Forage availability from a temperate pasture managed with intensive rotational grazing

Successful integration of rotational grazing into livestock production systems requires estimates of pasture growth rates for feed budgeting of daily animal intake. By matching livestock nutrient demand with forage availability, over-feeding of supplements can be minimized, which reduces feed costs and the need lo manage surplus nutrients, A three-year grazing study was carried out on a Kentucky bluegrass ( Poa pratensis L.)-dominant pasture to estimate the daily quantity of herbage available to cattle in an intensive, rotational grazing system. Herbage production, species composition, and forage quality were determined in each of the six grazing cycles in a year, from April until September. The average length of a grazing cycle was 28·6 d, with 2·7 d for duration of grazing on a paddock. Pre-grazing and post-grazing sward heights, measured with a plate meter, were 14 and 7 cm, and the corresponding herbage masses were 1955 and 775 kg DM ha⁻¹ respectively. Under adequate soil moisture during 1989, herbage available for daily intake was 53 kg ha⁻¹ from April until mid- August, declining to approximately 32 kg ha⁻¹ d⁻¹ by the end of September. Distribution of this herbage was fairly uniform until the end of August. However, a dry summer in 1991 reduced herbage availability to 15 kg ha⁻¹ d⁻¹. Bluegrass and white clover ( Trifolium repens L.) formed 70% of the herbage yield during the period April–June. Later in the season, dead matter and other species increased, reducing the contribution of bluegrass and clover to approximately 60% of total dry matter. While these pastures have the potential to provide significant amounts of forage for 5–6 months in a year, additional on-farm forage reserves are needed during periods of water stress.     

The utilization of N fertilizer, applied to perennial ryegrass/white clover pasture growing on a humus iron podzol in N.E. Scotland

Applications of N fertilizer (0.40, 80, 120 and 240 kg N ha⁻¹ year⁻¹ in dressings of 40 or 80 kg N ha⁻¹) were made to a perennial ryegrass/white clover ( Lolium perenne L. Trifolium repens L.) pasture growing on a humus iron podzol reclaimed from heather ( Calluna vulgaris L.) moor in 1982 and 1983. Where no N was applied, estimates of Ni fixation and mineralization were almost equal, being approximately 50 kg N ha⁻¹ year⁻¹ from each source. Apparent efficiencies of fertilizer use were generally low; for each dressing they ranged between –0·7 and 25·5 kg dry matter (DM)kgN⁻¹. Also, responses to N fertilizer were affected by previous dressings. The net N recovery in harvested herbage from application of 120 kg N ha⁻¹ year⁻¹ was 30 kg N ha⁻¹ year⁻¹.     

Sward dynamics of a smooth-stalked meadowgrass dominantwhite clover sward rotationally grazed by cattle and/or sheep

This experiment was carried out to study the responses of sward components (particularly white clover, Trifolium repens ) to grazing management in a natural sward dominated by smooth-stalked meadowgrass ( Poa pratensis ) syn. Kentucky bluegrass. Treatments during two grazing seasons (1989–90) were: cattle grazing alone (C); cattle grazing followed by topping (CT); cattle grazing followed by sheep grazing (CS); and sheep grazing alone (S). Mean target pre- and post-grazing herbage masses were 2200 and 1100 kg DM ha⁻¹, estimated by single-probe electronic capacitance meter. Sward component dynamics were monitored using turf dissections, marked white clover stolons, and ring-toss white clover leaf counts. Component and sward data for the C, CT, CS and S treatments respectively, were: number of white clover leaves m⁻², 1295, 1384, 1408, 900 (s.e. ± 108); number of leaves per growing point, 3·2, 3·4, 3·0, 2·8 (s.e. ± 0·2); herbage accumulation (t DM ha⁻¹), 5·16, 5·02, 5·87, 8·28 (s.e. ±0 08); rejected herbage (% pasture area) 39·7, 7·7, 16·0, 0 (s.e. ± 75); and annual net herbage production (t DM ha⁻¹) 3·39, 4·35, 4·99, 8·28 (s.e. ± 0.07). Swards grazed by sheep alone contained less white clover, but regrew quicker and produced more herbage than other treatments. Close topping or grazing by sheep following dairy cattle grazing decreased sward rejection by cattle. These treatments maintained more of the pasture in better condition for subsequent cattle grazing, resulting in greater net herbage production than where no post-cattle grazing treatment was used.     

