Stand structure of an uneven-aged fir–beech forest with an irregular diameter structure: modeling the development of the Belevine forest, Croatia

This study focuses on the problem of irregular diameter structure in a silver fir–beech selection (plenter) forest with a “surplus” of large diameter trees and a lack of natural regeneration and small diameter trees. We sampled 274 plots (900 m² each) in the Belevine research site (266.24 ha) in the mountain region Gorski Kotar (Croatia), where diameter (dbh) distribution, diameter increment, and natural regeneration were analyzed in detail. A low density of natural regeneration, weak annual recruitment of small (10 cm dbh) diameter trees (only five trees per hectare), delayed diameter growth of trees, and a low annual rate of trees reaching the next dbh class were attributed to the current irregular dbh structure. The stand development prediction for the next 50 years is based on a simulation model, which considers the current diameter structure, increment, recruitment, and future cutting regime. Intensive cutting in the first of five 10-year cutting cycles (intensity higher of 25%) is needed to initiate natural regeneration and to accelerate growth of young silver fir trees. In the next 50 years, the irregular diameter structure will be gradually improved.     

Floristic diversity and carbon stocks in the periphery of Deng–Deng National Park,Eastern Cameroon

Carbon is continuously being removed from the atmosphere by photosynthesis and stored in carbon pools(live,dead,and soil carbon)of forest ecosystems.However,carbon stock in dead wood and of trees with diameters at breast height(dbh)between 5 and 10 cm is often not considered in many studies carried out in the Congo Basin Forest.The relationship between tree diversity,life-forms and carbon stocks has received little attention.This study was carried out on the outskirts of Deng Deng National Park(DDNP)to determine tree diversity(dominant families,species richness and Shannon index),assess carbon stocks in the five carbon compartments(living tree,understory,fine roots,dead wood and litter)as well as to analyze the relationship between(1)carbon stocks and tree diversity;and,(2)between carbon stock and life-forms.The Shannon index of trees≥10 cm dbh ranged from 2.6 in riparian forest to 4.3 in secondary forest;and for the tree between 5 and 10 cm,it ranged to 1.56 in riparian forest to 3.68 in the secondary forest.The study site housed 16 species,7 genera and 3 families which are only found in trees of dbh between 5 and 10 cm.The average total carbon stock of the five compartments varied from 200.1 t ha-1 in forest residues to 439.1 t ha-1 in secondary forest.Dead wood carbon stock varied from 1.2 t ha-1 in riparian forests to 12.51t ha-1 in agroforests.The above ground carbon stocks for trees with diameter between 5 and 10 cm varied from 0.7 t ha-1 in young fallow fields to 5.02 t ha-1 in old secondary forests.This study reveals a low but positive correlation between species richness and total carbon stocks,as well as a significant positive relationship between life-forms and total carbon stocks.The findings highlight the need for more data concerning carbon content of dead wood,carbon of trees≥5 cm<10 cm dbh and the relationship between carbon stocks and tree diversity from other areas of the Congo Basin for a good understanding of the contribution of tropical forests to climate change mitigation.     

Comparative regeneration status in a natural forest and enrichment plantations of Chittagong （south） forest division, Bangladesh

于2002年4月至11月,对孟加拉Chittagong (南部)林区天然纯林和肥沃地人工林树种更新情况进行了比较研究。共设100个3m×3m的样地(天然林50个,人工林50个)。共记录天然林更新树种64个,苗木密度为24767株/hm2;人工林更新树种40个,苗木密度为18633株/hm2。天然林更新量最大的树种为Castanopsis spp.(2200 株/hm2),其次是Glochidion lanceolarium (2183 株/hm2);人工林更新量最大的树种为Dipterocarpus gracilis (2117株/hm2),其次是Anogeissus acuminata (2000株/hm2)。对天然林而言, Castanopsis spp 的相对密度最高(8.88%), Glochidion lanceolarium 的相对频度最高(7.36%),Syzygium spp. 的相对富有度最高(3.79%),Glochidion lanceolarium的重要值指数最高(18.24%);对人工林而言,Dipterocarpus gracilis的相对密度最高(11.36%),Glochidion lanceolarium的相对频度最高 (9.71%), Dipterocarpus gracilis的相对富有度最高(5.92%), Glochidion lanceolarium的重要值指数最高 (23.32%)。Dipterocarpus gracilis 常见种在天然林和人工林都有更新苗木,但人工林的苗木要高于天然林。天然更新的苗木非常密集,但由于人为的干扰,这些更新的苗木通常达不到径干生长期。表4参35。     

