THE EFFECT OF NITROGEN AND STOCKING RATE ON THE OUTPUT OF PASTURE GRAZED BY BEEF CATTLE

An experiment was conducted in two successire years to measure the effect of two levels of fertilizer N, 50 and 300 kg/ha (45 and 270 Ib/ac) on the productivity of pastures grazed by young beef cattle. Two stocking rates were imposed at the lower N level and 4 at the higher level. The responses per kg fertilizer N were approximately 1 kg liveweight gain, 20–24 Meal ME and 8–9 kg DM. Maximal yields of about 1000 kg gain/ha (890 Ib/ac) and 19,000 Meal ME/ha (7700 Mcal ME/acre) were recorded. Animal performance was similar on the low and the high N pastures. There was evidence that the chemical quality of pasture was lower on the low N pasture in the first year, but there was no difference in the second year. The numbers of dung pats per ha and the refusal of herbage due to fouling were reduced by Increasing the stocking rate.     

THE EFFECT OF HIGH NITROGEN LEVELS AND STOCKING RATES ON MILK OUTPUT FROM PASTURE

An experiment using 40 lactating dairy cows was conducted for 3 successive years to examine the effects of fertilizer N applied at 400 and 700 kg/ha (358 and 627 lb/ac) at stocking rates of 4.94 and 7.41 cows/ba (2 and 3 cows/ac). The animals were grazed in groups of 10 per treatment, using a fixed rotational grazing system, for 22 weeks in each year. Milk ontput/cow and/ha was significantly affected by stocking rate, the mean milk outputs being 2499 and 2218 kg/cow (5498 and 4880 lb/ cow) and 12313 and 16396 kg/ha (11032 and 14691 lb/ac) at the lower and higher stocking rates, respectively. Milk yield/cow at the lower stocking rate was not affected by N level. At tbe higher stocking rate, increasing the amount of N resulted in a 7.4% increase in milk output/cow and/ha. This response to N declined from 1.7% in the first year of the trial to 2.4% in the third year. The response in milk yield to N appeared to he greatest during the first part of the grazing season. Milk composition, liveweight change, and herbage availahilities and intakes were also recorded.     

THE EFFECT OF NITROGEN, STOCKING RATE AND GRAZING METHOD ON THE OUTPUT OF PASTURE GRAZED BY BEEF CATTLE

An experiment was conducted in 1971 and 1972 to study the effects of two levels of fertilizer N (50 and 504 kg/ha) on the productivity of pastures grazed by young beef cattle. Two stocking rates were imposed at the lower N level and three at the higher. In addition to rotational grazing, set-stocking and an integrated grazing-conservation system were included. The responses/kg N were approximately 1 kg live-weight gain, 19–24 Meal ME and 8–9 kg DM. Maximum yields of 1200–1300 kg live-weight gain/ha and 25,000–27,000 Meal ME/ha were recorded. Productivity of set-stocked pasture was similar to rotational grazing at the high N level, but lower at low N. The integrated system yielded comparable results to high-N rotational grazing. Performances and herbage intakes per animal reflected the stocking rates imposed.     

A FURTHER INVESTIGATION OF THE EFFECT OF NITROGEN AND STOCKING RATE ON THE PRODUCTIVITY OF PASTURE FOR BEEF CATTLE

The liveweight gain was measured of beef cattle on pastures stocked at 5.9 to 11.8 cattle/ha and receiving 50, 182 or 329 kg N/ha in a 12-week season. N levels and stocking rates were so adjusted that similar yields of herbage per animal were expected. Results showed that similar liveweight gains per animal, averaging 1.03 kg/day, occurred at all N levels. Liveweight gain/ha ranged from 547 to 1037 kg in 12 weeks and the corresponding quantities of metabolizable energy utilized were 9890 and 17590 Meal. The results confinn earlier work (4, 7).     