A plate meter inadequately estimated herbage mass in a semi-arid grassland

A resting plate meter was tested in the laboratory and on a field site to determine its effectiveness for estimating peak herbage mass on semi-arid grasslands. In laboratory tests, data from the plate meter closely predicted the herbage mass of four perennial bunch grasses (thirteen of sixteen tests with r ² ≥ 0·90) but the closeness of the prediction varied with the pressure of the plate. Field tests took place on a southern Arizona, USA semi-arid grassland in 2005, 2006, 2007 and 2008. Twenty teams of two or three people sampled the site; five teams in each year measured eight to thirteen plate heights and clipped the vegetation beneath. Consistent with a need for yearly calibration, a significantly different ( P  <   0·05) linear relationship was detected between 3 of the 4 years which was associated with differences in average herbage mass of dry matter (DM) (1525 kg ha⁻¹ for 2005, 2093 kg ha⁻¹ for 2006, 1338 kg ha⁻¹ for 2007 and 1370 kg ha⁻¹ for 2008). Plate height poorly predicted herbage mass within years ( r ²  = 0·21, 0·51, 0·49 and 0·41 respectively) with plate heights explaining half or less of the variability in field herbage mass and having a mean prediction error of 466 kg DM ha⁻¹. The plate meter technique had limited potential for estimating peak herbage mass in semi-arid grasslands.     

A comparison of quadrat, capacitance meter, HFRO sward stick, and rising plate for estimating herbage mass in a smooth-stalked, meadowgrass-dominant white clover sward

A single-probe capacitance meter (Pasture Probe), the Hill Farm Research Organization (HFRO) sward stick, a rising-plate meter, and cut quadrats were used to estimate herbage mass of swards that were rotationally grazed by cattle, cattle followed by topping, cattle followed by sheep, or sheep alone during 1989 and 1990 grazing seasons. Mean target pre- and post-grazing herbage masses were 2200 and 1100 kg dry matter (DM) ha⁻¹ respectively. Linear regressions, correlations and scatterplots were calculated relating meter and sward stick readings to herbage mass on an ash-free organic matter basis measured by cutting quadrats of herbage at ground level.
Mean coefficients of variation for quadrat, capacitance meter, sward stick and rising plate were 28·8, 15·5, 27·2 and 27·9% respectively for pre-grazing herbage mass measurements, and 20·2, 10·1, 21·4, and 18·4% respectively for post-grazing measurements. These coefficients indicate that the capacitance meter varied less in estimating pre- and postgrazing herbage mass than the other three methods.
Correlation coefficients relating cut quadrats to capacitance meter, sward stick and rising plate readings were 0·65, 0·70 and 0·72 for pre-grazing, and 0·36, 0·31 and 0·05 for post-grazing herbage mass measurements respectively.
The non-destructive methods provided quick herbage mass estimates at a level of precision adequate for making day-to-day grazing management decisions on farms.     

Factors affecting the stolon growth of white clover in ryegrass/clover patches

The effects of spatial location of white clover ( Trifolium repens L.) within a perennial ryegrass ( Lolium perenne L.)/white clover pasture on stolon and petiole extension were investigated in two experiments, where patch size containing white clover (0·5 m, 1·5 m and 4 m diameter), location within the patch (inside and edge) and cutting height (4 cm and 8 cm) were varied. Stolon extension rate was greater on the edge of a patch (12·1 mm week⁻¹) than inside the patch (7·2 mm week⁻¹). Patch size affected both stolon and petiole extension rate, which were both greater in small and medium-sized than in large patches. It is suggested that the fastest spread of white clover in patchy sward environments should occur from small patches, which could double in diameter during a growing season. Manipulating the heights of vegetation within and outside large patches affected light quality (red-far red; R/FR) at ground level, which was greater under shorter than taller swards and greater under the canopy of the grass matrix than the grass/white clover patch. However, the height differences between adjacent vegetation had little effect on stolon or petiole growth. In May only, stolon extension at the patch boundary was greatest when both patches and the grass matrix had a height of 8 cm.     