Plant diversity, population structure, and regeneration status in disturbed tropical forests in Assam, northeast India

We investigated the plant population structure and the phy-tosociological and regeneration status in two disturbed tropical forests in Assam Province, the Hojai Reserve Forest and Kumorakata Reserve Fo...     

江西德兴天然次生林林下幼苗更新特征

为了解实生和萌生幼苗个体数量及比例在各类植被中的变化情况,本文以江西德兴常绿阔叶林不同林分为研究对象,对各林型中幼苗的种类组成、数量、多样性与更新方式进行调查研究。结果表明：7种林分类型的乔木更新情况不同,德兴常绿阔叶林更新幼苗共35种,分属17科23属。更新苗数量最低的是拟赤杨＋马尾松林,仅为1 250株/hm^2,更新苗最多的林分类型为米槠＋甜槠林,为6 500株/hm^2;甜槠＋木荷林更新幼苗种类高达25种,居7种林分之首,马尾松＋木荷林下幼苗仅有10种;从演替更新年龄结构来看,马尾松＋木荷林和丝栗栲＋木荷林所在的群落,林下更新苗属于增长型锥体。其它5种林型属于下降型锥体。     

雪灾干扰下林窗对木荷幼苗更新的影响

林窗干扰是维持森林生态系统的重要驱动力之一,对种子萌发、幼苗等自然更新过程、森林物种组成和动态、森林生物多样性的维持具有重要作用。本研究以2008年雪灾干扰后的浙江江郎山木荷林为研究对象,对木荷林窗大小结构、幼苗更新、生长等进行调查研究,结果表明:扩展林窗以50 100 m2的林窗个数最多(占总数的45.45%),各等级林窗中以50 100 m2的林窗占总面积比例最大(占总面积的30.31%)。林窗中木荷幼苗的平均高度和地径较对照林分分别高1.44 cm和0.61 mm,幼树在林窗中的平均高度和地径则比对照林分中分别高45.37 cm和5.00 mm且差异显著;林窗大小对木荷幼苗、幼树的高度和地径生长影响显著,中林窗中幼苗的高度和地径均高于小林窗和大林窗中的幼苗且差异均显著(F=4.893,P=0.007;F=5.203,P=0.004;n=357);幼树的地径在不同大小林窗中差异显著(F=3.569,P=0.037;n=43)。林窗幼苗的更新密度随着林窗面积的增大而增大,在林窗面积达到76 m2时,更新密度达到最大值,而后随着林窗面积的增大下降;中林窗和小林窗中更新苗木以低矮植株(1级、2级)为主,面积100 m2大林窗中,木荷幼苗生长较快。与他人研究的森林天然林窗相比,雪灾干扰后改变了林窗的大小分布结构和面积,50 100 m2的林窗比例较大,一定程度上更利于幼苗更新,具有相对较大的林窗幼苗更新密度;不论林窗大小,林窗内的更新幼苗都比林内多,郁闭度较大的林内或大面积的空地上都不利于更新幼苗的生长。因此,从受灾木荷林窗大小结构、幼苗更新、生长等来看,中林窗是幼苗适宜更新的面积,为木荷灾后恢复与重建提供了科学依据。     

Regeneration ecology and population status of a critically endangered and endemic tree species （Ilex khasiana Purk.） in north-eastern India

Detailed studies on population structure and regeneration ecology of Ilex khasiana were carried out at five representative natural populations i.e. Mylliem (Population - I), Upper Shillong (Population - II), Shillong Peak (Population - III), Laitkor (Population - IV) and Nongpiyur (Population - V) to probe into the responsible affecting factors for its population structure and regeneration ability in these populations. Experimental observation under three controlled light conditions (full sunlight (100%), intermediate sunlight (50%) and low sunlight (30%)) showed that the growth status and survival rates of seedlings under intermediate and low light were better than those under the high light condition. The density-diameter distribution in population of adult trees (≥5cm dbh) exhibited a typical character of a regenerating population. A higher seedling mortality rate in natural populations of I. khasiana was most probably due to increase in light intensity following vegetation destruction and other anthropogenic disturbances that cause opening of forest canopy. The findings of the present study would be of immense value in formulating appropriate conservation measures for the species.     

