文献   

4.
J. Kumar  P. N. Bahl 《Euphytica》1992,60(3):197-199
Summary The efficiency of indirect selection for seed yield was compared with direct selection for yield per se in chickpea. A total of 2500 single F2 plants, derived from 50 crosses with 50 plants from each cross, were divided into five sub-populations (SP1 to SP5) of 500 plants each by including 10 plants from each of the 50 crosses. The five sub-populations were advanced upto F6 by exercising 10% selection intensity for four successive generations for number of pods per plant in SP1, number of seeds per pod in SP2, seed weight in SP3, seed yield in SP4 and random selection in SP5. The efficiency of direct and indirect selection for yield was evaluated by comparing groups of 50 F6 lines from each sub-population. SP1 and SP3 F6 lines showed higher mean grain yield than the other three methods. SP1 and SP3 were found to be almost equally efficient in developing F6 lines which were significantly superior to the check. This suggests that indirect selection for yield via pod number and seed weight is more efficient than direct selection for yield.  相似文献   

5.
Common bean (Phaseolus vulgaris L.) is grown in regions where water deficits during reproductive development significantly reduce yield. The purpose of this study was to evaluate the association of specific phenological and physiological traits with drought resistance in common bean. Five genotypes were grown under and near a rain shelter in 1988, and an additional 16 progeny lines were included in 1990. Drought stress determined by the drought intensity index was severe (0.78) in 1988 and more moderate (0.63) in 1990. Water stress reduced the expression of most traits with the exception of days to flower and leaf moisture retention capacity. Seed yield among genotypes was reduced from 22 to 71% due to drought. Yield under stress was correlated with yield under nonstress in 1990 and negatively correlated with the drought susceptibility index in 1988. Yield components which exhibited the largest differential genotypic responses to stress were pod and seed number, whereas seed size was more stable. Genotypic variation was detected in all the partitioning indexes, chiefly harvest index and relative sink strength, and the heritability estimates for these traits were high. The limited genetic variability observed among water relations traits and their role in water conservation would restrict their potential use in the selection for drought resistance in common bean. The differential correlations between phenological, biomass and partitioning traits and the indexes for yield and drought susceptibility would suggest that the most effective approach in breeding for drought resistance in common bean would be based first on selection for high geometric yield followed by selection among the high-yielding individuals for low to moderate levels of the drought susceptibility index. This revised version was published online in July 2006 with corrections to the Cover Date.  相似文献   

6.
The phenotypic expression and heritability of quantitative traits vary due to genotypic differences, environmental influences and genotype by environment interactions. The objective of this study was to determine variance components and heritabilities of seed yield and its components in cowpea. Field experiments were conducted at three locations, three planting dates using ten diverse cowpea genotypes during 2004/2005. The experiments were laid out in the randomized complete block design with three replications. Results indicated significant interactions (P ≤ 0.01) among genotypes, locations and planting dates. The genotypic variance contributed ≥50% of the total phenotypic variance for the numbers of days to 50% flowering, seeds per pod, productive branches per plant and seed yield. The heritabilities of the numbers of days to 50% flowering were estimated at 50%, pods per plant (23%), days to maturity (66%), productive branches per plant (53%), 100 seed weight (11%) and seed yield (55%). The presence of considerable degree of genotypic variance among tested genotypes under various environments suggests that success in cowpea breeding could possibly be achieved through direct phenotypic selection.  相似文献   

7.
Genetically superior common bean lines will be efficiently selected with the use of more precise experimental statistics. The objectives of this study were to evaluate the experimental precision of grain yield and primary grain yield components in experiments to register common bean cultivars and identify more appropriate statistics for the selection of genetically superior common bean lines. For this purpose, 21 experiments were performed in a randomized block design in southern Brazil. A total of 156 common bean genotypes of the Mesoamerican and Andean gene pool were assessed between 1998 and 2015. Experimental precisions of grain yield and primary grain yield components were evaluated using 11 statistics. Grain yield, number of pods per plant, number of grains per pod, and mass of 100 grains were evaluated with greater experimental precision by the F-test value for genotype, heritability, coefficient of relative variation, and selective accuracy. Mass of 100 grains presented the highest experimental precision among the traits evaluated in this study. The F-test value for genotype, heritability, coefficient of relative variation, and selective accuracy allow the selection of common bean lines with genetic superiority for grain yield and primary grain yield components. Selective accuracy is the most appropriate statistic to select common bean lines with genetic superiority for grain yield and is recommended for breeding programs.  相似文献   