Assessment of contrasting perennial ryegrasses, with and without white clover, under continuous sheep stocking in the uplands. 3. Herbage production, quality and intake

Herbage characteristics were monitored over 3 years (1985–1987) in three perennial ryegrass ( Lolium perenne L.) varieties as grass/only (200 kg N ha⁻¹) and grass/clover ( Trifolium repens L.) (75 kg N ha⁻¹) swards when continuously stocked with sheep. Although mean total annual herbage production was similar from the grass varieties, growth of Aurora, a very early-flowering variety, was higher than that of Aberystwyth S23 and Meltra (tetraploid) late-flowering types in spring. Herbage production from grass-only swards was 15% higher than from grass/clover swards.
In vitro organic matter digestibility (OMD) of Aurora and Meltra was 34 g kg ⁻¹ and 26 g kg ⁻¹ higher ( P < 0·01) than that of S23. Averaged over varieties, the OMD of grass/clover was 25 g kg ⁻¹ higher ( P < 0·05) than grass-only during the post-weaning period. Individual lamb digestible organic matter intake (DOMI) was 22% higher ( P < 0·05) on grass/clover than on grass-only during this period. Water-soluble carbohydrates content of Aurora was 42% higher ( P < 0·001) than that of both Meltra and S23.
Herbage characteristics were related to lamb performance reported previously. Lamb production was positively correlated with DOMI. However, the magnitude of differences in lamb performance between treatments was much greater than the extent of differences detected in herbage characteristics.     

The effect of offering a brewers grains/treated straw supplement to grazing dairy cows

In a two-year study, set stocked dairy cows were offered a mixture of brewers grains and sodium hydroxide-treated straw (1:1 on a dry matter basis)for 1 h daily. The experiments lasted for 20 and 19 weeks, mean supplementary (buffer) feed dry matter intakes were 3·0 and 3·4 kg d^-1 and mean sward heights (rising plate meter) were 4·9 and 4·7 cm in 1988 and 1989 respectively. The control group received no supplementation. The buffer feed significantly increased milk yield (26·3 vs 24·6 kg d^-1 and 28·6 vs 26·3kg d^-1 in 1988 and 1989 respectively), increased liveweight gain (0·27 vs 0·13 kg d^-1 and 0·27 vs 0·17 kg d^-1) and increased effective stocking rate (5·17 vs 4·69 cows ha^-1and 5·10 vs 4·53 cows ha^-1). There were no significant effects on milk composition. The greatest responses in milk production were found in the late grazing season.     

Herbage productivity of a range of grass species in association with white clover

Red fescue ( Festuca rubra L.) cvs Reptans, Dawson and Ruby, perennial ryegrass ( Lolium perenne L.) cvs Perma and Sprinter, crested dogstail ( Cynosurus cristatus L.) cv. Southlands, smooth meadow-grass ( Poa pratensis L.) cvs Parade and Arena, Yorkshire fog ( Holcus lanatus L.) commercial types I and II, creeping bent ( Agrostis stolonifera L.) cv. Penncross and rough meadow-grass ( Poa trivialis L.) cvs Omega Øtofte, Dasas and Sabre were each sown with white clover ( Trifolium repens L.) cv. Donna and the productivity of the fourteen mixtures assessed under simulated grazing (six harvests per year) for 3 years. A single fertilizer-N rate (50 kg ha⁻¹) in early spring was applied to all treatments each year.
Over the 3 years, mean annual dry matter (DM) production of the swards ranged from 5·25 t ha⁻¹ (creeping bent) to 8·71 t ha⁻¹ (red fescue cv. Reptans), with the species ranking order of: red fescue > perennial ryegrass > crested dogstail > smooth meadow-grass > Yorkshire fog > creeping bent. Rough meadow-grass only persisted over the first harvest year. Mean annual organic matter digestibility (OMD) values of the swards ranged from 0·712 (creeping bent) to 0·782 (ryegrass cv. Sprinter). Mixtures with the lowest proportion of grass (crested dogstail and smooth meadow-grass) generally gave the highest proportions of white clover. The creeping bent mixture had exceptionally low production of both components.
Red fescue, crested dogstail and smooth meadow-grass showed compatibility with clover and, therefore, have potential for use in grass/clover swards. Yorkshire fog had lower compatability with clover. Some breeding effort to improve selected characteristics of promising species, e.g. improvement of low OMD in red fescue, seems warranted.     

