Effects of livestock breed and grazing intensity on biodiversity and production in grazing systems. 1. Nutritive value of herbage and livestock performance

Reduction of grazing intensity and the use of traditional instead of commercial breeds has frequently been recommended to meet biodiversity and production goals in sustainable grazing systems in Europe. To test the impact of such practices across a range of contrasting grassland types, integrated measurements of foraging behaviour, agronomic production and botanical, structural and invertebrate biodiversity were made over three years on four sites in the UK, Germany, France and Italy. The sites in the UK and Germany were mesotrophic grassland with high productivity and low to moderate initial levels of plant diversity, and were grazed by cattle. The French site was a semi‐natural, species‐rich grassland grazed by cattle. The Italian site contained a wider range in plant diversity, from species‐rich to mesotrophic grassland, and was grazed by sheep. The treatments were: MC, moderate grazing intensity with a commercial breed – this was designed to utilize herbage growth for optimum livestock production; LC, lenient grazing intensity with a commercial breed – this was designed to increase biodiversity by not fully utilizing herbage growth; and LT, lenient grazing intensity with a traditional breed – this was also designed to increase biodiversity. Neither fertilizers nor pesticides were applied. The nutritive value of the herbage and the performance of the livestock were measured. Mean stocking rates were proportionately 0·30–0·40 lower and mean sward heights and herbage mass on offer were 0·30–0·50 higher on the LC and LT treatments compared with the MC treatment. The proportion of live and dead material, and leaves and stems in the herbage, its chemical composition and nutritive value were little affected by the treatments. Individual livestock performance, measured as liveweight gain, showed no consistent response to treatment. In Germany, performance on the MC treatment was slightly lower than on the LC and LT treatments but no such difference was found on the sites in the other countries. Livestock breed did not have a strong effect on livestock performance. In the UK and France the traditional breeds had a lower performance but this was not the case in Germany or Italy. Livestock performance per ha of the LC and LT treatments was up to 0·40 lower than of the MC treatment. It is concluded that biodiversity‐targeted extensive grazing systems have potential to be integrated into intensive livestock production systems because the individual livestock performance reaches a similar level compared to a moderate grazing intensity. Traditional breeds did not have a production advantage over commercial breeds on extensively managed pastures.     

Influence of defoliation regime on herbage production and characteristics of intake by dairy cows as affected by grazing intensity

Three contrasting defoliation regimes for dairy cows—four cuts annually, severe rotational grazing and lax rotational grazing integrated with cutting—were compared in terms of herbage production, selection and intake per cow. Lax and severe grazing were compared simultaneously and grazing intensity was characterized by means of changes in herbage mass and sward height during grazing.
Herbage growth and yield were similar under cutting, severe grazing and lax grazing/cutting (120 t organic matter (OM) ha⁻¹). Herbage quality and leaf production were highest with severe grazing, which also had a less marked seasonal pattern of growth. The herbage intake per cow was 111 kg OM d⁻¹ with severe grazing and proportionately (0-20) higher at lax grazing/ cutting. 050 of the herbage yield was harvested by cutting at lax grazing/cutting. Selectivity was described with high certainly by the nutrient content prior to defoliation and by the degree of defoliation. Models to quantify this are presented.
Grazing intensity could be described by the size and degree of utilization of the fouled area, and herbage intake was quantified by means of herbage height before and after grazing. Within the range of 80-240 mm sward height prior to grazing, height measurements led to realistic and more accurate estimates than those obtained by measuring organic matter disappearance.
Herbage utilization was depressed significantly by increasing maturity of herbage due to lower nutritive value, but in particular due to reduced green leaf content and increased reproductive development. If sward height did not exceed 250 mm at any time, good utilization by grazing could be obtained.     