Regeneration of Picea abies forests on highly productive peatlands—clearcutting or selective cutting?

The study was focused on severe forest regeneration problems that are typical for highly productive peatlands. The aim of the study was to give recommendations for practical forestry on how to renew the forests. Experiments with different forest regeneration methods on highly productive peatlands were set up in nine mature Norway spruce (Picea abies (L.) Karst.) forests in northern, central and southern Sweden. The treatments in the study were natural regeneration in shelterwoods (at densities of 140 and 200 stems ha^‐1), planting of bare‐root spruce seedlings after site preparation (mounding) in the shelterwoods and on clearcuts, and planting without site preparation on clearcuts.

Judging from the extent of windthrow in the denser shelterwoods and the stocking of natural regeneration under remaining shelter trees 4–5 years after final cut, shelterwood regeneration is a promising method. On clearcuts, planting without site preparation resulted in poor seedling survival, large extent of damage to the seedlings and small height increment. Planting in mounds on the clearcuts was much more successful. The best survival of planted seedlings was observed in the shelterwoods. Both the denser and the sparse shelterwoods seemed to give satisfactory protection to the seedlings.

Final recommendations on the most suitable forest renewal method(s) will be made in about five years after the removal of the shelter trees.     

红锥天然林生长规律与生物量的调查研究

华安金山林场红锥天然林调查表明,红锥生长快,３４～３８年生树高可达２１．９～２３．７ｍ、胸径可达２０～３０ｃｍ。全林保存密度５７２株／ｈｍ２,蓄积量３６６ｍ３／ｈｍ２;Ⅱ级立地级保存红锥６００株／ｈｍ２,蓄积量高达４８３ｍ３／ｈｍ２,生物量５００．８７ｔ／ｈｍ２,材积年生长量可达１３．４ｍ３／ｈｍ２;Ⅲ级立地红锥蓄积量３４２ｍ３／ｈｍ２,生物量３５４．６５ｔ／ｈｍ２。林分生产力很高,树高速生期为４～１８年生。胸径速生期为６～１４年生,材积速生期为１０～２７年生,３４～３８年生的红锥尚未达到数量成熟。表明红锥适应性强,不苛求立地,生长潜力大,极有发展前途。     

Relationships between diameter and height of trees in natural tropical forest in Tanzania

The relationship between tree height (h) and tree diameter at breast height (dbh) is an important element describing forest stands. In addition, h often is a required variable in volume and biomass models. Measurements of h are, however, more time consuming compared to those of dbh, and visual obstructions, rounded crown forms, leaning trees and terrain slopes represent additional error sources for h measurements. The aim of this study was therefore to develop h–dbh relationship models for natural tropical forest in Tanzania. Both general forest type specific models and models for tree species groups were developed. A comprehensive data set with 2 623 trees from 410 different tree species collected from a total of 1 191 plots and 38 sites covering the four main forest types of miombo woodland, acacia savanna, montane forest and lowland forests was applied. Tree species groups were constructed by using a k-means clustering procedure based on the h–dbh allometry, and a number of different non-linear model forms were tested. When considering the complexity of natural tropical forests in general and in particular variations of h–dbh relationships due to high species diversity in such forests, the model fit and performance were considered to be appropriate. Results also indicate that tree species group models perform better than forest type models. Despite the fact that the residual errors level associated with the models were relatively high, the models are still considered to be applicable for large parts of Tanzanian forests with an appropriate level of reliability.     

