Die Zukunft: krankheitsresistente Apfelsorten

Zusammenfassung Integrierte und biologische Anbauverfahren gewinnen zunehmend an Bedeutung. Die Anwendung von Pflanzenschutzmitteln gestaltet sich zunehmend schwieriger bezüglich ihrer Bereitstellung als auch im Hinblick auf zunehmende Umweltsensibilität der Bevölkerung. Im Obstbau ist eine der Alternativen der Anbau resistenter Sorten, die mit weit weniger Pflanzenschutzaufwand produziert werden können. Eines der Ziele der Pillnitzer Obstzüchtung war die Kombination von hoher Fruchtqualität mit guter Ertragsleistung und Resistenz gegenüber Krankheiten und Schädlingen. Resistente früh, mittel und spät reifende Sorten, die Re-Sorten^®, konnten ausgelesen werden. Sie garantieren einen hohen Resistenzgrad unter Feldbedingungen, sie sind aber nicht immun. Dreifachresistente Sorten mit Widerstandsfähigkeit gegenüber Schorf, Mehltau und Feuerbrand sind: Remo, Regia, Rewena und Rebella. Rebella besitzt weiterhin Resistenz gegenüber Rindenbrand, Rote Spinne und abiotische Stressfaktoren. Obwohl in Mitteleuropa seit 1984 verschiedentlich Resistenzdurchbrüche an schorfresistenten Sorten mit M.-floribunda-Resistenz (V_f) beobachtet wurden, blieben die Sorten mit anderer genetischer Basis befallsfrei, so u. a. Reglindis (V_A), Reka (V_r) oder Regia (V_r). Als künftiges Ausgangsmaterial für die Züchtung wurde in der Genbank Obst nach weiteren Resistenzträgern unter den Malusarten und alten Apfelsorten gesucht, das in Zukunft durch Pyramidisierung von Resistenzgenen bei der Züchtung neuer Sorten zu mehr Stabilität unter Feldbedingungen beitragen kann.     

Coincidences between different landscape ecological zones and growth forms of Cembran pine (Pinus cembra L.) in subalpine habitats of the Central Alps

The forgotten depots of the European Nutcracker (Nucifraga c. caryocatactes) often lead to the development of tufts of Pinus cembra. In many cases the other individuals of such tufts are not suppressed by the fittest one, rather there is an intraspecific coexistence up to the senescent stage of the trees. There are fusions of separate trunks, and so frequently the individual history of older trees can only be reconstructed by studying sutures, crown structures or trunk cross sections. Different types of trunk fusions are worked out. By means of transect counting the occurrence of these multiple trunk trees is documented quantitatively in different landscape ecological zones of the Engadin region (the Grisons, Switzerland). The data base is 3024 counted microsites of Pinus cembra individuals arising from seeds, including 5272 living individuals. These multiple trunk trees significantly play an important role in the landscape ecological zones of recent glacier recession and at the alpine timberline. Their growth forms have a higher biomechanical stability.     

Respective roles of recent hedges and forest patch remnants in the maintenance of ground-beetle (Coleoptera: Carabidae) diversity in an agricultural landscape

We compared three kinds of habitats: small remnants of native forests, recent hedges and barley crops, in order to investigate their respective roles in the maintenance of carabid-beetle diversity in a 950-ha area of an intensive agricultural landscape. Carabid faunas in remnants differed weakly from these found in hedges and crops. In particular, small remnants had few typical forest carabid species and a large number of open-area or ubiquitous species. Different approaches in the measurement of and -diversity (classical indices, and additive partitioning of Simpson's index) showed similar results: hedges supported a high -diversity but habitat types were quite similar overall, with weak differences between open and closed or disturbed and undisturbed habitats.A comparison of species dispersal powers in the various habitat types showed that species with a low dispersal power were rare in all habitats. However, wing development measured on two dimorphic species revealed, surprisingly, that brachypterous individuals were mainly present in hedges, which were expected a priori to be more disturbed, than remnants hence less suitable for the establishment of populations with a low dispersal power.These results suggest that small remnants do not behave as 'climax' habitats in this intensive agricultural landscape, probably because of their small size and strong isolation. We discuss the interest of new undisturbed habitats, such as recent hedges, for the maintenance of carabid diversity at both the local and landscape scale.     

