Effect of altering the grazing interval on growth and utilization of grass herbage and performance of dairy cows under rotational grazing

The effects of short grazing intervals in the early part of the grazing season on the growth and utilization of grass herbage, and the performance of grazing dairy cows, in a rotational grazing system were examined. Seventy-six cows were allocated to two grazing treatments: a normal rotation treatment (20-d rotations for the first 60 d) and a short rotation treatment (12-, 8-, 8-, 8-, 12- and 12-day rotations). Thereafter, both treatments had the same grazing interval and over the season as a whole both treatments received the same amount of nitrogen fertilizer and were stocked at the same rate. The short rotation treatment significantly reduced pre- and post-grazing sward heights and pre-grazing herbage mass in May and June. Total herbage production was significantly lower on the short than the normal rotation treatment as a result of a significant reduction in the growth rate of herbage in May and June. The short rotation treatment had a significantly lower milk output per cow. Grazing shorter swards, as a result of shorter rotations, significantly reduced herbage intake, reflecting reductions in intake per bite, grazing time and total bites per day. Treatment had no significant effect on herbage quality or pre- and post-grazing sward height in August and September, despite the increased grazing severity in May and June with the short rotations. The severity of rotation length in this instance had a detrimental impact on animal performance, whereas a more modest reduction in grazing interval may control herbage production, without reducing animal performance.     

The combined use of fertilizer nitrogen and white clover on grazed ryegrass swards

A 1-year grazing experiment with dairy cows is described in which the milk yield and herbage intake from a sward of S23 perennial ryegrass alone receiving fertilizer N at an annual rate of 360 kg ha⁻¹ were compared with those from a ryegrass-white clover var. Blanca sward given a total N application of 180 kg ha⁻¹. Both treatments gave virtually the same total milk yield of about 12500 kg ha⁻¹ over a 20-week grazing season.     

Concentrate supplementation of grazing dairy cows. 2. Effect of concentrate composition on herbage intake and milk production

The effect of feeding either traditional concentrates containing starch or high quality fibrous concentrates on the performance of grazing dairy cows was examined in a trial in which cows were given concentrates with either 350 g starch and sugars (kg dry matter (DM))^-1 (high-starch) or 100 g starch and sugars (kg DM)^-1 (high-fibre). The swards used consisted predominantly of perennial ryegrass and were usually aftermaths following cutting. Each area was grazed for 3 or 4 d at each grazing and a two-machine sward-cutting technique was used for estimating herbage intake.
The effect of concentrate composition on the herbage intake of grazing cows at a high daily herbage allowance of 28 kg OM above 4 cm cutting height was investigated in 1983 and 1984. With 54 kg OM d^-1 of high-starch concentrates the mean herbage intake was 11·5 kg OM d^-1 per cow while cows fed 5.3 kg d^-1 of high-fibre concentrates consumed on average 12–6 kg OM d^-1. The mean substitution rate of herbage by concentrates was reduced from 0·45 kg herbage OM (kg concentrate OM)^-1 with the high-starch concentrate to 0·21 with the high-fibre concentrates.
The effect of the treatments on milk production was studied in 1984. The cows consumed 5·5 kg OM d^-1 as concentrates and grazed at a lower herbage allowance of 19 kg OM above 4 cm cutting height. With high-fibre concentrates milk production and 4% fat-corrected milk production were 13 and 1·8 kg d^-1, respectively, higher than with the high-starch treatment. The daily live weight gain with the high-starch concentrates was 0·17 kg per cow more than with the high-fibre concentrates.     