Effects of gaps on regeneration of woody plants: a meta-analysis

Forest gaps, openings in the canopy caused by death of one or more trees, have a profound effect on forest regeneration and drive the forest growth cycle. It is therefore necessary to understand the effects of forest gaps on regeneration for modern forest management. In order to provide a quantitative assessment of the effects of forest gaps on regen-eration of woody plants, we conducted this review of gap effects on woody plant regeneration on the basis of 527 observations from 42 indi-vidual papers, and reported the results of these data in a meta-analysis. Overall, densities of regenerated woody plants were significantly greater （359%） in forest gaps than on the closed-canopy forest floor. The regen-eration density in gaps of plantation forests was significantly greater （P＆lt;0.05） than that of natural forest because the regeneration in gaps of plan-tation forests was improved by both gap effects and experimental meas-ures. Similarly, in comparison to natural gaps, regeneration was better enhanced in artificial gaps. Regeneration density exhibited a significantly positive correlation with gap size, but a negative correlation with gap age because the gap size decreased with increasing gap age. Shade tolerance of woody plants affected regeneration density in gaps and understory. Average regeneration density of shade-tolerant species exhibited a sig-nificantly positive response to gaps but densities remained lower in total than those of intermediate and shade-intolerant species. Gap effects on regeneration decreased in response to increasing temperature and pre-cipitation because of the limiting effects of lower temperature and moisture on woody plant regeneration. In summary, forest gaps enhance woody plant regeneration, and the effects of gaps varied by forest type, gap characteristics, environmental factors and plant traits. The results of this meta-analysis are useful for better understanding the effects and roles of gaps on forest regeneration and forest management.     

Relationship between annual rainfall and tree mortality in a tropical dry forest: Results of a 19-year study at Mudumalai,southern India

Variability in rainfall is known to be a major influence on the dynamics of tropical forests, especially rates and patterns of tree mortality. In tropical dry forests a number of contributing factors to tree mortality, including dry season fire and herbivory by large herbivorous mammals, could be related to rainfall patterns, while loss of water potential in trees during the dry season or a wet season drought could also result in enhanced rates of death. While tree mortality as influenced by severe drought has been examined in tropical wet forests there is insufficient understanding of this process in tropical dry forests. We examined these causal factors in relation to inter-annual differences in rainfall in causing tree mortality within a 50-ha Forest Dynamics Plot located in the tropical dry deciduous forests of Mudumalai, southern India, that has been monitored annually since 1988. Over a 19-year period (1988–2007) mean annual mortality rate of all stems >1 cm dbh was 6.9 ± 4.6% (range = 1.5–17.5%); mortality rates broadly declined from the smaller to the larger size classes with the rates in stems >30 cm dbh being among the lowest recorded in tropical forest globally. Fire was the main agent of mortality in stems 1–5 cm dbh, elephant-herbivory in stems 5–10 cm dbh, and other natural causes in stems >10 cm dbh. Elephant-related mortality did not show any relationship to rainfall. On the other hand, fire-related mortality was significantly negatively correlated to quantity of rainfall during the preceding year. Mortality due to other causes in the larger stem sizes was significantly negatively correlated to rainfall with a 2–3-year lag, suggesting that water deficit from mild or prolonged drought enhanced the risk of death but only with a time lag that was greater than similar lags in tree mortality observed in other forest types. In this respect, tropical dry forests growing in regions of high rainfall variability may have evolved greater resistance to rainfall deficit as compared to tropical moist or temperate forests but are still vulnerable to drought-related mortality.     

Influence of distance to forest edges on natural regeneration of abandoned pastures: a case study in the tropical mountain rain forest of Southern Ecuador

In spite of its high diversity the forests in Southern Ecuador are highly endangered by deforestation. One of the main reasons for the loss of forests is the conversion into pastures. Due to their fast degradation, the pastures are abandoned after several years and form an increasing area of unproductive land. The remoteness from existing forest edges is discussed as one reason for the very slow natural reforestation of these areas. In this study we analyzed the regeneration of a secondary forest after approx. 38 years of succession in relation to the distance from the surrounding forest. We revealed that regeneration was rather slow. Especially larger trees with dbh > 10 cm were very scarce. Only Dioicodendron dioicum, Graffenrieda emarginata and Clusia sp. achieved larger diameters. The basal area of the secondary forest is still far beyond the original level in the primary forest. The number of species on plot level and the Shannon index were significantly lower in the secondary forest compared to the primary forest. The total number of species decreased from 47 to 31 with increasing distance from the forest edge and the similarity of species composition to the upper story declined to a level of 56.4 (Sörensen). Alzatea verticillata, Macrocarpea revoluta and Palicourea andaluciana had significantly higher abundances in the succession stages than in the natural forest. The most abundant species in all regeneration plots, G. emarginata and Purdiea nutans, seem to be generalists as they did not show preference either to natural forest or successional stages.     