Can we manage tropical landscapes? – an answer from the Caribbean perspective

Humans have used Caribbean island landscapes for millennia. Theconversion of wild lands to built-up lands or to agricultural lands in thesetropical countries follows predictable patterns. Conversion of moist forestlifezones and fertile flatlands is faster than conversion of wet and rain forestlife zones and low fertility steep lands. In Puerto Rico, these trends areleading to increased built-up areas, environmental surprises, and increaseddependence on external subsidies. Changes over the past 50 yr alsoinclude a reversal in deforestation and increase in forest patch size in spiteof increasing human population density. Present forests have different speciescomposition than the original ones but are indistinguishable in physiognomy andbasic function. The reversal of deforestation and forest fragmentation trends,if accompanied by an understanding of the forces that cause the reversal, canresult in the development of tools for landscape management. Tropical landscapemanagement requires understanding and application of natural resiliencemechanisms of ecosystems, greater use of ecological engineering approaches toinfrastructure development, enforcement of zoning laws, enlightened economicdevelopment policies, and an understanding and agreement of a conservationvision among all sectors of society. Mixing species in new combinations to formnew ecosystems is a necessary step in the development of future landscapes.Landscapes have always changed over time, but we sense,somewhat uneasily, that the processes now at work are more powerfully focusedand possibly irreversible.Our present assumption is that generally tropical lands aremuch more sensitive and less resilient than those in the temperatezones....the long-term world trend in land uses is a cascadingloss in capacity (i.e., degradation) in every broad category of landuse. J.F. Richards 1990, p 176 and 177.This revised version was published online in May 2005 with corrections to the Cover Date.     

Comparison of available soil water capacity estimated from topography and soil series information

We present a simple and generalized method to predict Available Soil Water Capacity (ASWC-TOP) for a given area using a topographic index, defined as ln(/tan), where is the upslope area draining past a certain point per unit width of slope, and is the local surface slope angle. The estimated results (ASWC-TOP) were then compared with the available soil water capacity calculated from soil series information provided by Soil Conservation Service, U.S. Department of Agriculture (ASWC-SCS). The model implementation was tested with three study cases: the Seeley-Swan valley, Montana, with pixel resolutions of 100 m and 1 km, respectively; and the state of Montana, U.S.A., with a pixel resolution of 1 km. A linear relationship exists between ASWC-SCS and ln(/tan). Standard errors between ASWC-TOP and ASWC-SCS were about 4.4 cm in the Seeley-Swan valley and 5.5 cm in the state. The number of pixels with absolute residuals 4 cm between ASWC-TOP and ASWC-SCS accounted for 68.2, 64.4, and 51.9% for the valley 100 m, valley 1 km, and the state respectively. Some of the mismatches between ASWC-TOP and ASWC-SCS may indicate an improvement using this method compared to existing data because the topographic method reflects the higher spatial variation of the inputs. The increasing availability of digital elevation data, at various resolutions, may provide an alternative to soil series for estimating ASWC. The accuracy of ASWC-TOP depends on estimation of mean and maximum ASWC for a study area.     