Effect of grazing management on herbage protein concentration,milk production and nitrogen excretion of dairy cows in mid‐lactation

The objective of this experiment was to use diurnal and temporal changes in herbage composition to create two pasture diets with contrasting ratios of water‐soluble carbohydrate (WSC) and crude protein (CP) and compare milk production and nitrogen‐use efficiency (NUE) of dairy cows. A grazing experiment using thirty‐six mid‐lactation Friesian x Jersey cows was conducted in late spring in Canterbury, New Zealand. Cows were offered mixed perennial ryegrass and white clover pastures either in the morning after a short 19‐day regrowth interval (SR AM) or in the afternoon after a long 35‐day regrowth interval (LR PM). Pasture treatments resulted in lower pasture mass and greater herbage CP concentration (187 vs. 171 g kg^?1 DM) in the SR AM compared with the LR PM but did not affect WSC (169 g kg^?1 DM) or the ratio of WSC/CP (1·0 g g^?1). Cows had similar apparent DM (17·5 kg DM cow^?1 d^?1) and N (501 g N cow^?1 d^?1) intake for both treatments. Compared with SR AM cows, LR PM cows had lower milk (18·5 vs. 21·2 kg cow^?1 d^?1), milk protein (0·69 vs. 0·81 kg cow^?1 d^?1) and milk solids (1·72 and 1·89 kg cow^?1 d^?1) yield. Urinary N concentration was increased in SR AM, but estimated N excretion and NUE for milk were similar for both treatments. Further studies are required to determine the effect of feeding times on diurnal variation in urine volume and N concentration under grazing to predict urination events with highest leaching risk.     

Manipulation of herbage production by altering the pattern of applying nitrogen fertilizer

The redistribution of herbage production during the growing season to synchronize herbage supply with feed demand by livestock by altering the application pattern of a range of nitrogen (N) fertilizer rates was studied. Application rates of N were 50, 150 and 250 kg N ha^?1 per annum and patterns were with 0·60 of N fertilizer applied before June (treatment RN) and with only 0·20 of N fertilizer before June (treatment IN). Treatments were imposed in a cutting (simulated grazing) experiment (Experiment 1), which was conducted for 2 years and a grazing experiment (Experiment 2) which was conducted for 3 years. In both experiments, herbage production was reduced in April and May and increased in the June–October period on treatment IN relative to RN, but annual herbage production was not significantly affected except in the third year of Experiment 2, when treatment RN had significantly (P < 0·05) higher herbage production than treatment IN. Crude protein (CP) concentration of herbage was lower in April and May on treatment IN than treatment RN. However, CP concentration of herbage was rarely below 150 g kg^?1 DM and so it is unlikely that livestock productivity would be compromised. On treatment IN, concentrations of CP in herbage were higher in the late summer than on treatment RN, which may increase livestock productivity during July and August when livestock productivity is often lower. Altering the strategy of application of N fertilizer did not affect in vitro dry matter digestibility of herbage.     

Sodium fertilizer application to pasture. 10. A comparison of the responses of dairy cows with high and low milk yield potential

Current recommendations for the intake of sodium of lactating dairy cows are related to milk yield. A study was conducted to compare the responses of cows of high and low milk production potential to the application of sodium fertilizer to grazed perennial ryegrass pasture. The application of sodium fertilizer increased the intake of herbage dry matter (DM), the time that cows spent grazing and the biting rate. It also increased the concentration of sodium, magnesium and calcium in herbage and decreased the concentration of potassium. Applying sodium fertilizer increased milk yield and milk fat concentration and decreased somatic cell count in the milk of cows of low-production potential only, whereas it increased persistency of milk production in the cows of high-production potential. The concentration of lactose in milk increased in both groups after the application of sodium fertilizer. It is concluded that the optimum dietary sodium concentration for grazing cows does not increase with milk yield, and that most immediate advantage will be gained from increasing the sodium concentration in herbage for low-yielding cows.     