THE USE OF GRAZING CONTROL FOR INTENSIVE FAT-LAMB PRODUCTION

The effects of three stocking rates on fat-lamb production were examined. These stocking rates were imposed upon four grazing systems. The grazing managements had no significant¹ effects upon the rate of liveweight gain or the total liveweight of lamb produced per acre, nor were the ewes' liveweights significantly affected. Reasons are suggested. The lambs' rate of liveweight gain was related negatively and the total liveweight of lamb produced per acre was related positively to stocking rate. The ewes' liveweights were similarly affected, there being the highest losses of liveweight under the high stocking rate. In all cases significant differences existed between the stocking rates and there was a significant linear relationship between them. It is concluded that high stocking rates are essential to obtain the maximum fat-lamb production on a 'per acre' basis.     

EFFECT OF HIGH FERTILIZER NITROGEN AND STOCKING RATES ON LIVEWEIGHT GAIN PER ANIMAL AND PER HECTARE

The effect of increasing annual fertilizer N application rate from 400 to 800 kg/ha (357 to 714 lb N/ac) and stocking rate from 5.0 to 7.5 animals per forage ha (2.02 to 3.04 animals per forage ac) on herbage availability, digestible OM intake and live weight gain was investigated in a grazing experiment repeated in 3 consecutive years. A 21×1 day rotational paddock grazing system was used in which the grazing area was increased in the ratio of 1:1.5:3 on two predetermined occasions. Increasing the fertilizer N application rate increased herbage availability by 12% but failed to increase live weight gain. Increasing stocking rate decreased herbage availability per ha and per animal, and also live weight gain per animal, but increased live weight gain per forage ha by 17.6%.     

A NOTE ON THE ZERO-GRAZING OF BEEF CATTLE

Groups of 8 steers weighing 300–400 kg (660–880 Ib) were rotationally grazed on a ryegrass-dominant pasture for 115 days. The systems compared were zero grazing (Z), field grazing conducted at the same stocking rate (FC), and field grazing conducted at a stocking rate varied with the intention of giving the same liveweight gain per uiimal as zero grazing (FV). Mean daily liveweight gains were: Z, 0.98; FC, 0.78; FV, 0.90 kg/animal (2.2, 1.7 and 20 Ib/animal), and liveweight gains/unit area were in the ratio 100:78:85. Organic-matter intake, measured on four occasions, was, on average, Z, 6.54; FC, 6.18; FV, 687 kg/head daily (14.4, 13.6 and 15.1Ib). From these results it appears that a comparison of zero grazing and field grazing made at the same stocking rate is likely to underestimate the potential of field grazing for beef production from grass.     

THE EFFECTS OF LEVEL OF NITROGEN FERTILIZER ON BEEF PRODUCTION FROM GRAZED PERENNIAL–RYEGRASS/WHITE–CLOVER PASTURES

The treatments in this summer–grazing experiment were designed as a 2 × 3 factorial, there being two levels of nitrogenous fertilizer (nitro–chalk), supplying 0 and 336kg of N/ha per annum (NO and N3, respectively) and three stocking rates. The fertilizer was applied in 7 dressings at approximately three–weekly intervals. The animals used were Hereford x Friesian steers; these were slaughtered at the end of the grazing season. The stocking rates were calculated on the basis of expected herbage production. The experiment was laid out in 6 randomized blocks which were grazed in rotation. Two blocks were cut for conservation before grazing each year; up to the end of June only 4 blocks were used for grazing.
There was little contamination of the swards with unsown species. With the NO swards the clover contents declined each year, but remained fairly high at over 20% in 1965. Clover contents also declined each year in the N3 swards and by 1965 were only 2 to 4%.
The quantities of herbage cut for conservation were higher from the swards receiving N. The live–weight gains of the cattle (per head) were higher at the lower stocking rates, the effect of stocking rate being more noticeable in the July to Oct. period. Live–weight gains per ha increased at the higher stocking rates. In 1963 and 1965, as stocking rates increased carcass weights per animal decreased and carcass quality, as indicated by the commercial grades, was poorer. In 1964, a dry spell from July to Oct. necessitated the removal of cattle from the experiment and the effects of treatment on carcass weight and quality were not apparent.     