Ergebnisse der Leistungsprüfung von Kirschensorten auf wuchsreduzierenden Unterlagen in Ungarn

Zusammenfassung In Ungarn gibt es ein reges Interesse an intensivem Obstanbau. Der Erfolg dieses Anbaus hängt in sehr großem Maße von den Wechselwirkungen zwischen dem Standort, den Sorten-Unterlagen-Kombinationen und dem Anbauverfahren ab.Um die Wirkung wuchsreduzierender Kirschenunterlagen aus ausländischen Züchtungsinstituten unter ungarischen Klimaverhältnissen untersuchen zu können, wurde im Forschungsinstitut für Obst- und Zierpflanzenbau Budapest-Érd ein Vergleichsversuch durchgeführt. Der Versuch wurde mit Cerasus avium C. 2493, Weiroot 13, Weiroot 53, Weiroot 72, Weiroot 154, Weiroot 158, GiSelA 5 und P-HL-A-Unterlagen 1997 in der Versuchsanstalt des Forschungsinstituts für Obst- und Zierpflanzenbau Budapest-Érd aufgepflanzt. Als Kontrolle diente die Sämlingsunterlage Cerasus mahaleb Cema, deren Anteil im ungarischen Süßkirschenanbau bei 70–80% liegt. Die Unterlagen wurden mit drei Süßkirschensorten (Germersdorfi , Linda, Katalin) und mit einer Sauerkirschensorte (Piramis) veredelt.Nach der Wüchsigkeit bis zum Ende des 8. Standjahres konnten die Unterlagen in drei Gruppen eingeteilt werden: starkwüchsige (Cerasus mahaleb Cema, Cerasus avium C. 2493, Weiroot 13), mittelstarkwüchsige (Weiroot 158, Weiroot 154, P-HL-A) und schwachwüchsige Unterlagen (Weiroot 72, Weiroot 53, GiSelA 5). Die stärkste Neigung, Wurzelschosse zu treiben, zeigten Weiroot 13 und Weiroot 154.Während der Untersuchung der generativen Eigenschaften wurde beobachtet, dass es Unterschiede in der Blütezeit der Sorten-Unterlagen-Kombinationen gibt. Auf GiSelA 5 veredelte Sorten blühten 1 oder 2 Tage früher als die Kontrolle. Es gab nur geringe Unterschiede in der Reifezeit der Kombinationen. Den größten Ertrag hatte die Sorte Linda, gefolgt von Katalin, Germersdorfi 3 und Piramis. Der größte Fruchtdurchmesser wurde bei der Süßkirschensorte Germersdorfi 3 gefunden, der größte Anteil an Früchten größer als 26 mm bei den Bäumen auf Weiroot 72 (23%), der geringste Anteil großer Früchte bei den Bäumen auf GiSelA 5. Bei Piramis wurde der größte Fruchtdurchmesser auf Cerasus avium C. 2493 gemessen.     

Pseudoreplication: a sine qua non for regional ecology

We question whether classical experimentation is adequate for real progress in landscape or regional ecology. One cannot do classical experimentation unless one can replicate the treatment. There is conflict between the need to replicate and the need to study processes at appropriately large scales.Because of the difficulties in doing controlled field experiments at regional scales, we propose that landscape ecologists take greater advantage of natural field experiments. Natural experiments must be coordinated, standardized, and synchronized over space and through time, and will require the cooperation of multiple investigators. Distributed computer networks can help provide the automated region-wide monitoring which will supply natural experiments with pre-treatment data.Regions or landscapes need not be replicated, and indeed, cannot be. One can achieve a relational understanding between a system's response and environmental characteristics. This understanding is not definitive, but allows for the development of testable hypotheses, in the classical sense. The confounding of space, time, and/or other environmental factors in pseudoreplicated natural experiments only allows for the development of hypotheses - how-possibly explanations. Discrimination among competing hypotheses can be done at smaller scales and used to infer processes occurring at larger scales. Use of natural and controlled field experiments in complementary roles is a more promising approach than views of one or the other as methodologically inferior.     

Bestimmung der S-Allele in Brennkirschensorten (Prunus avium L.)