Implications of nitrogen fertilizer applications and extended grazing for the N economy of grassland

The response of swards which have been previously grazed to N fertilizer applied in early February was studied in two experiments in Northern Ireland. The effect of N fertilizer applied at a range of dates in autumn and spring on swards for out-of-season utilization was studied in a further experiment. Deep soil coring was also undertaken, subsequent to grazing with dairy cows, in grazed and protected areas in November and March to investigate the effect of out-of-season grazing on soil mineral N levels.
Dry-matter (DM) yield response to early spring N application in previously grazed swards was low, with no effect on DM yield in February or March. Progressively delaying N application (and commencement of herbage accumulation) in autumn from 8 September until 18 October reduced herbage availability in late autumn and early spring but increased leaf lamina content. The greater the amount of herbage accumulated to 1 December, the lower the tiller density in the following April.
N fertilizer had a greater impact on soil mineral N in spring than in late autumn/early winter, suggesting that fertilizer N was more prone to loss in the latter. Soil mineral N was not significantly affected by out-of-season grazing.
It is concluded that in well-fertilized, previously grazed swards response to N for out-of-season herbage is low and the probability for N loss is increased. Herbage quality will decline and the sward may be damaged if about 2 t DM ha⁻¹ or more of harvestable herbage accumulates for use in winter or in early spring.     

Effect of herbage mass and allowance on sward characteristics,milk production,intake and rumen volatile fatty acid concentration

The objective of this study was to examine the effect of herbage mass and daily herbage allowance (DHA) on sward characteristics and animal performance, dry‐matter intake, rumen pH and volatile fatty acid production of unsupplemented spring‐calving dairy cows throughout the main grazing season. Sixty‐eight Holstein‐Friesian dairy cows were randomly assigned across four treatments (n = 17) in a 2 × 2 factorial design. Two swards were created with different levels of pre‐grazing herbage mass [allocated above 4 cm (>4 cm); 1700 kg DM ha^?1 (medium; M) or 2200 kg DM ha^?1 (high; H)] and two levels of DHA (>4 cm; 16 or 20 kg DM per cow d^?1). An additional eight lactating ruminally cannulated Holstein–Friesian dairy cows were randomly assigned to each treatment in a replicated 4 × 4 Latin square design. Sward and animal measurements were collected across four periods each of 1 week duration in April and May (PI) and July and August (PII). Maintaining the medium‐mass sward across the season improved the nutritive value of the sward in the latter part of the grazing season compared with high‐mass swards, thus resulting in increased animal intakes and milk production throughout PII. The higher organic matter digestibility of the medium‐compared with high‐masses during PII indicates that grazing severity and herbage mass in the spring to mid‐summer period will determine sward quality parameters in the late summer period.     

GrazeIn: a model of herbage intake and milk production for grazing dairy cows. 2. Prediction of intake under rotational and continuously stocked grazing management

Decision support tools to help dairy farmers gain confidence in grazing management need to be able to predict performance of grazing animals with easy‐to‐obtain variables on farm. This paper, the second of a series of three, describes the GrazeIn model predicting herbage intake for grazing dairy cows. The model of voluntary intake described in the first paper is adapted to grazing situations taking account of sward characteristics and grazing management, which can potentially affect intake compared to indoor feeding. Rotational and continuously stocked grazing systems are considered separately. Specific effects of grazing management on intake were quantified from an extensive literature review, including the effect of daily herbage allowance and pre‐grazing herbage mass in rotational grazing systems, sward surface height in continuously stocked grazing systems, and daily time at pasture in both grazing systems. The model, based on iterative procedures, estimates many interactions between cows, supplements, sward characteristics and grazing management. The sensitivity of the prediction of herbage intake to sward and management characteristics, as well as the robustness of the simulations and an external validation of the GrazeIn model with an independent data set, is presented in a third paper.     