Effect of altering the grazing interval on growth and utilization of grass herbage and performance of dairy cows under rotational grazing

The effects of short grazing intervals in the early part of the grazing season on the growth and utilization of grass herbage, and the performance of grazing dairy cows, in a rotational grazing system were examined. Seventy-six cows were allocated to two grazing treatments: a normal rotation treatment (20-d rotations for the first 60 d) and a short rotation treatment (12-, 8-, 8-, 8-, 12- and 12-day rotations). Thereafter, both treatments had the same grazing interval and over the season as a whole both treatments received the same amount of nitrogen fertilizer and were stocked at the same rate. The short rotation treatment significantly reduced pre- and post-grazing sward heights and pre-grazing herbage mass in May and June. Total herbage production was significantly lower on the short than the normal rotation treatment as a result of a significant reduction in the growth rate of herbage in May and June. The short rotation treatment had a significantly lower milk output per cow. Grazing shorter swards, as a result of shorter rotations, significantly reduced herbage intake, reflecting reductions in intake per bite, grazing time and total bites per day. Treatment had no significant effect on herbage quality or pre- and post-grazing sward height in August and September, despite the increased grazing severity in May and June with the short rotations. The severity of rotation length in this instance had a detrimental impact on animal performance, whereas a more modest reduction in grazing interval may control herbage production, without reducing animal performance.     

Effect of different grazing management systems on the herbage mass and pasture height of a Nardus stricta grassland in western Scotland, United Kingdom

The aims of this study were to examine the effect of three grazing treatments (year‐round stocking rates of 0·8 ewes ha^?1, 0·5 ewes ha^?1 and 0·5 ewes ha^?1 plus grazing cattle in summer), imposed for 4 years, on the herbage mass and surface height of a Nardus stricta‐dominated grassland in western Scotland and to obtain estimates of annual productivity of this grassland. Nardus stricta‐dominated grassland comprised proportionately 0·20 of the grazing area. Stocking rate of sheep had no significant effect on the herbage mass of the grassland in the first 2 years of the experiment, although mean summer pasture heights were significantly higher under the lower stocking rate of sheep. The pasture on the treatment with cattle grazing in summer had a significantly lower herbage mass and lower surface height than the two sheep‐only grazing treatments. Year‐to‐year variation in the herbage mass and surface height of herbage in summer was greater than the effect of treatments. Despite changes in surface height, the structural diversity of the grasslands was not increased by the treatments. The annual production of vascular plant material ranged from 417 g DM m^?2 in 1994 to 628 g DM m^?2 in 1996.     

Evaluation of the effects of ewe prolificacy potential and stocking rate on herbage production,utilization, quality and sward morphology in a temperate grazing system

This study investigated the effect of ewe prolificacy potential (PP; predicted number of lambs born ewe^?1 year^?1), stocking rate (SR; ewes/ha) and their interaction on herbage dry matter (DM) production, utilization, quality and sward morphology within a temperate grass‐based lamb production system. The study had a 2 × 3 factorial design, consisting of two ewe PP as dictated by sire breed (180 medium prolificacy potential (MP—Suffolk crossbred) and 180 high prolificacy potential ewes (HP—Belclare crossbred)) and three SR: low (LSR; 10 ewes/ha), medium (MSR; 12 ewes/ha) and high (HSR: 14 ewes/ha). Each treatment was managed in a rotational grazing system, with LSR, MSR and HSR treatments grazing to target post‐grazing sward heights (PGSH) of 4.55, 4.15 and 3.75 cm respectively. Herbage DM production (above target PGSH) and utilization were highest at the HSR, intermediate at the MSR and lowest at the LSR (p < .001). Ewe PP had no effect on herbage DM production, utilization, quality or sward morphology (p > .05). The proportion of leaf in the sward (above target PGSH) was 4% greater in MSR and HSR compared with LSR (p < .05). In conclusion, findings demonstrate the potential to support increased ewe PP through the selection of ewe genotypes of a genetically higher PP and lower mature live weight and increased SR within a temperate grass‐based lamb production system.     

Changes in the herbage nutritive value and yield associated with threshold responses of vegetation to grazing in Mongolian rangelands

We examined the changes in the nutritive value and yield of herbage along a grazing gradient, where abrupt changes in community composition occurred, at multiple ecological sites in Mongolian rangelands. At grassland sites, changes in the herbage nutritive value could be attributed to rapid replacement of perennial grasses or forbs with weedy annual forbs along a grazing gradient. Crude protein (CP) concentration increased sharply in approaching the source of grazing gradient, whereas neutral detergent fibre and metabolizable energy (ME) concentrations decreased sharply. As ME can be utilized as a main index of herbage nutritive value, these results indicated its overall loss with severe grazing. Consequently, gradual increases in the yields of CP and ME in the direction of the gradient source at the grassland sites did not necessarily indicate the improvement of rangeland condition. In contrast, at shrubland sites, we could not generally detect any significant trends in the herbage nutritive value. The yield of ME gradually decreased in the direction of gradient source, suggesting that grazing affects herbage yield rather than herbage feed value at shrubland sites. Thus, the nutritive value and yield of herbage can be modified greatly in association with nonlinear responses of vegetation to livestock grazing.     