8.
With the objective of selecting superior recombinant lines of snap bean, four segregating F2 populations were selected by early generation testing and advanced by single seed descent. In a randomized complete block design with two replications within sets 120 F6:7 lines were obtained and evaluated for seven traits of agronomic interest. The mean pod yield of the F6:7 generation was 72% higher than in generation F2, confirming the efficiency of early selection. There was genetic variability in the F6:7 lines within the sets. The selection of superior genotypes was possible by the high estimates of narrow-sense heritability. Highest gains in simultaneous selection of traits were expressed by the indices proposed by Mulamba & Mock for the genetic standard deviation and by Williams, based on tentatively attributed arbitrary weights. Line 3 of the F6:7 generation, with a yield of 8,050.0000 kg ha−1 and pod fiber content of 0.3650% is highly promising to be release as new cultivar.  相似文献   

9.
Summary Three S1-recurrent selection programs (designated HG, HP, and HGP) for increasing protein yield of oat (Avena sativa L.) were conducted for five cycles of selection. The selection criteria in each program (line of descent) emphasized different components of protein yield; high grain yield in HG, both high protein concentration of the groats (caryopsis) and high grain yield in HP, and high protein yield per se in HGP. Heading date and height were restricted to no net change. The objectives of this study were to evaluate the agronomic performance of these three lines of descent and the correlated responses in agronomic traits due to selection. Thirty to sixty random S0-derived lines from each cycle and ten check lines were grown in hill-plot experiments at two locations in 1989. Grain yield increased in all three lines of descent; HG showed the greatest rate of gain followed by HGP and then HP (0.20, 0.10, and 0.07 Mg ha-1 per cycle, respectively). Test weight decreased in HGP by 2.6 kg m-3 per cycle and in HP by 4.0 kg m-3 per cycle, but remained constant in HG. Cycle 5 (C5) means for straw yield, height, and seed weight were not significantly different from the C0 means for any line of descent. Groat percent remained unchanged in HGP, increased from 70.1 to 71.7% in HG, and decreased from 69.9 to 67.6% in HP. Heading date decreased by 3 d in HG and HGP, but remained unchanged in HP. HG compares most favorably with commercial cultivars because of its high yield and acceptable agronomic traits, while HP tends to be low in test weight, seed weight, and groat percent.Abbreviations BM- above ground biomass - GP- groat (caryopsis) percent, fraction of average seed weight that is groat rather than hulls - GPC- groat-protein concentration - GPY- groat-protein yield - GTY- groat yield - GY- grain yield - HD- heading date - HG- selection for protein yield through high grain yield - HGP- selection for protein yield per se - HP- selection for protein yield through high grain yield and high groat-protein concentration - HT- height - PY- protein yield - SDWT- seed weight - SY- straw yield - TWT- test weight  相似文献   

10.
The selection of cultivars for the predominant cropping systems of small farms in the tropics depends to a large extent on the information obtained by testing their performance across the different systems. The main objective of this experiment was to measure the genotype × cropping system (G × CS) interaction for yield and selected agronomic traits of climbing beans (Phaseolus vulgaris L.) grown as sole crop and intercropped with two morphologically contrasting maize (Zea mays L.) cultivars. A secondary objective was to identify the most efficient and productive bean–maize intercrop combinations. Seven climbing bean genotypes were grown as sole crop and intercropped with two maize varieties, BH 140 (Mix. 1) and Guto (Mix. 2), in a factorial arranged Randomized Complete Block Design with three replications at Bako Agricultural Research Center in western Ethiopia. Main effects due to genotype and cropping system (except days to flowering) were significant for all bean traits considered. The genotypes × cropping system interaction terms were also significant for the number of seeds per pod, 100-seed weight, harvest index and seed yield. While bean seed yield significantly correlated with the number of seeds per pod (in Mix. 1) and with harvest index (in both mixtures), positive and significant correlations occurred with the number of pods per plant and 100-seed weight under sole cropping system. The correlation between bean seed yields of Mix. 1 and Mix. 2 and between Mix. 2 and sole crop were positive and significant. No such relationship was found between Mix. 1 and sole crop. The results suggest that selection of suitable climbing bean cultivars for intercropping with maize varieties predominantly grown in the area should be made under the associated culture of the two crops. Intercropping contributed to a significant reduction in seed yield of the bean genotypes due mainly to its adverse effects on the numbers of pods per plant and seeds per pod. The index tLER1 identified most bean–maize genotype combinations of Mix. 2 as biologically more efficient system than Mix. 1. On the other hand, tLER2 values of more than 1.00 for all treatments of Mix. 2 demonstrated higher overall productivity of the intercrop system when the bean genotypes were grown in association with a late-maturing and high yielding maize hybrid BH 140.  相似文献   