THE RELATIONSHIP BETWEEN INTENSITY OF GRAZING AND THE HERBAGE CONSUMPTION AND GROWTH OF CALVES

Experiments were conducted in 1967 and 1968 in which HerefordXFriesian (Experiment 1) and Friesian (Experiment 2) steer calves horn in April were turned out to graze at one week or 3 months of age, respectively, and maintained at three stocking densities in the ratio 1:2:3 animals per unit area. The calves grazed paddocks of S23 perennial ryegrass in rotation, and were moved when the height of grazed stubble at the medium stocking density was reduced to 8 cm. The rate of liveweight gain and herbage intake per head declined as stocking rate increased. When the results of the two experiments were compared, the weight gain of the calves was more closely related to the weight of herbage residues than to the height of the grazed sward. The rate of liveweight gain was depressed when the amount of herhage left after grazing fell helow 2000–2500 kg OM/ha (1800–2250 Ib/ac).     

THE EFFECT OF NITROGEN APPLICATION TO PASTURE ON BEEF PRODUCTION

Five grazing experiments each lasting 2 or 3 years were made between 1955 and 1967, all starting in the first year of ryegrass/cocksfoot/clover or ryegrass/clover leys. A high and a low rate of N, 235 and 45 Ib/ac on average (263 and 51 kg/ha) were compared for beef production. High- and low-N treatments gave mean clover contents for the grazing season of 8 and 24 % on a dry-weight basis, respectively. High N consistently gave a smaller liveweight gain/animal than low N, on average 1±92 and 2±08 Ib/day (0±87 and 0±94 kg/day), respectively. Liveweight gain/ac was 20% greater for high N than for low N, and in terms of net energy the production from high- and low-N, respectively, was 18,500 and 15,000 MJ/ac (45,700 and 37,100 MJ/ha). Data from these experiments, together with published results, were used to calculate a regression of liveweight gain response on N rate and an equation was derived from this to express the output in terms of profit. At 1971 prices profit was maximal at λ0±9/ac (λ2/ha) with 112 Ib N/ac (125 kg N/ha); it was considerably greater at 1973 prices when higher rates of N were justified.     

A PORTABLE-CORRAL TECHNIQUE FOR MEASURING THE EFFECT OF GRAZING INTENSITY ON YIELD, QUALITY AND INTAKE OF HERBAGE

A grazing experiment using four stocking rates of sheep, equivalent to 34, 45, 57 and 68 sheep/ha on an assumed 200-day grazing season, was conducted using a portable grazing corral technique. Plots were subjected to fotir grazing periods between mid May and early September. The effect of stocking rate on herbage yield and quality and the influence of these factors and feed intake on liveweights of the sheep were recorded. The mean daily herbage organic matter available over the trial was 9.6, 5.8, 3.3 and 23% of the total liveweight of the sheep at each of the four stocking rates. The highest grazing pressure led to the production of high-quality herbage, but also led to reduced productivity, low feed intake and liveweight losses. The most lenient grazing pressure failed to provide adequate herbage utilization. Even at the most intensive stocking pressure, only 66% of the herbage available to ground level was utilized in grazings after July. Intake results suggested that sheep of 45 kg liveweight required 1000–1200 g digestible organic matter per day to maintain body weight. Despite the higher in vitro digestibility of herbage on offer at the higher stocking rates, intake was limited through lack of herbage; a high degree of correlation existed between herbage availability and herbage intake over the four stocking rates and at all grazings. It is concluded that the portable corral technique is well suited for grazing studies and the assessment of sward response to varying stocking rates and is particularly useful where facilities for more extensive studies are limited.     

THE EFFECT OF STOCKING RATE ON ANIMAL PRODUCTION FROM CONTINUOUS AND ROTATIONAL GRAZING SYSTEMS

Liveweight and wool production data are presented from a grazing management experiment in which continuous stocking was compared with a four-paddock rotational system at three stocking rates on a sown pasture. Significant differences were recorded in liveweight and wool production between stocking rates. The overall effect of management was non-significant but its interaction with stocking rate and year were both significant. At the intermediate stocking rate (8 wethers/ac, 20 wethers/ha) increases in both liveweight and wool production were recorded for the rotational management system; in this treatment there was also less need for supplementary feeding. Greater wool production was recorded from the continuously grazed treatments at both the high and the low stocking rates but there was little effect on liveweight or the need for supplementary feeding.     