Effects of cattle grazing on herbage quality in a herbaceous Mediterranean rangeland

The main objective of this study was to determine to what extent grazing pressure and timing modulate the seasonal progression of herbage quality in hilly Mediterranean grassland systems. The study was conducted during six consecutive years between 2003 and 2008 at the Karei Deshe experimental farm, in eastern Galilee, Israel, dominated by rich hemicryptophytic grassland. Treatments included two different grazing intensities, heavy and moderate, with 1·1 and 0·55 cows ha^?1, respectively, and management that included a continuous and a seasonal stocking system that was divided at both intensities into early and late grazing. Herbage samples were analysed for digestibility, crude protein, neutral detergent fibre and acid detergent fibre contents. Significant differences in herbage quality were found between seasons and years. Herbage quality was significantly higher in paddocks grazed continuously or early in the season. Herbage quality increased with increasing grazing intensity as younger herbage and continued re‐growth were maintained during the green season. The greater difference between herbage qualities was found at the peak of the growing season. The significant differences found in herbage quality emphasize the importance of the decision‐making process aimed at improving cattle grazing management in Mediterranean rangelands and its consequences for the sustainability of the system.     

Impact of deferred grazing and fertilizer on herbage production,soil seed reserve and nutritive value of native pastures in steep hill country of southern Australia

Developing sustainable grazing management systems based on perennial species is critical to preventing land degradation in marginal land classes. A field study was conducted from 2002 to 2006 to identify the impacts of deferred grazing (no defoliation of pastures for a period generally from spring to autumn) and fertilizer application on herbage accumulation, soil seed reserve and nutritive value in a hill pasture in western Victoria, Australia. Three deferred grazing strategies were used: short‐term deferred grazing (no defoliation between October and January), long‐term deferred grazing (no defoliation from October to the autumn break) and optimized deferred grazing (withholding time from grazing commenced between annual grass stem elongation and seed head emergence and concluded in February/March). These treatments were applied with two fertilizer levels (with or without fertilizer at 50 kg phosphorus ha^?1 and 2000 kg lime ha^?1 applied in year 1 only) in a factorial arrangement and two additional treatments: continuous grazing (CG) and no grazing (NG) in year 1. The deferred grazing treatments on average produced herbage dry matter of 4773 kg ha^?1, the NG produced 4583 kg ha^?1 and the CG produced 3183 kg ha^?1 in year 4 (2005–06) of the experiment. Deferred grazing treatments with and without fertilizer application produced an average of 5135 and 4411 kg DM ha^?1 respectively. Averaged over 4 years, deferred grazing increased the germinable seed pool of perennial grasses by 200% and annual grasses by 50% (except optimized deferred grazing that considerably decreased the annual grass seed pool) compared with the CG. The best of the deferred grazing strategies increased the digestibility of pastures by 7% compared with the CG. The results demonstrated that deferred grazing from spring to autumn followed by rotational grazing could be an effective tool to increase herbage production and soil seed pool and improve the digestibility of native pastures in the steep hill country of southern Australia.     

The effect of stocking rate and calving date on grass production,utilization and nutritive value of the sward during the grazing season