Zusammenfassung Brennkirschen sind Süßkirschensorten, welche spezielle Eigenschaften für die Branntweinproduktion und die industrielle Verarbeitung besitzen. Ihr Hauptanbaugebiet in Deutschland befindet sich in Baden-Württemberg und in Teilen Bayerns. In den letzten Jahren wurde über einen geringen Fruchtbehang in neu gepflanzten Brennkirschenanlagen berichtet. Süßkirschen sind selbstinkompatibel und benötigen deshalb eine Bestäubersorte. Ursache ist ein gametophytischer Selbstinkompatibilitätsmechanismus, bedingt durch einen S-Lokus. Mit Hilfe molekularer Marker wurden deshalb Untersuchungen zu den S-Allel-Kombinationen in 62 Brennkirschensorten bzw. -herkünften aus dem Raum Oberkirch, Mittelbaden, durchgeführt. Im Ergebnis konnten zwei bisher noch nicht beschriebene S-Allele und 14 neue Inkompatibilitätsgruppen ermittelt werden. Ursache für den schlechten Fruchtansatz in neuen Kirschenanlagen mit den Hauptsorten Dolleseppler und Benjaminer sind die gleichen S-Allel-Kombinationen (S₁S₇) in beiden Sorten. Als mögliche Befruchtersorten werden die Sorten Schwarze Schüttler (S₆S₇), Hartwälder (S₄S₇), Feuerkirsche (S₄S₁₄) und die ehemalige Sorte Dolleseppler der Baumschule Kiefer (S₁S₄) empfohlen.     

Using diffusion models to simulate the effects of land use on grizzly bear dispersal in the Rocky Mountains

Timber harvests proposed for Trail Creek Watershed, in southwestern Montana, U.S.A., have been opposed because grizzly bear (Ursus arctos horribilis) dispersal from northern Montana wildernesses into the Greater Yellowstone Ecosystem may be less likely. We used an individual-based model to simulate grizzly bear responses to: 1) region-level management practices represented by ownership patterns, and 2) watershed-level changes in habitat availability due to proposed harvests and road building. We assigned permeabilities (i.e., values that represent how easily a bear can move through a patch) to ownership blocks (region-level) and habitat patches (watershed-level) based upon a literature review, and used a correlated random-walk diffusion model to simulate movements. Simulated bears were placed into rasterized landscapes in a stratified random manner. At the regional level, bears moved 1500 times (i.e., 1530 km), and their destinations were tallied. At the watershed level, the number of moves required for bears to leave the watershed were tallied. Sensitivity analyses were used to determine the variability of the results with respect to changes in some parameters of interest (i.e., permeabilities of private lands, harvest permeabilities, and disturbance indices).With the permeability of private land set at 50 (range: 0 to 99), simulated grizzlies did not disperse from the Scapegoat and Bob Marshall Wildernesses into Yellowstone National Park (0 of 10000 simulated individuals). Under the assumptions of this model, a linkage between the wildernesses in northern Montana and Yellowstone does not appear to exist. However, a significant number of simulated grizzlies (41%) dispersed from Anaconda Pintler Wilderness, which is near Trail Creek Watershed, into the wilderness ES in eastern Idaho. A linkage may exist between these sites.At the watershed-level, removal of forest habitat under proposed Harvest I (1.77% of the watershed cut) led to simulated grizzlies using slightly more moves (i.e., 5.6%, P=0.042) to exit the watershed than under existing conditions. Harvests of 3.5% of the watershed (planned Harvest II) did not alter the number of moves required to exit the watershed (P=0.068). When disturbances associated with roads and harvests were also examined, large increases in number of movements required to exit the watershed occurred ( 151%, P=0.002). These analyses suggest that grizzly bears would be disturbed while timber harvests were ongoing, but that long-term changes in movement would not occur if roads were closed following harvests. The analyses demonstrate the utility of applying individual-based diffusion models to landscape-level movements of animals, and identifies the need for telemetry studies to determine movement rates through specific habitats.     