The sustainability of selection cutting in a late successional broadleaved community forest in Bhutan

This study assessed the sustainability of selection cutting in a community forest (CF) in Bhutan. The harvesting approach differed from cable crane logging operations in an adjacent commercially managed forest by creating much smaller canopy openings. This had many implications for natural regeneration of preferred species. The study was conducted in a late successional broadleaved CF containing 32 genera of tree species dominated by Quercus and Castanopsis and managed for timber, firewood, non-wood forest products and forest grazing. The study was based on a comparison of two forest inventories to assess forest structure and regeneration, a study of stumps to quantify harvesting intensities, and a household survey to quantify livestock holdings and grazing patterns. The study examined different intensities of selection cutting in three blocks of the CF and found that higher intensities of selection cutting did not have a negative impact on: (a) natural regeneration of seedlings and saplings of preferred timber species; (b) the diameter distribution of all species and use categories except for Quercus; (c) the diversity of tree genera; and (d) the percentage of remaining trees with favorable bole shape and form. These results contrasted with findings in the adjacent commercially managed forest. Community management of broadleaved forests with selection cutting appeared to be sustainable and avoid some of the unresolved silvicultural problems associated with commercially managed forests in Bhutan. However more research is needed on the sustainable management of Quercus spp.     

Optimizing dosage and preventing leaching of imidacloprid for management of hemlock woolly adelgid in forests

Hemlock woolly adelgid (Adelges tsugae Annand) threatens native hemlock (Tsuga canadensis and T. caroliniana) in the eastern United States. Imidacloprid used as a soil-applied systemic insecticide is effective against A. tsugae in natural forests and in landscapes. Non-target impacts are a special concern because hemlock is ecologically important; often growing next to streams which contain aquatic species that are sensitive to imidacloprid, some of which are protected or endangered fauna. Environmental risk can be mitigated by determining the minimum effective dosage in forests. Adelgid populations responded to imidacloprid dosage with approximately a linear relationship between the percent population reduction (probability scale) vs. log of dosage; 50% reduction in populations could be achieved with 0.15 g per 2.5 cm of trunk diameter at breast height (dbh), or 10% of the maximum labeled dosage. However, effectiveness was found to vary based on dbh; the dosage predicted to give ∼90% reductions in adelgid populations is given by the equation log(dosage) = 0.0153 × dbh − 1.074, where the dosage is g imidacloprid per 2.5 cm of trunk dbh, and dbh is measured as cm. For trees less than 82 cm dbh, these dosages are less than the typical 1.5 g imidacloprid per 2.5 cm dbh currently used. Formulation into tablets rather than as a wettable powder reduced the maximum imidacloprid concentration in soil solution at 5 cm depth from 80 to 1.6 ppm in columns constructed with the organic soil found beneath forest hemlocks. A combination of optimum dosing of trees and adoption of the tablet formulation when treating trees in sensitive habitats should minimize the risk of contaminating aquatic resources with imidacloprid.     

Effects of management, environment and landscape conditions on establishment of hardwood seedlings and saplings in central Japanese coniferous plantations