A 2‐year whole‐farm system study compared the accumulation, utilization and nutritive value of grass in spring‐calving grass‐based systems differing in stocking rate (SR) and calving date (CD). Six treatments (systems) were compared over two complete grazing seasons. Stocking rates used in the study were low (2·5 cows ha^?1), medium (2·9 cows ha^?1) and high (3·3 cows ha^?1), respectively, and mean CDs were 12 February (early) and 25 February (late). Each system had its own farmlet of eighteen paddocks and one herd that remained on the same farmlet area for the duration of the study. Stocking rate had a small effect on total herbage accumulation (11 860 kg DM ha^?1 year^?1), but had no effect on total herbage utilization (11 700 kg DM ha^?1 year^?1). Milk and milk solids (MS; fat + protein) production per ha increased by 2580 and 196 kg ha^?1 as SR increased from 2·5 to 3·3 cows ha^?1. Milk production per ha and net herbage accumulation and utilization were unaffected by CD. Winter feed production was reduced as SR increased. Increased SR, associated with increased grazing severity, resulted in swards of increased leaf content and nutritive value. The results indicate that, although associated with increased milk production per ha, grazed grass utilization and improved sward nutritive value, the potential benefits of increased SR on Irish dairy farms can only be realized if the average level of herbage production and utilization is increased.     

Herbage mass and nutritive value of herbage of extensively managed temperate grasslands along a gradient of shrub encroachment

Semi-natural grasslands often serve as important reserves of biodiversity. In Europe extensive grazing by livestock is considered an appropriate management to conserve biodiversity value and to limit shrub encroachment. However, little is known about the influence of shrubs on agronomic values. A gradient analysis of shrub-invaded temperate grasslands (from shrub-free to pioneer forest) in Germany was carried out to test the hypothesis that herbage mass and variables describing nutritive value of herbage decrease with increasing shrub encroachment. The herbage mass of dry matter (DM), variables describing the nutritive value of herbage, composition of the vegetation and mean of Ellenberg's indicator values were analysed with respect to the extent of shrubs. There was a reduction of herbage mass of DM from 3570 to 210 kg ha⁻¹ with increasing shrub encroachment. Metabolizable energy concentration of herbage ranged from 8·9 to 10·2 MJ kg⁻¹ DM and crude protein concentration from 72 to 171 g kg⁻¹ DM, both measures being positively correlated with shrub occurrence. Increasing shrub occurrence was associated with a decrease in water-soluble carbohydrates concentration (from 151 to 31 g kg⁻¹) and a reduction in the indicator 'forage value'. The results indicate a potentially large agronomic value for shrub-encroached temperate grasslands.     

The effects of variation in sward characteristics upon the ingestive behaviour and herbage intake of calves and lambs under a continuous stocking management

Groups of calves and lambs grazed on plots of S23 perennial ryegrass, in successive periods of 2, 2 and 3 weeks, at stocking rates which resulted in a slow change in herbage mass on each plot and measurements were made of ingestive behaviour (biting rate, bite size and grazing time) and herbage intake. As the green herbage mass was progressively reduced from approx. 3000 to 1000 kg OM per ha between periods, biting rate and grazing time increased, but insufficiently to offset the rapid fall in bite size so that herbage intake declined by 24% and 39% for calves and lambs respectively. Patterns of response were similar in calves and lambs, though in all parameters except biting rate the lambs appeared to be more sensitive than the calves to changes in sward conditions and there was no evidence that lambs were better adapted to grazing on short swards than calves. Intake increased in both calves and lambs up to levels of 2500–3000 kg green OM per ha, though this response may have been attributable in part to the reinforcing effects of increasing herbage mass and diet digestibility. Lambs consistently selected a more digestible diet than calves. Both calves and lambs increased grazing time substantially in response to limiting sward conditions, in contrast to results in earlier strip-grazing trials with calves, thus supporting the suggestion that the lack of response under strip-grazing is a characteristic of the management imposed.     

Effects of ruminal fill on short-term herbage intake rate and grazing dynamics of beef heifers

The impact of manipulating ruminal fill (RF) on intake rate of herbage and grazing dynamics was measured with three rumen‐cannulated beef heifers grazing Bermudagrass pastures individually. The treatments compared were removal of proportions of rumen contents of 0 (treatment RF0), 0·33 (treatment RF33), 0·66 (treatment RF66) and 1·00 (treatment RF100). Treatments were randomly applied in a 3 × 4 Youden‐square design. The rumens were emptied before and after planned grazing sessions (30 min) to set up the treatments, and to estimate intake rate and bite mass, respectively. Measurements were made of bite rate, bites per feeding station, feeding stations per minute, intake per feeding station, time per feeding station, eating and searching step rates and times. Apparent bite area and area grazed per feeding station were calculated. Ruminal fill affected short‐term intake rate and changed grazing dynamics. As RF increased, step rates, searching times, bite mass, apparent bite area, bites per feeding station, area grazed per feeding station, time per feeding station and intake per feeding station decreased (P < 0·01) while step times, eating step rates and bite depth increased (P < 0·01). The results of the present study indicate that RF is an important factor governing the intake characteristics and behaviour of grazing beef heifers.     