Eintritt für Selbstpflücke, ein Kontinent ohne Frigopflanzen und ‘Elsanta‘

Zusammenfassung Die im Wachstum begriffene australische Erdbeerindustrie ist geprägt durch optimale Klimabedingungen und einfachen Zugang zu den Ressourcen, hohe Intensivierung und — wie in Kalifornien — hohe Erträge über 30 t/ha. Der Markt ist fast autark, Im- und Exporte frischer Erdbeeren spielen keine Rolle. Mit einem derzeitigen Produktionsvolumen von 40.000 t/Jahr auf ca. 1.300 ha wird eine Ausweitung auf 71.000 t bis zum Jahre 2008 angestrebt. Dies entspricht — bei ca. 20 Mio. Einwohnern Australiens — einer Steigerung des Erdbeerkonsums von 2,0 kg auf ca. 2,7 kg/Kopf und Jahr im Jahr 2008. Der gegenwärtige Erdbeerkonsum in Deutschland liegt bereits bei ca. 2,7 kg/Kopf und Jahr, wobei die Erdbeerimporte von ca. 120.000 t die Eigenproduktion von ca. 100.000 t auf 10.420 ha Anbaufläche — im Gegensatz zu Australien — übersteigen. Die drei australischen Hauptanbaugebiete konzentrieren sich auf Queensland im Nordosten, Perth im Westen und Victoria im Südosten Australiens. Der überwiegend einjährige Anbau erfolgt mit 45.000–66.000 Grünpflanzen/ha als Einzel- oder Doppelreihe in Dammkultur und Mulchfolie mit integriertem Fertigationsschlauch. Nach telefonischem Verkauf erfolgt die Lieferung im Kühl-LKW direkt an die Supermärkte oft weit entfernter Großstädte. Als tagneutrale Erdbeersorten dominieren mit 90% amerikanische Sorten wie Selva, Camarosa und Camino Real aus Kalifornien sowie Sweet Charlie und Festival aus Florida. Interessante Neuzüchtungen aus dem Jahre 2003 sind Rubygem, Sugarbaby, Brighteyes und Harmony aus Queensland als Kurztagssorten. Zusammen mit tagneutralen Sorten aus dem südlichen Züchtungsprogramm in Victoria stieg ihr Marktanteil in den letzten drei Jahren von 8% auf 10%. Primäre Zuchtziele des nördlichen Züchtungsprogramms in Queensland sind Kurztagssorten mit frühem Reifebeginn und hohem Ertrag über 1,2 kg/Pflanze. Das südliche Züchtungsprogramm um Victoria zielt dagegen sowohl auf Kurztagssorten als auch Tagneutralität sowie eine zusätzlich lange Ernteperiode über mindestens 10 Wochen. Während die Zuchtziele der äußeren Fruchtqualität wie Fruchtgröße, Fruchtfestigkeit und Glanz mit denen Deutschlands übereinstimmen, unterscheiden sich die der inneren Fruchtqualität wie süßer Geschmack bei gleichzeitiger Säurearmut und hoher Fruchtfleischfestigkeit.     

Bewertung von Pyrus-Arten auf Befall mit Birnengitterrost (Gymnosporangium sabinae Dicks.)

Zusammenfassung Birnengitterrost breitet sich in nicht kommerziell genutzten Anlagen zunehmend aus, vor allem dort, wo Pflanzenschutzmaßnahmen unterbleiben. In unserer Untersuchung wurde nach alternativen Lösungen gesucht, den Befall zu minimieren. Dazu wurden Pyrus-Arten und -Hybriden ohne Pflanzenschutz angezogen und der natürliche Befall bewertet. Die Arten Pyrus korzhinskyi, P. betulifolia, P. cordata und die Hybride P. salicifolia Pendula zeigten den geringsten Befall. Sie könnten als Züchtungsausgangsmaterial eingesetzt werden. Ebenso scheint eine Prüfung als Park- oder Landschaftsgehölz sinnvoll.     