Effects of spring fertilizer nitrogen and sward height on production from perennial ryegrass/white clover swards grazed by beef cattle

An experiment was carried out to compare the effects of two compressed sward height treatments, each at two fertilizer nitrogen (N) treatments (0 and 50 kg ha⁻¹ in spring), on the date of turnout and liveweight gain of steers grazing a perennial ryegrass/white clover ( Lolium perenne/Trifolium repens ) sward sited on clay loam in south-west England. The sward height treatments were 6 cm all season, and 4 cm in the spring rising to 6 cm in June; these were maintained using continuously variable stocking with Hereford × Friesian steers. Cattle were turned out on average 11·5 days earlier on the 4-cm sward height compared with the 6-cm sward height treatment. Liveweight gain early in the season was lower on the 4-cm swards than on the 6-cm swards. Liveweight gain ha⁻¹ over the whole season was similar for the two sward height treatments. Fertilizer N did not affect turnout date or liveweight gain.     

The voluntary intake and in vivo digestibility of herbage harvested from indigenous hill plant communities

Voluntary intake and in vivo digestibility of herbage harvested from five indigenous communities– Agrostis-Festuca, Nardus and Molinia grasslands and Eriophorum vaginatum- and Trichophorum caespitosum -dominated varieties of blanket bog–and also from sown Lolium perenne and Trifolium repens swards were measured over two years. First growths of each sward were harvested at three stages of maturity and stored at — 20°C before feeding. Voluntary intake was measured while feeding ad libitum to 15% excess of the intake on the previous day and digestibility when feeding at 80% of voluntary intake.
There was a common positive relationship (r²= 0·86***) between intake (adjusted for year of feeding, range 72·8–16·7 g organic matter (kg LW)^0·75 d^-1) and organic matter digestibility (range 0·796–0·371), covering Lolium perenne and most of the indigenous communities. However, there were significant differences between the regressions of intake on the concentration of neutral detergent fibre for ryegrass and the indigenous communities, and also between the regressions for individual communities.
The results demonstrated the general value of organic matter digestibility as a predictor of intake potential, and the high digestibility and intake potential of immature vegetation from the grass and grass heath communities.     

Supplementary forage for grazing sheep. 1. Effects on lactating ewes and lambs

In two experiments, ewes in weeks 1–20 of lactation received either pasture only or pasture plus ad libitum conserved forage. Herbage height was maintained at 3·3–3·4 cm on both treatments by manipulation of stocking rate in the 24 plots (twelve per treatment) used for the experiment with additional ewes and lambs. In experiment 1, silage, which was of lower metabolizable energy (ME) and crude protein (CP) contents than the grazed herbage, was consumed at an average of 375 g dry matter (DM) ewe⁻¹ d⁻¹. In experiment 2 a strawmix, containing (g kg ⁻¹ freshweight) 480 g barley straw, 300 g concentrate and 220 g molasses, again with a lower ME and CP content than the grazed herbage, was consumed at an average of 165 g DM ewe ⁻¹ d ⁻¹. In both experiments ewes not receiving forage lost weight whereas those with forage gained weight and tended to have increased body condition scores. Offering silage or strawmix did not affect lamb liveweight gain or stocking rate.     

Morphological development and forage quality changes in crimson clover (Trifolium incarnatum L.)

The development of efficient forage management and utilization is facilitated by a knowledge of the changes in nutritive value and patterns of dry matter (DM) distribution within plants. The objectives of this experiment were to determine the patterns of DM accumulation and the change in nutritive value during maturation of crimson clover ( Trifolium incarnatum L.), a cool-season annual forage legume adapted to mild winter areas and acid soils. Crimson clover was established on a Dystric Eurocrept soil with pH 5·4 in Galicia, north-western Spain, during three consecutive years (1990–92). Shoot samples were taken at approximately biweekly intervals in the spring and summer after autumn sowing, and were separated into stems, leaves and seed pods. DM accumulation, crude protein (CP) and acid-detergent fibre (ADF) concentrations and in vitro organic matter digestibility (IVOMD) were determined for each plant part at each stage. Average annual DM production at initial flowering was 4·2 t ha^–1, whereas the CP and ADF concentrations and IVOMD values were 134 g kg^–1 DM, 302 g kg^–1 DM and 0·720 respectively. The nutritive value declined with age, coinciding with a reduction in leaf:stem ratio (from 0·53 at the vegetative stage to 0·13 at the ripe seed stage). This decline was more rapid than that reported for other legumes, possibly because of the low leaf:stem ratio of crimson clover. However, as it is used in the Atlantic coastal areas of north-western Spain, where it is harvested around initial flowering, it can provide forage of good quality.     