11.
The study was carried out to determine combining ability and association of yield and yield components among crosses derived from seven selected cowpea (Vigna unguiculata (L.) Walp) parents. Twenty-one hybrids were generated from diallel crosses excluding reciprocals. Hybrids along with seven parents were studied for combining ability and phenotypic correlations for seed yield and eight yield components. The result indicated that the general combining ability (gca) and specific combining ability (sca) were significant for most characters indicating the importance of both additive and non-additive genetic components. Additive gene action was important for eight characters except pod number, whereas non-additive gene action was not important for nodule number and grain yield. Parental lines IT86D-716 and IT81D-985 were found to be good general combiners for grain yield and other traits. The most promising specific combiners for yield and yield components were from crosses including IT87D-697-2 × IT86D-716, IT88D-867-11 × IT86D-716, IT93K-624 × IT87D-697-2, and IT87D-697-2 × IT92KD-405-1. Significant positive phenotypic correlations were observed between seed yield with pod length (POL), number of pods per plant (PON), and number of seeds per pod (SPD). POL, pod number per plant, SPD, and grain yield were identified as the best selection criteria that could be used in cowpea breeding programs.  相似文献   

12.
The biomass allocation pattern of plants to shoots and roots is a key in the cycle of elements such as carbon, water and nutrients with, for instance, the greatest allocations to roots fostering the transfer of atmospheric carbon to soils through photosynthesis. Several studies have investigated the root to shoot ratio (R:S) biomass of existing crops but variation within a crop species constitutes an important information gap for selecting genotypes aiming for increasing soil carbon stocks for climate change mitigation and food security. The objectives of this study were to evaluate agronomic performance and quantify biomass production and allocation between roots and shoots, in response to different soil water levels to select promising genotypes for breeding. Field and greenhouse experiments were carried out using 100 genotypes including wheat and Triticale under drought‐stressed and non‐stressed conditions. The experiments were set‐up using a 10 × 10 alpha lattice design with two replications under water stress and non‐stress conditions. The following phenotypic traits were collected: number of days to heading (DTH), number of productive tillers per plant (NPT), plant height (PH), days to maturity (DTM), spike length (SL), kernels per spike (KPS), thousand kernel weight (TKW), root biomass (RB), shoot biomass (SB), root to shoot ratio (R:S) and grain yield (GY). There was significant (p < 0.05) variation for grain yield and biomass production because of genotypic variation. The highest grain yield of 247.3 g/m2 was recorded in the genotype LM52 and the least was in genotype Sossognon with 30 g/m2. Shoot biomass ranged from 830 g/m2 (genotype Arenza) to 437 g/m2 (LM57), whilst root biomass ranged between 603 g/m2 for Triticale and 140 g/m2 for LM15 across testing sites and water regimes. Triticale also recorded the highest R:S of 1.2, whilst the least was 0.30 for wheat genotype LM18. Overall, drought stress reduced total biomass production by 35% and R:S by 14%. Genotypic variation existed for all measured traits useful for improving drought tolerance, whilst the calculated R:S values can improve accuracy in estimating C sequestration potential of wheat. Wheat genotypes LM26, LM47, BW140, LM70, LM48, BW152, LM75, BW162, LM71 and BW141 were selected for further development based on their high total biomass production, grain yield potential and genetic diversity under drought stress.  相似文献   

13.
Morpho-agronomic traits and microsatellite markers were used to survey genetic diversity in 115 common bean genotypes that included 70 Indian landraces, 24 released varieties and 21 exotic accessions. Twelve morpho-agronomic traits, namely, days to 50% flowering, leaflet length, leaflet width, pod length, pod width, number of pods per plant, days to maturity, seed length, seed width, number of seeds per pod, 100 seed weight and seed yield per plant were studied. Field data of two consecutive years were subjected to multivariate analysis as proposed by Mahalanobis’s D2-statistics, Tochers method of clustering and combined analysis of variance. Seventeen microsatellite markers were also used to examine genetic diversity at molecular level that showed polymorphic information content (PIC) in the range of 0.00–0.684. Dendrograms based on Euclidean distances and UPGMA analysis showed the presence of majority of released varieties into single cluster, which pointed toward their low genetic base in comparison to indigenous landraces and exotic germplasm. Significant correlation existed between morphological genetic distance and microsatellite genetic distance tested by Mantel test (r = 0.876).  相似文献   