Effects of post‐grazing herbage height and concentrate feeding on milk production and major milk fatty acids of dairy cows in mid‐lactation

Milk fatty acids (FA) were compared in mid‐lactation dairy cows in four feeding systems combining grazing management and supplementation. The four treatments were factorial combinations of compressed herbage grazed to 3·7 or 4·6 cm post‐grazing height, with or without concentrate feeding (3·6 kg cow^?1 d^?1). Milk yield and composition were measured for four groups of eight Friesian × Jersey dairy cows over 3 weeks in mid‐lactation for cows that had grazed treatments for 64 d from early spring. Milk yield was higher in cows fed concentrate plus herbage (23·9 kg d^?1 cow^?1) than cows fed herbage only (20·3 kg d^?1 cow^?1). Milk fat percentage was higher in cows fed herbage only (5·5%) than that fed herbage plus concentrate (5·1%). Milk protein percentage was higher in cows fed herbage plus concentrate (4·0%) than that fed herbage only (3·7%). The concentrations of conjugated linoleic acids c9, t11, C18:0, C18:1 t11 and C18:2 t9, c12 FA were lower where concentrate was fed. The concentrations of C18:1 t10, C18:1 t5, t8 and C18:2 c9, c12 FA were higher where concentrate was fed. The concentrations of C18:1 c6, C18:1 c9, C18:1 t9 and C18:3 c6,9,15 were unaffected by concentrate feeding. Post‐grazing herbage height had no significant effect on milk yield or concentration of milk FA. Provided dairy cows are harvesting leafy material of similar nutrient and FA concentration, post‐grazing herbage height does not appear to alter milk FA and the supply of high energy concentrates is more influential on milk FA profiles.     

Influence of defoliation regime on herbage production and characteristics of intake by dairy cows as affected by grazing intensity

Three contrasting defoliation regimes for dairy cows—four cuts annually, severe rotational grazing and lax rotational grazing integrated with cutting—were compared in terms of herbage production, selection and intake per cow. Lax and severe grazing were compared simultaneously and grazing intensity was characterized by means of changes in herbage mass and sward height during grazing.
Herbage growth and yield were similar under cutting, severe grazing and lax grazing/cutting (120 t organic matter (OM) ha⁻¹). Herbage quality and leaf production were highest with severe grazing, which also had a less marked seasonal pattern of growth. The herbage intake per cow was 111 kg OM d⁻¹ with severe grazing and proportionately (0-20) higher at lax grazing/ cutting. 050 of the herbage yield was harvested by cutting at lax grazing/cutting. Selectivity was described with high certainly by the nutrient content prior to defoliation and by the degree of defoliation. Models to quantify this are presented.
Grazing intensity could be described by the size and degree of utilization of the fouled area, and herbage intake was quantified by means of herbage height before and after grazing. Within the range of 80-240 mm sward height prior to grazing, height measurements led to realistic and more accurate estimates than those obtained by measuring organic matter disappearance.
Herbage utilization was depressed significantly by increasing maturity of herbage due to lower nutritive value, but in particular due to reduced green leaf content and increased reproductive development. If sward height did not exceed 250 mm at any time, good utilization by grazing could be obtained.     

GrazeIn: a model of herbage intake and milk production for grazing dairy cows. 3. Simulations and external validation of the model

GrazeIn is a model for predicting herbage intake and milk production of grazing dairy cows. The objectives of this paper are to test its robustness according to a planned arrangement of grazing and feeding scenarios using a simulation procedure, and to investigate the precision of the predictions from an external validation procedure with independent data. Simulations show that the predicted effects of herbage allowance, herbage mass, herbage digestibility, concentrate supplementation, forage supplementation and daily time at pasture are consistent with current knowledge. The external validation of GrazeIn is investigated from a large dataset of twenty experiments representing 206 grazing herds, from five research centres within Western Europe. On average, mean actual and predicted values are 14·4 and 14·2 kg DM d^?1 for herbage intake and 22·7 and 24·7 kg d^?1 for milk production, respectively. The overall precision of the predictions, estimated by the mean prediction error, are 16% (i.e. 2·3 kg DM d^?1) and 14% (i.e. 3·1 kg d^?1) for herbage intake and milk production, respectively. It is concluded that the GrazeIn model is able to predict variations in herbage intake and milk production of grazing dairy cows in a realistic manner over a wide range of grazing management practices, rendering it suitable as a basis for decision support systems.     