A study of the nutritive value of white clover (Trifolium repens L.) in relation to different stages of phenological maturity in the primary growth phase in spring

This paper reports the effects of onset of phenological maturity on the nutritive value of white clover ( Trifolium repens L.). The study comprised (i) examination of an extensive data set on nutritive value and (ii) investigation of the constituents of nutritive value, in vivo feeding value, protein degradability and metabolizable protein content of white clover harvested at three stages of maturity (early-flowering, full-flowering, ripe seed stages) during the primary growth phase in spring in Australia. The data set on nutritive value showed a consistent pattern of high nutritive value during cool season months, progressive decline through spring and uniformly lower nutritive value over summer. Results from laboratory determinations, in sacco degradability studies and a digestibility trial on white clover harvested at early-flowering, full-flowering and ripe seed stages were consistent with results from the data set on nutritive value. Onset of maturity during the primary growth phase in spring was accompanied by large changes in nutritive value: neutral-detergent fibre (NDF) increased from 184 to 301 g kg⁻¹ dry matter (DM), nitrogen (N) declined from 36 to 20 g kg⁻¹ DM, in vitro digestibility declined from 0·74 to 0·65 and metabolizable protein content declined from 144 to 67 g kg⁻¹ DM from early-flowering to ripe seed stage. These nutritive value changes were accompanied by a decline of in vivo digestibility at the rate of 0·0032 d⁻¹ and an 0·2 reduction in voluntary intake.     

The impact of grazing severity on perennial ryegrass/white clover swards stocked continuously with beef cattle

An experiment was carried out to examine the changes in perennial ryegrass ( Lolium perenne L.) and white clover ( Trifolium repens L.) populations in mixed swards, under different grazing severities over three successive grazing seasons. In year 1, three paddocks were erected on a sward with a low initial content of clover (block 1). Sward heights were measured using a rising-plate meter, and were maintained at overall mean heights of 3·0, 5·5 or 7·0 cm by variable stocking with 8-month-old steers. In year 2, a further three paddocks were erected on an adjacent area with a high initial content of clover (block 2), and were maintained at the same three heights by similar management. Botanical analyses were carded out on samples collected at four times during the season. Maintaining swards at 5·5 or 7·0 cm led to a large proportion of the area being infrequently grazed. Block I paddocks had higher initial tiller densities, which increased as sward height was reduced, while block 2 paddocks, with their lower initial tiller density, showed little effect of sward height on tiller density. Initially, clover stolon growing-point densities and stolon masses increased more rapidly in the taller swards. Later, however, large losses in the clover populations occurred on all paddocks during long wet winters and there was a general reversal in these trends for stolon growing-point densities and stolon masses, 3·0>5·5>7·0. By year 3, swards with differing     

Plant interference and chemical toxins in upland pastures

Supernatants derived from desiccated plant material gathered from Agrostis/Festuca vegetation had an inhibitory effect on the germination and early development of seedlings of Trifolium repens cv. S184. Two compounds, o -hydroxyphenylpropionic acid and benzoic acid, were identified chromatographically and their structures confirmed by mass spectrometry. Commercial preparations of the two compounds were effective inhibitors at a concentration of 10⁻² and 10⁻³ m respectively when T. repens was used as the phytometer species. Benzoic acid lost its inhibitory effect when the pH of the two solutions was adjusted to pH 5·5. The roots of seedlings of white clover were severely distorted by o -hydroxyphenylpropionic add at 10⁻³ m . It is likely that phenolic acids, produced from the surface trash created by slot seeding procedures, interfere with the establishment of white clover in upland pastures.     