14.
The objective of this study was to characterize physiologically wheat cultivars released in different decades and identify selection criteria for the continued genetic progress in Brazil. Ten cultivars released from 1940 up to 2009 were tested during 2010 and 2011 crop seasons. The following traits were evaluated: grain yield (GY), thousand-kernel weight (TKW), grain number per m−2 (GN), plant height (PH), harvest index (HI), above-ground biomass (BIO), relative Chlorophyll content and leaf gas exchanges. The increase in grain yield was 29 kg ha−1 yr−1 a genetic gain of 0.92%, annually. Grain yield improvement was largely associated with HI (0.94**), number of grains m−2 (0.93**), BIO (0.88**) and reduced PH (−0.93**). The post-anthesis Chlorophyll content, stomatal conductance and pre/post-anthesis photosynthetic rate were positively correlated with GY. Genetic gains of Brazilian wheat are mainly related to the increases of HI, GN, and BIO. These improvements were achieved by reducing PH and raising gas exchanges and chlorophyll content.  相似文献   

15.
Common bean (Phaseolus vulgaris L.), an important food crop in Europe, America, Africa and Asia, is thought to fix only small amounts of atmospheric nitrogen. It contributes significantly to the sustainability of traditional cropping systems because of the predominance of small-scale farmers who cultivate beans in those areas. The objectives of this work were to evaluate bush bean varieties under common agronomic cropping systems and to evaluate breeding lines under low N-fertility sole cropping and intercropping systems. The purpose of the study was to characterize the genotype and cropping system's variability in symbiotic and plant characters and to identify the most suitable genotypes to establish an effective symbiosis with indigenous strains of Rhizobium. No significant differences among the bush bean varieties evaluated under typical fertilization practices were observed for N2-fixation and plant traits except for seed nitrogen. Significant differences among the bean lines studied under low N-fertilization conditions were detected for plant growth,plant component and N2-fixation traits. A significant interaction of bean genotype x cropping system was found for number of nodules per plant and nodule moisture on the bush bean varieties studied, and for days to emergence, days to flowering, end of flowering, shoot length, root dry weight and shoot nitrogen on the bean lines evaluated. Nodulation parameters were correlated positively with the yield components, shoot and root parts and duration of flowering, and correlated negatively with seed crude protein, pod and seed dimensions and seed dry weight. These observations indicate that it may be possible to increase both the symbiotic N2-fixation and seed yield through plant breeding. This revised version was published online in August 2006 with corrections to the Cover Date.  相似文献   

16.
J. Rubio    M. T. Moreno    J. I. Cubero  J. Gil 《Plant Breeding》1998,117(6):585-587
Five pairs of near isogenic lines (NILs) for the gene for single vs. double pod were used to study the effect of this character on yield and its components (seeds per plant, yield per plant and seed weight). The NILs were sown in 2 years at five locations in ihe first year and four locations in the second. Combined analysis of variance was applied to t he data and no significant differences between isolines were found, but there were significant genotype– location interaction effects for yield and seeds per plant. For both characters the double–podded lines were more stable than the single–podded lines. Although there were significant differences between the five genotypes lor yield per planl and seed weight, the differentes between single– and double–podded lines were almost nil. The results indicate that the double–pod gene has a positive effect on the stability of yield and is not linked to any other gene responsible Tor seed size in chickpea  相似文献   

17.
Phenotypic selection under spaced planting is frequently used in forage species. This study aimed to compare the predicted efficiency of direct selection for lucerne seed or forage yield under dense planting with indirect selection based on the same traits or seed yield components evaluated under spaced planting. Sixteen genotypes randomly chosen from a representative sample of locally adapted germplasm were grown for two years as individual clones spaced at 75 cm (density =1.78 plants/m2) and in dense plots formed by one row of four clones spaced at 10 cm (density = 50 plants/m2) using a randomized complete block design with three replications. Indirect selection based on seed yield under spaced planting was just 19% less efficient than direct selection, owing to moderate genetic correlation between plant densities (rg = 0.66) and somewhat higher broad‐sense heritability under spaced planting than under dense planting. The relative efficiency of indirect selection for seed yield in density based on individual seed yield components under spaced planting ranged from modest to very low and was always below 45%. The efficiency of indirect selection for dry matter yield based on yield response under spaced planting was moderate for total yield (64%) and very low for second‐year yield (32%) relative to direct selection.  相似文献   