FACTORS LIMITING ANIMAL PRODUCTION FROM GRAZED PASTURE*

Animal production from intensively managed pasture (240 units N/ac or 300 kg N/ha applied annually) should in theory reach 11,000 Ib/ac (12,500 kg/ha) of milk or 1550 lb/ac (1750 kg/ha) of liveweight gain per year. The figure for theoretical milk production is achieved in experiments, although rarely approached in commercial practice; the figure for liveweight gain is never approached, either experimentally or commercially. On commereial farms in Britain, the average stocking rate for grazed lowland pasture in 1971 was about 0·74 cow equivalents/ac (l±82/ha), compared with a theoretical target of 1·9/ac; in 1969 fertilizer N application was about 48 units/ae (60 kg/ha). The main purpose of this paper is to suggest reasons for the differences between practical, experimental and theoretical levels of production. Economic factors may deter farmers from intensifying their grassland management, but the deterrent to high stocking rates may be the fear of running out of grass. In practice, ungrazed herbage is used as a buffer, but other feeds might be used for this purpose. Concentrates, pelleted roughages and even long dried grass and silage are too attractive to grazing beef cattle to be offered to appetite; but silage might be used as a long-term buffer. Fouling of pasture reduces efficiency of harvesting, but attempts to overcome this effect, by conditioning of grazing animals, spreading excreta or by alternation of grazing and cutting, have largely been unsuccessful. High production per unit area cannot be achieved without high production per animal. Legumes have often been found to increase production per animal, and a technique has been developed for growing red-clover and grass in adjoining areas and grazing them together. The clover was grazed satisfactorily by dairy cows (and caused no bloat) but failed to increase milk yield per animal. Recent calorimetric studies of grass have shown that the net energy value of digestible organic matter is variable, and in particular is low for late-season herbage.     

THE EFFECT OF SUPPLEMENTARY BARLEY UPON GROWTH AND EFFICIENCY OF FOOD CONVERSION IN CALVES KEPT AT HIGH GRAZING INTENSITY

Three groups of Hereford × Friesian steer calves, bom in early January 1969, were strip-grazed on eqoal areas of a sward of S321 perennial ryegrass from May to September inclusive, and received kibbled barley at the rate of nil, 3/4% or 1 1/2% of liveweight per day. Adjustments were made to the number of animals per group in order to maintain the same high grazing intensity (defined as 8 cm stubble height in grazed areas) on all treatments. The animals receiving supplementary barley gained weight at a rate 17–19% higher than the controls, but there was no significant difference in growth rate between the high and low levels of supplementation. The effficiency of conversion of barley was only 5–12 kg per animal liveweight gain/100 kg barley OM. The high level of supplementation resulted in an increase in stocking rate of 36 % over that of the control group, and an increase in liveweight gain per unit area of land grazed of 63%. The additional liveweight gain due to supplements, expressed per unit area of land grazed, was relatively constant at 21–24 kg liveweight gain/ 100 kg barley OM consumed.     

A COMPARISON BETWEEN SPRING AND AUTUMN PASTURE FOR BEEF CATTLE AT EQUAL GRAZING PRESSURES

Two comparisons between spring and autumn pasture for beef cattle were made. Animals used in all comparisons were of similar breed and weight and were subjected to the same feeding regime for 6–8 weeks before turn-out. The pastures were grazed on an equal grazing pressure basis between season comparisons. The pastures received similar rates of fertilizer N between seasons and had similar lengths of rest period for regrowth. Intakes of digestible OM were greater per unit of metabolic liveweight in spring than in autumn. Daily liveweight gains in spring were high, being 1·09 and 1·37 kg (2·4 lb and 3·0 lb), but were lower from autumn pasture at 0·98 and 0·71 kg (2·2 lb and 1·6 lb). Weather was implicated as a factor affecting daily gain from autumn pasture. Greater herbage yields in spring supported 42 and 204 more grazing days per ha which together with the greater gains per animal supported 80–120% more liveweight gain/ha.     