Using spatial analysis to drive reserve design: a case study of a national wildlife refuge in Indiana and Illinois (USA)

Our refuge design strategy involves a landscape approach formulated to complement existing management efforts and employ restoration in the heavily degraded Kankakee River watershed in northeastern Indiana and northwestern Illinois. The watershed historically contained an approximately 400,000 ha wetland (Grand Marsh), a diverse riverine system, oak savanna, and prairie. Today only fragments of these habitats remain. The U.S. Fish and Wildlife Service developed a preliminary project proposal (PPP) during the summer of 1996 to protect and restore habitat within the watershed by establishing the Grand Kankakee Marsh National Wildlife Refuge. The Indiana and Illinois Gap Analysis projects provided the resources to move beyond the expert workshop approach to a data-driven method for initial reserve design. Using visual analysis of various Gap Analysis data layers, we established Focus Areas that formed the basis for the Environmental Assessment and the Economic Impact Study required under the National Environmental Policy Act (NEPA). We have initiated Phase II of the reserve design analysis using preliminary results of the Indiana Gap Analysis project and C-Plan software in conjunction with the New South Wales National Parks and Wildlife Service.     

A model of arctic tundra vegetation derived from topographic gradients

We present a topographically-derived vegetation model (TVM) that predicts the landscape patterns of arctic vegetation types in the foothills of the Brooks Range in northern Alaska. In the Arctic there is a strong relationship between water and plant structure and function and TVM is based on the relationships between vegetation types and slope (tan ) and discharge (), two independent variables that can be easily derived from digital terrain data. Both slope and discharge relate to hydrological similarity within a landscape: slope determines the gravitational hydrological gradient and hence influences flow velocity, whereas discharge patterns are computed based on upslope area and quantify lateral flow amount. TVM was developed and parameterized based on vegetation data from a small 2.2 km² watershed and its application was tested in a larger 22km² region. For the watershed, TVM performed quite well, having a high spatial resolution and a goodness-of-fit ranging from 71–78%, depending on the functions used. For the larger region, the strength of the vegetation types predictions drops somewhat to between 56–59%. We discuss the various sources of error and limitations of the model for purposes of extrapolation.     

Bird assemblages in forest fragments within Mediterranean mosaics created by wild fires

The role of habitat heterogeneity as a key factor in determining species pools in habitat mosaics has been acknowledged, but we still know little on the relative importance of the different ecological processes acting within such complex landscapes. We compared species richness and distribution in forest fragments imbedded in shrub-lands to those in continuous forests or in continuous shrublands. We examined the consistency of our data with the predictions of two hypotheses: 1) the Habitat fragmentation hypothesis which states that fragmentation has negative effects on the species from the original continuous habitat; 2) the Habitat supplementation /complementation hypothesis which stipulates that the presence of a matrix habitat around the fragments will mitigate negative effects on the species from the original habitat (supplementation) or allow the presence of species that depend on the presence of both the fragment and matrix habitats (complementation). We show that: 1) species richness in forest fragments did not differ from species richness in segments of continuous forests of equal area; 2) the bird community of forest fragments got impoverished in some forest species but a higher proportion of species common in continuous forests were not affected by fragmentation; 3) fragment communities had a significant proportion of common species that were scarce in, or absent from both continuous forests and shrublands. While, a few forest species supported predictions from the fragmentation hypothesis, occurrence patterns observed in several other species were consistent with either the supplementation or the complementation hypotheses. Our results suggest that there is no single hypothesis that properly captures the consequences of a shift from continuous forests to a mosaic of forest fragments and shrublands and that different ecological mechanisms act in conjunction to determine species pools in habitat mosaics. Habitat heterogeneity at a local scale appears a key factor in maintaining bird diversity in fire driven Mediterranean landscapes.     