Setting management limits for the production and utilization of herbage for out-of-season grazing

Three experiments were carried out on perennial ryegrass‐dominant swards to provide a basis for recommendations for the limits to (a) building up and timing of utilization of a herbage ‘bank’ for out‐of‐season grazing and (b) duration and intensity of early spring grazing in the United Kingdom and Ireland. In experiment 1, the effect of regrowth interval (from 7 September, 20 October, 17 November or 15 December) in autumn on herbage accumulation, leaf turnover and on subsequent spring growth was investigated. Swards regrown from early September reached maximum herbage mass (about 3 t ha^–1 DM) and leaf lamina content in mid‐November, by which time senescence rate exceeded rate of production of new leaves. New leaf production and senescence rates were greater in swards remaining uncut until December than in those cut in October or November. Time of defoliation up to December had no effect on spring herbage mass in the subsequent spring. Defoliating in March reduced herbage mass in late May by less than 20%. Experiment 2 investigated the progress in herbage growth and senescence in swards regrowing from different times in late summer and autumn to produce herbage for utilization beyond the normal grazing season. Treatments in a randomized block design with three replicates were regrowths from 19 July, 8 August, 30 August and 20 September. Based on a lower ceiling of leaf and total herbage mass being reached with progressively later regrowths, beyond which leaf senescence generally exceeded leaf production and herbage mass declined, it was concluded that currently recommended rotation lengths for this period should extend from 3 weeks in late July to 8 weeks for swards previously grazed in mid‐September. In both experiments, leaf senescence commenced earlier (by one leaf‐age category) than previously published estimates and so brought forward the time at which senescence rates balanced leaf growth rates. In experiment 3, designed to evaluate the effect of daily grazing period and intensity in early spring on herbage regrowth, dairy cows grazed successive plots (replicates) for 2 or 4 h each day at two intensities (target residual heights of 5 or 7 cm) in March to mid‐April. Regrowth rate was similar in all treatments including the ungrazed control, despite soil moisture content being relatively high on occasions. Tiller density was significantly reduced in May by grazing plots in early or mid‐April. It is concluded that in autumn there are limits to which rotation lengths should be extended to produce herbage for out‐of‐season grazing owing to attainment of ceiling yields. Although utilization in early spring may reduce herbage availability in spring, out‐of‐season utilization need not reduce herbage growth rates in early spring.     

Does pre‐grazing herbage mass really affect herbage intake and milk production of strip‐grazing dairy cows?

The effect of pre‐grazing herbage mass (HM) on herbage intake and milk production of strip‐grazing dairy cows is usually studied at the same herbage allowance (HA). In the literature, the effect of HM seems to be affected by the cutting height above which HA is estimated. The aim of this 2 × 3 factorial study was to determine whether the effect of HM (low vs high HM) on herbage intake, milk production and grazing behaviour of dairy cows is affected by the HA estimation height (ground level vs 2·5 vs 5 cm). Two HMs were compared in three different ways: at same HA above ground level (SHA₀), at same HA above 2·5 cm (SHA₂) and at same HA above 5 cm (SHA₅). During two consecutive years, twenty‐four Holstein‐Friesian dairy cows in mid‐lactation were assigned to one estimation height in an incomplete switchback design, with two 14‐d periods. There was an interaction between HM and estimation height for herbage intake and milk production. The effect of HM on herbage intake was positive, null and negative when HMs were compared at SHA₀, SHA₂ and SHA₅, respectively. This study may have practical implications on future research for studying, directly or indirectly, the effect of pre‐grazing HM under strip‐ or rotational‐grazing management, and on modelling herbage intake at grazing.     