THE USE OF LIVEWEIGHT-GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

A 21-acre grazing trial which ran continuously for three years to evaluate grass and grass/legume pasture with and without inorganic fertilizers is described. Techniques for conducting grazing trials where large treatment differences are expected and the interpretation of results from these trials are discussed. Good practical management of stock and pastures was shown to be necessary in these grazing trials and where it was only possible to establish one true replicate some measure of statistical control was achieved by comparing regression coefficients of cumulative animal production.
The results showed that up to 475 1b liveweight gain/acre per annum (532.48 kg/ha) could be obtained from pastures grazed by small East African Zebu steers. The addition of the legumes Stylosanthes gracilis and Centrosema pubescens to grass mixtures gave significant and economic increases in liveweight gain. Grass/legume swards receiving phosphorus and sulphur produced liveweight gains equivalent to the animai production from grass swards receiving 140 1b N/ac per annum (156.94 kg/ha).     

The effects of stocking rate and nitrogen fertilizer on a perennial ryegrass-white clover sward

The effects of stocking rate and N fertilizer on a mixed sward of perennial ryegrass ( Lolium perenne) and white clover ( Trifolium repens ) set-stocked with sheep were examined. Sward production and composition, and sheep production were studied.
Increasing the stocking rate over the range 25–55 yearling sheep ha⁻¹ reduced herbage accumulation by about 40%, whether or not N fertilizer was applied. Increasing the stocking rate increased the density of ryegrass tillers, but reduced the density of clover stolons and the clover content of the swards. Applications of N fertilizer (200 kg N ha⁻¹ a⁻¹) increased herbage accumulation by about 20% but substantially reduced the clover content.
Liveweight gain per animal and per unit area were greater at the lower stocking rates where the clover content and nutritive value of the diet were greatest. Wool growth per unit area was greater at the higher stocking rates. Applications of 1M fertilizer increased liveweight gain at stocking rates above 25 sheep ha⁻¹, but had no effect on wool production at any stocking rate.
The results demonstrate that a stable and productive grass-clover association was maintained under conditions of set-stocking at around 23 yearling sheep ha⁻¹and that at this stocking rate, which appears to be about the biological optimum, there was no advantage in using N fertilizer.     

INTENSIVE SET-STOCKING OF DAIRY COWS

Farm-scale trials with Ayrshire dairy cows were conducted in South-West Scotland over two seasons to compare set-stocking and paddock grazing at similar stocking rates and N fertilizer rates. At a stocking rate of 1±9–2±0 cows/ac throughout the grazing season and a fertilizer N rate of about 400 units/ac, milk production per head from set-stocked cows was equal to that from paddock-grazed cows. No health problems arose as a result of applying fertilizer to the fields which were set-stocked. These trials hare established that set-stocking is a viable system for the intensive management of dairy cows on grassland in the UK, but further work is necessary to determine whether paddock-grazing has a higher potential for milk production than set-stocking.     

THE USE OF LIVEWEIGHT-GAIN TRIALS FOR PASTURE EVALUATION IN THE TROPICS

An experiment is described in which liveweight gains were measured from two grass/legume pastures using small East African Zebu animals and also progeny of small East African Zebu cows crossed with Bos taurus bulls. Significantly higher production was obtained from cross-bred stock, which averaged 706 lb liveweight gain at per annum (791 kg/ha per annum), compared with 509 lb liveweight gain/ac per annum (571 kg/ha per annum) from Zebu stock, indicating that pasture management had advanced beyond the genetic potential of the local Zebu stock. The problems of designing, conducting and interpreting the results of such grazing trials are discussed.     

THE EFFECTS OF LEVEL OF NITROGEN FERTILIZER ON BEEF PRODUCTION FROM GRAZED PERENNIAL RYEGRASS/WHITE–CLOVER PASTURES

The treatments in this summer–grazing experiment were designed as a 3X3 factorial; 3 levels of nitrogenous fertilizer, supplying 0, 336 and 672 kg N/ha per annum, and 3 stocking rates. The fertilizer was applied in 6 to 8 dressings at intervals of 3 to 4 weeks. The animals used were fattening Hereford X Friesian steers. The stocking rates were calculated on the basis of data available on herbage production under N and irrigation treatments and varied according to the level of N input. The experiment was laid out in 6 randomized blocks, which were grazed in rotation. Blocks were removed from the grazing cycle in spring and early summer of each of the 3 years (1962—-4) over which the experiment was conducted and cut for conservation: yields were recorded. Details are given of: grazing management; results of studies of soil–N levels; the quantities of herbage removed for conservation: the quantity, chemical and botanical composition of the herbage available for grazing; and herbage consumption.