18.
A. N. Aryeetey  E. Laing 《Euphytica》1973,22(2):386-392
Summary The inheritance of yield components was studied in a cross of two varieties of cowpea (Vigna unguiculata), and the relationships among the components and yield were examined in the F2 of a cross and in a trial of 22 varieties. The main objective was to determine the suitability of using the components as selection criteria for increasing seed yield. All the components appeared to be under polygenic control and transgressive segregation in the F2 was observed for pod length and number of seeds per pod. Narrow sense heritability estimates ranged from 19.8% for number of pods per plant to 60.3% for pod length. Most of the genotypic correlations between pairs of yield components were significant. Correlation of yield per plant was negative with pod length, but positive with the other components. Number of pods per plant was consistently correlated with yield. However because of low heritability for number of pods per plant, this component may only be used as a preliminary selection criterion while the final selection is based on yield itself.  相似文献   

19.
Two experiments were conducted under field conditions to evaluate reproductive abscission, seed yield and yield components of three cowpea [Vigna unguiculata (L.) Walp.] genotypes. In the first experiment, level of abscission and yield of two cultivars, California Blackeye Pea No. 5 (CA-5) and Speckle Purple Hull (SPH), and one experimental line (AZ-54) were studied. In the second experiment, effect of drought stress on abscission at three nodal positions, seed yield, and yield components of CA-5 were studied. Abscission in both experiments was determined by counting scars left by dropprd reproductive structures including floral buds, open flowers, and immature pods. Abscission of CA-5 and AZ-54 in the first experiment ranged between 68 and 76 % while that of SPH ranged between 86 and 89 %. CA-5 and AZ-54 retained two to three pods per peduncle, and SPH retained only one mature pod per peduncle. Average seed yields of SPH and AZ-54, respectively, were 45 and 50 % of CA-5. Drought stress in the second experiment did not affect production of floral buds Peduncle?1 (average of 10) but significantly increased percent reproductive abscission and decreased pod retention of CA-5. Abscission in the bottom two-third nodes increased from 82 % in well-irrigated plants to 93 % in non-irrigated plants. This increase in abscission corresponded to nearly 60 % reduction in pod retention. The number of pods per peduncle in the bottom two-third nodes decreased from 1.9 in well-irrigated plants to only. 77 in non-irrigated plants. The increase in abscission and decrease in pod retention with increasing intensity of drought was greatest in the bottom one-third nodes. Drought stress did not affect abscission and pod retention in the top one-third nodes. Stress also decreased peduncles plant?1, seeds pod?1, and dry matter and seed yield plant?1 but did not affect seed weight and harvest index. The decrease in seed yield was largely due to reductions in pods plant?1 and seeds pod?1. The reduction in the number of pods and, therefore, seed yield due to stress was because of reductions in the number of peduncles plant?1 and increases in reproductive abscission. It is concluded external conditions that increase abscission beyond that of normal occurrence affect seed yield adversely.  相似文献   

20.
Paolo Ranalli 《Euphytica》1996,87(2):127-132
Summary The effect of a recurrent selection procedure for improving seed yield per plant was evaluated in common bean. Progress was registered after three cycles of recurrent selection with S2 progeny evaluation performed on the grain yield/plant. Gain from selection was estimated by field testing 45 random S2 lines from each of the C0, C1 and C2 populations. Response to selection for seed yield/plant was 6 g from C0 to C1 and 4.4 g from C1 to C2. The seed yield improvement was due principally to an increase in pods/plant; little contribution was provided by the 1000-seed weight and seeds/pod traits. Broad-sense heritability and genotypic variance for grain yield remained high in all cycles which suggests further gain from additional selection cycles. Grain yield/plant is significantly correlated to pods/plant and seeds/pod traits in all cycles, indicating that plants with favourable combinations of these traits could be identified. The proposed recurrent selection procedure appears to be effective in improving the population and extracting superior genotypes for varietal development.  相似文献   

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