Human influence on the abundance and connectivity of high-risk fuels in mixed forests of northern Wisconsin,USA

Though fire is considered a natural disturbance, humans heavily influence modern wildfire regimes. Humans influence fires both directly, by igniting and suppressing fires, and indirectly, by either altering vegetation, climate, or both. We used the LANDIS disturbance and succession model to compare the relative importance of a direct human influence (suppression of low intensity surface fires) with an indirect human influence (timber harvest) on the long-term abundance and connectivity of high-risk fuel in a 2791 km² landscape characterized by a mixture of northern hardwood and boreal tree species in northern Wisconsin. High risk fuels were defined as a combination of sites recently disturbed by wind and sites containing conifer species/cohorts that might serve as ladder fuel to carry a surface fire into the canopy. Two levels of surface fire suppression (high/current and low) and three harvest alternatives (no harvest, hardwood emphasis, and pine emphasis) were compared in a 2×3 factorial design using 5 replicated simulations per treatment combination over a 250-year period. Multivariate analysis of variance indicated that the landscape pattern of high-risk fuel (proportion of landscape, mean patch size, nearest neighbor distance, and juxtaposition with non fuel sites) was significantly influenced by both surface fire suppression and by forest harvest (p > 0.0001). However, the two human influences also interacted with each other (p < 0.001), because fire suppression was less likely to influence fuel connectivity when harvest disturbance was simultaneously applied. Temporal patterns observed for each of seven conifer species indicated that disturbances by either fire or harvest encouraged the establishment of moderately shade-tolerant conifer species by disturbing the dominant shade tolerant competitor, sugar maple. Our results conflict with commonly reported relationships between fire suppression and fire risk observed within the interior west of the United States, and illustrate the importance of understanding key interactions between natural disturbance, human disturbance, and successional responses to these disturbance types that will eventually dictate future fire risk.This revised version was published online in May 2005 with corrections to the Cover Date.     

Für eine Verdopplung der Ca-Gehalte der Äpfel sind auch unter günstigen Bedingungen mindestens 12 Spritzungen mit Kalziumchlorid erforderlich

Zusammenfassung Siebenjährige Apfelbäume der Sorten Elstar, Gala, Karmijn de Sonnaville, Pinova und Pilot wurden 2003 von Juni bis September 12-mal mit 400 l/ha CaCl₂-Lösungen gesprüht. Kontrollen (Fu) waren Bäume, die nur mit Fungiziden (Discus, Dithane Ultra oder Kumulus) behandelt wurden. Bei den Behandlungen FuCa wurde den Fungiziden CaCl₂ (ansteigend von 5–25 g/l) zugegeben. Die Behandlungen FuCaTe enthielten zusätzlich das Netzmittel Plantacare 1200 UP, um die Benetzung der Früchte (Oberflächenspannung 29–30 mN/m) zu verbessern. Die Ca-Konzentrationen in den Kontrollfrüchten (Schale plus 5 mm Fruchtfleisch) variierten von 3,6 (Elstar) bis 9,0 mg/100 mg Frischsubstanz (Pilot). Durch die Fruchtdüngung (FuCa) stiegen die Konzentrationen auf 6,3 bzw. 13,3 mg/100 g Frischsubstanz (FS). Zugabe des Netzmittels (FuCaTe) erhöhte die Ca-Gehalte weiter auf 6,8 bzw. 14,9 mg/100 g FS. Bezogen auf die Behandlung Fu entspricht das einer Erhöhung der Ca-Konzentrationen der Früchte auf 163 bzw. 193%. Der Tensideffekt auf die Ca-Konzentrationen der Früchte wird auf die schnellere Penetration des CaCl₂ infolge besserer Benetzung zurückgeführt, da ein Einfluss auf die Retention der Spritzbrühe ausscheidet. Von der applizierten Ca-Dosis in Höhe von 26 kg/ha wurden nur 2,3 kg in den Früchten (50 t/ha) wieder gefunden, was einer Wiederfindungsrate von 8,8% entspricht. Die beobachtete Erhöhung der Ca-Konzentrationen der Früchte durch 12 Spritzungen dürfte in der Nähe des maximal Möglichen liegen, da die Witterungsbedingungen günstig und die Ca-Konzentrationen der Spritzbrühen hoch waren.     