The effects of offering a range of forage and concentrate supplements on milk production and dry matter intake of grazing dairy cows

An experiment was undertaken to examine the effect of supplement type on herbage intake, total dry matter (DM) intake, animal performance and nitrogen utilization with grazing dairy cows. Twenty‐four spring‐calving dairy cows were allocated to one of six treatments in a partially balanced changeover design with five periods of four weeks. The six treatments were no supplement (NONE), or supplementation with either grass silage (GS), whole‐crop wheat silage (WS), maize silage (MS), rapidly degradable concentrate (RC) or slowly degradable concentrate (SC). Cows were rotationally grazed with a mean herbage allowance of 20·5 kg DM per cow per day, measured above 4 cm. Forage supplements were offered for approximately 2 h immediately after each morning milking, with cows on NONE, RC and SC treatments returning to the grazing paddock immediately after milking. Cows on treatment MS had a significantly higher supplement DM intake than the other treatments but a significantly lower grass DM intake than the other treatments, resulting in no significant difference in total DM intake when compared with cows on treatments WS, RC and SC. Concentrate type had no significant effect on herbage intake, milk yield, milk composition or yield of milk components. The yield of milk fat and milk protein was significantly higher on treatments MS, RC and SC compared with treatments NONE, GS and WS. The results indicate that despite a relatively high substitution rate, maize silage can be a useful supplement for the grazing dairy cow.     

Relationship between digestibility and faecal nitrogen in sheep and cows offered herbage ad libitum

A comparison was made of the digestibility, voluntary intake and concentration of N in faecal organic matter in castrated male sheep and non-lactating cows offered herbage ad libitum. The sheep ate 22% more herbage DM per kg liveweight than the cows. In general, the sheep and cows digested the organic matter of the herbage to the same extent, although on two occasions significantly higher digestibilities were observed in cows than in sheep. Similar concentrations of N in faecal organic matter in sheep and cows were matched by similar digestibilities of herbage organic matter. Regression equations relating herbage organic matter digestibility to faecal N concentration were not significantly different between sheep and cattle.     

Sodium fertilizer application to pasture. 9. The effects of combined or separate applications of sodium and sulphur fertilizers on herbage composition and dairy cow production

An experiment was conducted to examine the response of herbage grazed by dairy cows to sodium fertilizer applied with or without sulphur fertilizer. The residual effects of applying sodium fertilizer in the previous year were also evaluated. The application of sodium or sulphur fertilizer did not affect herbage growth or height, but the application of sulphur fertilizer increased crude protein content of herbage. The increase in sodium content with application of sodium fertilizer was small, but was greater when sodium fertilizer had also been applied in the previous year. Herbage potassium was increased when sodium fertilizer was applied in the year of the experiment, but only if sodium had not been applied in the previous year. Application of sulphur fertilizer increased herbage sulphur content and reduced the contents of boron, chromium, molybdenum and nickel. Cows grazing pasture that had received sodium fertilizer had increased milk yields and the content of lactose in milk, whereas those grazing pasture that had received sulphur fertilizer application had reduced milk yields and the content of milk fat.     

Incorporating grazing behaviour measurements in models to predict herbage intake by grazing dairy cows

Models to predict herbage intake were constructed using 168 dairy cow records from three grazing experiments. Variables included fell into three categories: animal state, sward state and animal behaviour. Linear regression models of varying complexity were obtained by removing variables from the best fitting model to reflect progressive lack of information availability on farms. Thus, behavioural variables were removed first, followed by sward surface height and milk fat concentration. Models were subject to outlier analysis and collinearity tests. Equivalent models were constructed using ridge regression to minimize collinearity problems. They were tested using 20 Holstein–Friesian dairy cows continuously stocked on a perennial ryegrass sward. A `best practice' treatment [7 cm sward surface height (SSH), 6 kg day⁻¹ concentrate (C)] was used together with treatments of SSH5/C6, SSH7/C8, SSH7/C0 and SSH9/C6. The best model accounted for 0.37 of the variance in the estimation data and contained the following variables: concentrate intake, milk yield, milk fat concentration, days in milk, sward surface height and chewing rate while ruminating. Model performance against test data was generally poor. This was mainly because of consistent underprediction of herbage intake, caused in part by the higher average herbage intakes in the test data compared with the estimation data.     