The effects of allowance of deferred herbage and concentrate supplementation when offered to ewe lambs,and system of allocation when offered to ewes in late gestation,on animal performance

The effects of allowance of extended (deferred) grazed herbage (AEGH) and herbage allocation management (HAM) were studied in ewe lambs (248) and late‐gestation ewes (152), respectively, on commercial farms in south‐east Ireland in 2005–06. In Experiment 1, which consisted of four treatments, the effects of AEGH (0·75, 1·25 and 1·75 kg DM per head daily) and concentrate supplementation (0·5 kg per head daily with the 0·75 kg DM herbage allowance) on lamb performance during the extended grazing (16 December to 3 March) and subsequent grazing (4 March to 11 August) seasons were evaluated. Increasing AEGH increased herbage intake linearly (P < 0·001) and live weight (P < 0·001) at the end of extended grazing and the subsequent grazing season. In Experiment 2, single‐ and twin‐bearing ewes were allocated to either a conventional (single‐ and twin‐bearing ewes grazed separately) or leader–follower system (twin‐ and single‐bearing ewes, as leaders and followers respectively) of HAM from 30 January to 24 March. The same quantities of herbage were offered daily for each system. System of HAM affected ewe condition score at lambing but did not alter (P > 0·05) subsequent lamb birth or weaning weights. It is concluded that increasing AEGH to ewe lambs increased liveweight gain during extended grazing and resulted in heavier animals in mid August of the subsequent grazing season. For ewe lambs each 1 kg concentrate DM had the same feed value as 2·4 kg DM AEGH. Use of a leader–follower system for ewes in late pregnancy did not alter lamb birth weight or subsequent performance.     

THE EFFECTS OF CUTTING AND GRAZING SYSTEMS ON HERBAGE PRODUCTION FROM GRASS SWARDS

Studies were made on two perennial ryegrass–dominant swards of the effects of cutting and grazing systems of herbage production. A motor scythe was used for cutting and sheep in small enclosures for grazing. In both systems, herbage production was measured by a ground-level sampling technique, using sheep shears, and the difference between pre-treatment and post-treatment herbage was expressed as ‘utilized yield’. Correction for soil contamination is ensured by quoting all results on an organic-matter basis. Grazing treatments averaged 14–16% more organic matter and 36–45% more crude protein than cutting treatments in both experiments. These differences were ascribed to recycling of N under grazing. Infrequent defoliation gave higher yields than frequent, and severe defoliation higher yields than lenient under both cutting and grazing systems. The trends shown by cutting and grazing intensities are comparable and can be quantified. There is need to examine closely the relationships between the effects of cutting and grazing techniques on herbage production, since such knowledge would allow a more accurate prediction of the relevance to the grazing situation of results obtained under cutting.     

Selective defoliation by sheep according to slope and plant species in the hill country of New Zealand

The objective of the study was to evaluate the grazing behaviour by sheep in hill country paddocks in New Zealand which had received two long‐term fertilization and stocking rate treatments [high fertility–high stocking rate (HH); low fertility–low stocking rate (LL)]. Herbage accumulation and selective grazing were evaluated within low slope (LS), medium slope (MS) and high slope (HS) categories. Transects lines were placed and tillers of Agrostis capillaris and Lolium perenne in the LS category; A. capillaris, Anthoxanthum odoratum and L. perenne in the MS category; and A. capillaris and A. odoratum in the HS category were marked. The leaf length of each marked tiller was measured and used to determine selective grazing over 3 weeks during each season. The highest herbage accumulation rates were during spring and the lowest in summer and winter. The LS category showed the highest herbage accumulation rates and HS the lowest. Except for autumn, the marked tillers were more frequently grazed in the HH than in the LL paddock. During summer, autumn and spring, grazing frequency in the slope categories was in the order LS > MS > HS. During winter sheep did not discriminate between slope category. During summer, autumn and spring, sheep did not selectively graze the species studied but this was not the case during winter. Overall, sheep selectively grazed L. perenne. In all the seasons L. perenne consistently had the longest leaves but within species there was no consistent relationship between leaf length and probability of being grazed. Selective grazing changed through the year according to herbage accumulation rate. Sheep concentrated grazing in the category LS when herbage accumulation rate was high, but they did not discriminate between slope categories in winter when herbage accumulation rate was low.     