Response of permanent and reseeded grassland to fertilizer nitrogen. 1. Herbage production and herbage quality

The productivity of permanent swards of mixed species composition was compared with that of sown Lolium perenne , cv. Melle at five fertilizer-N rates (0, 150, 300, 450 and 900 kg N ha⁻¹) and with L. perenne/Trifolium repens , cv. Grasslands Huia at 0 kg N ha⁻¹. The investigation was conducted under two cutting frequencies at sixteen sites in England and Wales, representing a range of grassland environments.
Annual total herbage dry matter (DM) production from both permanent and reseeded swards increased with successive increments of fertilizer-N up to 450 kg N ha⁻¹. Herbage DM production from reseeded swards in the first year after sowing was signficantly higher than from the permanent swards, at all fertilizer-N treatments. In subsequent years the production advantage of the L. perenne reseeds was maintained only at the higher N rates, though sown L. perenne/T. repens was the most productive sward type at 0 N. Average differences in modified acid-detergent fibre suggested small advantages in herbage quality to the reseeded swards.
It is concluded that, while reseeded swards are more productive in the year after sowing, many permanent swards are capable of high levels of production and that reseeding to a L. perenne sward cannot always be justified, particularly for grassland receiving low or moderate inputs of fertilizer-N.     

Annual cool-season legumes for forage production in mild winter areas

The forage potential of several annual winter legumes – crimson clover ( Trifolium incarnatum L.), faba beans ( Vicia faba L.), forage peas ( Pisum sativum L.), serradella ( Ornithopus sativus Brot.) and vetches ( Vicia sativa L. and Vicia villosa Roth.) – was evaluated over two growing seasons in the wet and mild winter areas of Galicia (north-west Spain) with a moderately acid soil, at the two normal harvesting dates in the region, April and May.
The results indicate that, in April, serradella and crimson clover (in the warmer location), can produce dry-matter (DM) yields of 4–5 t ha⁻¹, which is similar to other winter forages in the same area, with an average growing season of 212–237 d, corresponding to about 339–420 accumulated degree–days above 10°C (GDD₁₀). In May, mainly in the warmer location, fava beans, peas and vetches yielded on average 4·7–8·5 t ha⁻¹ DM, with an average growing season of 166–206 d but with only 233–278 GDD₁₀ units. Faba bean, with an average of 8·5 t ha⁻¹ DM in the May harvest, was the highest producing forage legume. The results also suggest that some cool-season legumes could fit into double-cropping systems based on summer crops, such as maize, because of their forage yields and nutritive value. However, in order to produce consistent DM yields, the date of sowing is crucial; they must either be sown much earlier than the dates in this study or the harvests must be delayed until the end of April.     

Yield and nitrogen utilization by Lolium perenne and Trifolium repens on a limed stagnohumic-gley soil in a pot experiment

In a pot experiment the pH of a hill peaty soil was adjusted to cover the range 3.8 to 6.8 with Ca(OH)₂. Yield and N content of the harvested plant material was greatest at pH 6.8 in pots sown with only Lolium perenne , and greatest at soil pH 6.1 in pots sown with Trifolium repens and L. perenne. T. repens established only at soil pH between 4·3 and 6·1. A small reduction in sward yield and a large and highly significant decrease in total N recovered by the harvested material, which occurred with increase in soil pH from 3·8 to 4·3, was attributed to a combination of increased N immobilization in the soil microbial biomass and to the poor performance of both sown and self-propagating indigenous species around pH 4·3.     

Yield and stability of yield of single- and multi-clover grass-clover swards in two contrasting temperate environments

Diversity of clovers in grass-clover swards may contribute to greater herbage yields and stability of yield. This possible effect was evaluated in an experiment carried out over three harvest years at two contrasting sites, differing in precipitation and soil composition, using mixed swards containing either one, two or three clover species sown together with timothy ( Phleum pratense L.) and meadow fescue ( Festuca pratensis L.). The clover species were red clover ( Trifolium pratense L.), white clover ( Trifolium repens L.) and alsike clover ( Trifolium hybridum L.) sown in various proportions in a total of ten treatments. All swards were fertilized with nitrogen with amounts that increased from year to year, and three harvests were taken in three consecutive years. There was a significant interaction between site and species mixture on total dry matter (DM) yields (range 27–32 tonnes ha⁻¹) and DM yields of clovers (range 5–15 tonnes ha⁻¹); red clover as a single species or in a mixture was superior at the dry site while multi-clover species mixtures were superior at the wet site. Alsike clover was the least productive species of clover. Stability of yield of clovers was generally higher by including white and red clover in the seed mixture but total DM yield was not.