Effect of regrowth interval on the productivity of swards defoliated by cutting and grazing

Over three grazing seasons (1984-1986) a sward of perennial ryegrass, cv. Talbot, which received a total of 336 kg N ha^-1 each season, was cut or grazed with ewes at 3- or 4-week intervals on a rotational basis.
Sward productivity was higher under cutting than under grazing irrespective of the interval between defoliations. Under cutting, mean herbage organic matter (OM) yields over both intervals were 8·66, 9·62 and 8·17 t ha-¹ in 1984, 1985 and 1986 respectively while under grazing the corresponding yields were 7·65, 8·63 and 7·50 t ha^-1. The mean annual yield of herbage defoliated at 3-week intervals was 7·50, 8·64 and 7 ·20 t OM ha^-1 compared with 8·80, 9·60 and 8·46 t OM ha^-1 for swards defoliated at 4-week intervals in the three years respectively.
The nitrogen (N) content of both the available and the residual herbage was consistently higher under grazing than under cutting. Available herbage contained 31·3 and 27·7 g N kg OM^-1 and residual herbage 26·1 and 22·7 g N kg OM^-1 under grazing and cutting respectively.
The mean yield of N under cutting was 284 kg ha^-1 compared with 304 kg ha^-1 under grazing. Defoliation interval had no effect on N yield, the overall mean yield being 294 kg ha^-1 under both 3- and 4-week defoliation intervals. The effect of the treatments on tiller population was slight and inconclusive.
The process of grazing reduced yield probably as a result of damage to the sward through trampling; the positive effect of excretal N on yield was minimal on account of the short grazing periods.     

Sodium fertilizer application to pasture. 8. Turnover and defoliation of leaf tissue

The effects of application of sodium fertilizer on the turnover and defoliation of leaf tissue were investigated in a perennial ryegrass (Lolium perenne) pasture grazed by dairy cows. Eight plots were allocated to treatments either with or without sodium fertilizer, with the former receiving 32 kg Na ha^–1 applied in five applications of NaNO₃ over the grazing season. An equivalent amount of nitrogen was given to the controls as ammonium nitrate, the application of which was reduced in the sodium treatment to equate nitrogen fertilizer applications for the two treatments. In nine periods between April and September, marked tillers were recorded to measure leaf turnover, leaf lamina growth and specific leaf weight and, when combined with tiller density measurements, gave an estimate of herbage flux for the sward. The defoliation and net growth of the marked tillers were monitored at 3-day intervals and the data were combined with tiller density and specific leaf weight data to determine the intake of the expanding, penultimate and oldest live leaf laminae. Sodium fertilizer application did not affect the rate at which leaves appeared, but it retarded their rate of disappearance. The extension rate and the specific weight of green laminae were both increased by sodium fertilizer application and therefore the net gravimetric growth rate was increased. Tiller density was not affected by sodium fertilizer application and hence the estimated herbage growth and net herbage flux were increased by sodium fertilizer application. Application of sodium fertilizer did not affect lamina length, and in both treatments the penultimate laminae were approximately twice as long as expanding and oldest live laminae. Defoliation frequency decreased from the expanding to the oldest live laminae in the control treatment without sodium. Sodium fertilizer application increased the frequency of defoliation of the oldest live leaf and also increased the length of the expanding leaf that was defoliated. For penultimate leaf laminae sodium fertilizer application reduced the defoliation frequency and length of foliage grazed. The dry-matter (DM) intake of the oldest live laminae was increased by the application of sodium fertilizer. It is concluded that sodium fertilizer application increases net herbage growth both by increased extension rate of leaf laminae and specific leaf weight and by delayed laminae senescence, and that it increases herbage DM intake by increasing the defoliation frequency of the oldest live leaf laminae.