Setting management limits for the production and utilization of herbage for out-of-season grazing

Three experiments were carried out on perennial ryegrass‐dominant swards to provide a basis for recommendations for the limits to (a) building up and timing of utilization of a herbage ‘bank’ for out‐of‐season grazing and (b) duration and intensity of early spring grazing in the United Kingdom and Ireland. In experiment 1, the effect of regrowth interval (from 7 September, 20 October, 17 November or 15 December) in autumn on herbage accumulation, leaf turnover and on subsequent spring growth was investigated. Swards regrown from early September reached maximum herbage mass (about 3 t ha^–1 DM) and leaf lamina content in mid‐November, by which time senescence rate exceeded rate of production of new leaves. New leaf production and senescence rates were greater in swards remaining uncut until December than in those cut in October or November. Time of defoliation up to December had no effect on spring herbage mass in the subsequent spring. Defoliating in March reduced herbage mass in late May by less than 20%. Experiment 2 investigated the progress in herbage growth and senescence in swards regrowing from different times in late summer and autumn to produce herbage for utilization beyond the normal grazing season. Treatments in a randomized block design with three replicates were regrowths from 19 July, 8 August, 30 August and 20 September. Based on a lower ceiling of leaf and total herbage mass being reached with progressively later regrowths, beyond which leaf senescence generally exceeded leaf production and herbage mass declined, it was concluded that currently recommended rotation lengths for this period should extend from 3 weeks in late July to 8 weeks for swards previously grazed in mid‐September. In both experiments, leaf senescence commenced earlier (by one leaf‐age category) than previously published estimates and so brought forward the time at which senescence rates balanced leaf growth rates. In experiment 3, designed to evaluate the effect of daily grazing period and intensity in early spring on herbage regrowth, dairy cows grazed successive plots (replicates) for 2 or 4 h each day at two intensities (target residual heights of 5 or 7 cm) in March to mid‐April. Regrowth rate was similar in all treatments including the ungrazed control, despite soil moisture content being relatively high on occasions. Tiller density was significantly reduced in May by grazing plots in early or mid‐April. It is concluded that in autumn there are limits to which rotation lengths should be extended to produce herbage for out‐of‐season grazing owing to attainment of ceiling yields. Although utilization in early spring may reduce herbage availability in spring, out‐of‐season utilization need not reduce herbage growth rates in early spring.     

Seasonal changes in herbage mass and nutritive value of a range of grazed legume swards under Mediterranean and cool temperate conditions

Seasonal changes in herbage mass and herbage quality of legume‐based swards under grazing by sheep or cattle were investigated at four locations in climatically different zones of Europe: Sardinia (Italy), southern France, northern Germany and south‐west England (UK). At each location standard treatments were applied to legumes typical of species widely used in each locality: Medicago polymorpha in Italy, Medicago sativa in France, and Trifolium repens in Germany and in UK. At each site comparisons were made of two other legumes: Trifolium subterraneum and Hedysarum coronarium in Italy, Onobrychis sativa and Trifolium incarnatum in France, Trifolium pratense and Lotus corniculatus in Germany, and Trifolium ambiguum and L. corniculatus in UK. Legumes were sown in mixture with locally appropriate companion grasses, and measurements were made over two or three grazing periods. In Italy M. polymorpha swards gave the greatest herbage mass in grazing period 1 but H. coronarium was more persistent. At the French site all legumes established poorly with no significant herbage mass differences between treatments. At both the UK and German sites L. corniculatus maintained a high proportion of legume in the sward; T. repens showed poor persistence under continuous sheep grazing in UK but persisted under cattle grazing in Germany, while T. ambiguum was slow to establish in the UK, and T. pratense proved to be of comparable herbage mass to the standard T. repens‐based sward in the last year of the experiment. The concentration of crude protein and in vitro digestibility of organic matter in the dry matter of herbage showed greater within‐season variation than between treatments at each site. It is concluded that, in addition to currently used species, legume‐based swards containing H. coronarium, O. sativa and L. corniculatus all have potential to contribute to forage production for low‐input grazing and their use merits further consideration in systems of livestock production in Europe.     