There is an increasing need to restore natural hardwood forests in landscapes dominated by monocultural conifer plantations. A convenient restoration approach is to exploit natural regeneration processes. Natural regeneration, however, is affected by diverse interacting factors, for which better understanding is required, in order to optimize restoration programs. To identify optimal management practices for improving natural regeneration of hardwood trees in coniferous plantations, we examined the effects of multiple factors on the abundance of seedlings, small saplings and large saplings (height <0.3, 0.3-1.3 and ?1.3 m, respectively) of hardwood tree and shrub species in both line thinned (LT) and unthinned (UT) plantations of sugi (Cryptomeria japonica) and hardwood forests (HF) in central Japan. The effects of management practices (number of the times of weeding and cleaning, thinning method, years after thinning and forest age), environment (slope position, slope angle and canopy openness), and landscape conditions (distance from nearest hardwood forest, altitude and landuse before planting) on the number of hardwood individuals were examined by using the data obtained from the LT plantations. We also compared hardwood density between LT and UT plantations to examine the effect of line thinning. Finally, we examined species composition of LT plantations and HF to identify hardwood forest components in the thinned plantations. The effects on hardwood regeneration of environmental conditions, landscape factors and management practices applied in the plantations varied, depending on the size class and life form of the regenerating species. The abundance of large saplings of tall tree species was affected by several management factors, especially number of the times of weeding. Landscape conditions (distance from the nearest hardwood forest and altitude) affected the abundance of small saplings and seedlings of tall tree species, but not the other classes. Seedlings and small saplings of many tall tree species that contribute to hardwood forest canopies were less abundant in the LT plantations. The results show that numerous factors affect the establishment and abundance of naturally regenerating hardwood tree species, and suggest that successful establishment during early plantation stages can have long-lasting effects on natural regeneration of tall tree species.     

Tree Life History Patterns and Forest Dynamics

Abstract

Forest trees are usually classified into broad ecological groups (e.g., pioneers vs. mature-phase species) based on the importance of natural disturbance for their regeneration. These classifications only tie into account a small component of the plant's life-cycle, usually seeds and seedlings, and therefore may over-simplify the role of natural disturbance in the evolution of life-history traits. We present a conceptual model for the study of plant demography within heterogeneous environments, and illustrate its use by reviewing how natural disturbance influences the demography of several lowland tropical tree species at Los Tuxtlas, Mexico. Our model assumes that habitat patches such as treefall gaps in tropical forests can affect a plant at all stages of its life-cycle, and reveals many potential life history patterns in relation to forest dynamics. We propose that only detailed demographic studies of whole life histories and genetics of tree populations, as well as forest disturbance regimes, will allow us to elucidate convergent life histories and the existence of discrete ecological groups among tropical forest trees.     

Long-term growth response of black spruce advance regeneration (layers), natural seedlings and planted seedlings to scarification: 25th year update

Careful logging around advance growth and tree planting following site preparation or not (fill planting) have been widely used to regenerate black spruce (Picea mariana) stands in the boreal forest of Canada. An experiment was established in 1990 to compare these regeneration modes and natural seeding over a 25-year period following two types of scarification (cone- or disk-trenching) that were applied at two intensities (single- or double-pass). Without scarification, the three types of regeneration had slow height growth during the first 10 years (3.4–5.8?cm/year). Thereafter, planted seedlings (6.1–9.8?cm/year) and pre-established layers (5.8–8.5?cm/year) had 2–3 times higher growth than natural seedlings (2.6–3.1?cm/year). Scarification improved height growth of the three types of regeneration, but planted seedlings responded much earlier, and more strongly over the 25 years (+9.4?cm/year) than layers (+2.8?cm/year) and natural seedlings (+2.7?cm/year). A shift in the proportion of layers and natural seedlings was observed following treatment. Long-term monitoring revealed that double-pass scarification was beneficial to layer growth from years 15 to 20, while no positive effect was observed on natural or planted seedlings. Our results suggest that decades may be required for natural regeneration to respond to scarification in cold soils of the boreal forest.     

呼伦贝尔沙地樟子松天然更新苗影响因素的研究

通过对呼伦贝尔沙地樟子松天然林天然更新苗特征及其影响因素的研究,结果表明:母树密度不同,天然更新苗数量和年龄结构存在明显差异;樟子松天然更新苗年龄结构在时间上具有明显的不连续性,樟子松天然更新苗密度因林分类型的差异明显不同。在疏林地(树龄30 a),平均更新苗密度每2 500 m2为13株,在火烧迹地平均更新苗密度每600 m2为28.5株,樟子松单株样地平均更新苗密度每600 m2为118.8株。不同年龄的天然更新苗保存密度和年平均降水量有显著或极显著关系,且随着林分郁闭度的增加,研究区天然更新苗密度呈现降低的趋势。