Fuzzy k-means classification of topo-climatic data as an aid to forest mapping in the Greater Yellowstone Area, USA

Previous attempts to quantify topographic controls on vegetation have often been frustrated by issues concerning the number of variables of interest and the tendency of classification methods to create discrete classes though species have overlapping property sets (niches). Methods of fuzzy k-means classification have been used to address class overlap in ecological and geographical data but previously their usefulness has been limited when data sets are large or include artefacts that may occur through the derivation of topo-climatic attributes from gridded digital elevation models. This paper presents ways to overcome these limitations using GIS, spatial sampling methods, fuzzy k-means classification, and statistical modelling of the derived stream topology. Using data from a ca. 3600 km² forested site in the Greater Yellowstone Area, we demonstrate the creation of meaningful, spatially coherent topo-climatic classes through a fuzzy k-means classification of topo-climatic data derived from 100 m gridded digital elevation models (DEMs); these classes were successfully extrapolated to adjacent areas covering a total of ca. 10000 km². Independently derived land cover data and middle infrared corrected Landsat TM derived estimates of Normalised Difference Vegetation Index (M_NDVI) at 575 independently sampled sites were used to evaluate the topo-climatic classes and test their extrapolation to the larger area. Relations between topo-climatic classes and land cover were tested by ² analysis which demonstrated strong associations between topo-climatic class and 11 of the 15 cover classes. Relations between M_NDVI and topo-climatic classes proved to be stronger than relations between M_NDVI and the independent cover classes. We conclude that the fuzzy k-means procedure yields sensible and stable topo-climatic classes that can be used for the rapid mapping of large areas. The value of these methods for quantifying topographic controls on biodiversity and the strength of their relations with computed NDVI values warrant further investigation.     

A proximity-based approach to assessing habitat

Planning for either a single species, multiple species, or ecosystems is greatly dependent on spatial interactions in the landscape. Problems exist for evaluating wildlife habitat changes over large ranges of space and time. This paper illustrates the use of Geographic Information Systems (GIS) to evaluate habitat for a single species, ruffed grouse (Bonasa umbellus), following a time series of forest harvests. A habitat suitability model for ruffed grouse is utilized on a two-township study area in north-central Minnesota to assess the habitat suitability changes over time using an even-aged area-control harvesting plan. The results are presented as a habitat quality change map and a contingency table, representing the movement of habitat class areas between time periods resulting from the proposed harvesting. We developed a neighborhood definition to allow for spatially varying habitat values. This work illustrates the ability to look ahead and around in estimating the impact on wildlife habitat resulting from alternative future management activities.Now with the U.S. Forest Service, Sitka, AK 99835.     

Effects of patch shape on the number of organisms

This study examined effects of habitat patch shape on the abundance of organisms. The effects of patch shape were considered in terms of (1) immigration and emigration of organisms. (2) the amount of available resources in a patch and (3) spatial and temporal heterogeneity of the organisms and environment. I hypothesized that (1) the number of organisms would increase as patch shape elongates because organisms are more likely to encounter an elongated patch, (2) the number of organisms in a patch would remain constant for all patch shapes where the number of organisms in a patch was limited by the amount of resources, because patch shape does not change the patch area that is directly associated with the amount of patch resources, and (3) spatial and temporal variation of the abundance of organisms would increase as patch shape elongates because an elongated patch is more likely to interact with the variable surrounding matrix.Common millipedes,Oxidus gracilis, and their habitat, plywood boards of five shapes (width:length ratio; 11, 14, 19, 136, 1144) with an area of 900 cm² were placed in forest and old field and the number of millipedes appearing under the boards was monitored. Significantly higher mean number of millipedes under the boards was observed at a patch with an elongated shape in the forest and the old field. A significant positive correlation was observed between perimeter length of a patch and the number of millipedes in the old field. The temporal and spatial variation of the number of millipedes was high in the old field. The spatial and temporal variation was higher for boards with elongated shape.