Some effects of herbage composition, as influenced by previous grazing management, on milk production by cows grazing on ryegrass/white clover pastures. 2. Milk production in late spring/summer: effects of grazing intensity during the preceding spring period

Three experiments (2a, 2b and 2c) were carried out to examine the effects of different grazing intensities imposed during spring on subsequent herbage composition and milk production measured in summer.
Increased intensity of grazing imposed during spring (including the period of reproductive development in ryegrasses) produced swards in early summer with lower pre-grazing herbage masses, which contained higher concentrations of green leaf, clover and digestible nutrients, but lower concentrations of grass stem and senescent material.
In the first and second experiment cows were given a common daily allowance of total herbage dry matter (DM). The cows grazing on low-mass swards in early summer produced larger daily yields of milk, fat and protein than those grazing on the high-mass swards.
In the third experiment, cows were given a common daily allowance of green leaf DM from three swards which differed in pre-grazing herbage mass and in herbage composition. The allowances of total DM required differed widely between the treatments. There were no significant differences in milk yields between the swards despite the large differences in herbage composition.
The practical implications of these results are discussed.     

Comparison of herbage production under continuous stocking and intermittent grazing

The possibility of increasing the herbage utilized over a grazing season was investigated in a study comparing continuously stocked steady-state swards maintained at optimum height (3.5 cm) with intermittently grazed swards. The intermittent systems were designed (a) to allow periodic increase in leaf area and hence growth rate, (b)to ensure that the accumulated herbage was eaten before it senesced, and (c) lo retain high tiller density by alternating periods of herbage accumulation with periods of continuous stocking. Two treatments (no animals or animal numbers reduced to half those on the 3.5 cm steady-state treatment) were used during the 17-18-d periods of herbage accumulation. Grazing down was completed in 3–4 d, after which two treatments (14 d or 28 d) were used for the intervening periods of continuous stocking when sward height was maintained at 3.5 cm. Herbage production was estimated using the tissue turnover technique, with tiller population densities and rates of growth, senescence and net production per tiller measured at frequent intervals. Intermittent grazing treatments where animals were removed during herbage accumulation resulted in changes in tiller size and number, and in growth rates, but not senescence rates, per tiller such that short-term deviations in the net rate of herbage production occurred compared with the continuously stocked control. The periods of advantage during phases of herbage accumulation were counterbalanced by those of disadvantage during the subsequent steady-state phases. Where animal numbers were reduced during herbage accumulation, sward conditions differed little from those of the continuously stocked control, implying that intake per individual animal was increased. It was concluded that intermittent grazing systems offered no advantage over simpler continuous stocking systems, provided that a flexible approach to conservation was incorporated to allow control of sward conditions on the grazed area.     

Release of cell contents and comminution of particles of perennial ryegrass herbage during ingestion by dairy cows fed indoors or grazing

The effect of feeding indoors fresh perennial ryegrass vs. grazing on ingestive behaviour, release of cell contents and comminution of particles during ingestion, as well as on gas production of ingested boli fermented in vitro, was studied. Indoor feeding and grazing were compared using four dairy cows according to a triple reversal design with six periods. Chemical and morphological composition of the ingested herbage was similar for both indoor feeding and grazing treatments. The intake rate was markedly higher indoors compared with grazing [52·1 vs. 22·9 g dry matter (DM) min^?1] with heavier boli and less saliva added per gram of DM intake. The proportions of intracellular nitrogen and chlorophyll released during mastication after ingestion of herbage fed indoors were lower, and the median size of the particles in the boli was larger (5·97 vs. 4·44 mm) compared with grazing. As a result, the rate of gas production in vitro was also lower for herbage fed indoors compared with grazing (0·423 vs. 0·469 mL min^?1 g^?1 incubated DM). Indoor feeding or grazing may have limited consequences in vivo on the kinetics of availability of nutrients for micro‐organisms in the rumen, because the consequences of the more extensive physical damage suffered by herbage ingested at grazing could be compensated by a lower intake rate.