Variation in vegetation composition,biomass production,and carbon storage in ridge top forests of high mountains of Garhwal Himalaya

Abstract

The present study was aimed to anticipate how forest composition, regeneration, biomass production, and carbon storage vary in the ridge top forests of the high mountains of Garhwal Himalaya. For this purpose five major forest types—(a) Pinus wallichiana, (b) Quercus semecarpifolia, (c) Cedrus deodara, (d) Abies spectabilis, and (e) Betula utilis mixed forests—were selected on different ridge tops in the Bhagirathi Catchment Area of the Uttarkashi District of Garhwal Himalaya. The highest species richness (10 species) and stand density (804 ± 184.5 stems ha^?1) were recorded in Abies spectabilis forests, whereas lowest species richness (4 species) and species density (428 ± 144.7 stems ha^?1) were found in Quercus semecarpifolia forests. The total basal cover (TBC) values were maximum (91.1 ± 24.4 m² ha^?1) in Cedrus deodara forests and minimum (26.5 ± 11.7 m² ha^?1) in Pinus wallichiana forests. The highest total biomass density (TBD) (464.2 ± 152.5 Mg ha^?1) and total carbon density (TCD; 208.9 ± 68.6 Mg C ha^?1) values were recorded for Cedrus deodara forests; however, lowest TBD (283.4 ± 74.8 Mg ha^?1) and TCD (127.5 ± 33.7 Mg C ha^?1) values for Quercus semecarpifolia forests. Our study suggests that Abies spectabilis-dominated forests should be encouraged for biodiversity enrichment and reducing carbon emissions on ridge top forests of high mountains.     

Influence of volcanic ash deposits on the radial growth of trees in Central Mexico: the case of Parícutin volcano

The impact of volcanic eruptions on humans and infrastructure has been significant and widely documented. However, the impact on forests covered by ash or tephra has not been as widely known or documented. This research evaluated the ashfall effects of the 1943–1952 Parícutin eruption on the radial growth of trees located near the top and flanks of the Tancítaro stratovolcano, ca.10 km southwest of the Parícutin volcano. We carried a dendrochronological sampling out in a temperate forest dominated by Pinus hartwegii and Abies religiosa trees. A ring-width chronology was built with 68 increment cores from 47 trees showed two statistically significant suppression events. The first event occurred from 1943 to 1946; it was caused by falling ash of the eruptive columns from Parícutin during the first 6 months of its eruption (February–June 1943). The second event occurred from 1818 to 1823, and it is attributed to the 1818 eruption of Volcán de Colima. The 1943–1946 suppression event observed in the tree rings demonstrates that trees are useful as palaeoenvironmental archives to evaluate the influence of volcanic eruptions in recent centuries in the Trans-Mexican Volcanic Belt. Our results indicate that falling ash immediately impacted the growth of trees in intertropical mountain zones of the Michoacan state located in the Michoacán-Guanajuato Volcanic Field. Such impact means that future eruptions may cause damage to crops and trees at the local and regional levels. Our findings contribute to evaluating volcanic hazards and risks in areas with active volcanoes and monogenetic volcanic fields.

     

Twentieth-century decline of large-diameter trees in Yosemite National Park,California, USA

Studies of forest change in western North America often focus on increased densities of small-diameter trees rather than on changes in the large tree component. Large trees generally have lower rates of mortality than small trees and are more resilient to climate change, but these assumptions have rarely been examined in long-term studies. We combined data from 655 historical (1932–1936) and 210 modern (1988–1999) vegetation plots to examine changes in density of large-diameter trees in Yosemite National Park (3027 km²). We tested the assumption of stability for large-diameter trees, as both individual species and communities of large-diameter trees. Between the 1930s and 1990s, large-diameter tree density in Yosemite declined 24%. Although the decrease was apparent in all forest types, declines were greatest in subalpine and upper montane forests (57.0% of park area), and least in lower montane forests (15.3% of park area). Large-diameter tree densities of 11 species declined while only 3 species increased. Four general patterns emerged: (1) Pinus albicaulis, Quercus chrysolepis, and Quercus kelloggii had increases in density of large-diameter trees occur throughout their ranges; (2) Pinus jeffreyi, Pinus lambertiana, and Pinus ponderosa, had disproportionately larger decreases in large-diameter tree densities in lower-elevation portions of their ranges; (3) Abies concolor and Pinus contorta, had approximately uniform decreases in large-diameter trees throughout their elevational ranges; and (4) Abies magnifica, Calocedrus decurrens, Juniperus occidentalis, Pinus monticola, Pseudotsuga menziesii, and Tsuga mertensiana displayed little or no change in large-diameter tree densities. In Pinus ponderosa–Calocedrus decurrens forests, modern large-diameter tree densities were equivalent whether or not plots had burned since 1936. However, in unburned plots, the large-diameter trees were predominantly A. concolor, C. decurrens, and Q. chrysolepis, whereas P. ponderosa dominated the large-diameter component of burned plots. Densities of large-diameter P. ponderosa were 8.1 trees ha⁻¹ in plots that had experienced fire, but only 0.5 trees ha⁻¹ in plots that remained unburned.     

小兴安岭典型林分类型土壤有机碳含量状况比较分析

对3种典型林型（原始红松林、蒙古栎林、云冷杉林）的林下土壤有机碳含量及相关物理状况进行分析测定,并进行比较的结果表明：蒙古栎林容重最小,孔隙度最高,含水率最高,对土壤物理性质改良效果好;3种林分类型土壤有机碳含量平均值依次为：原始红松林〉云冷杉林〉蒙古栎林,差异极显著（P〈0．01）,原始红松林土壤有机碳含量最高,表层（0～10cm）土壤有机碳含量迭（26．73±3．15）g/kg;3种林型样地表层有机碳含量占总有机碳含量的质量分数比重很大,为41．22％～45．01％,而人类各种活动主要作用于土壤表层,可见减少人为对森林生态系统的干扰活动,可提高森林生态系统固碳能力。     

Seed and microsite limitation for seedling recruitment of <Emphasis Type="Italic">Quercus wutaishanica</Emphasis> on Mt. Ziwuling,Loess Plateau,China

The relative influence of seed and microsite availability in the seedling emergence and recruitment of Liaodong oak (Quercus wutaishanica), which dominates the native hardwood forest in Loess Plateau, was examined by seed sowing experiments. Experiments were performed in 216 plots (30 × 30 cm), located in three typical stands (Pinus tabulaeformis plantation, Liaodong oak forest and grassland, 72 plots per stand). A seed augmentation experiment was performed in an oak forest and a conifer plantation, and a seed introduction experiment was carried out in a grassland. Three main factors were considered: seed addition, disturbance and shrub cover. The seedling number and growth in every plot were recorded continuously over 3 years. GLM multivariate was used to analyze the relationships between seedling emergence/recruitment and the explanatory variables in every stand. The results showed that seed addition in undisturbed or disturbed plots always caused significantly increased seedling emergence and recruitment in oak and conifer forests. It demonstrated that the recruitment of Liaodong oak was limited by seed and microsite availability under closed forest, and litter is one of the factors leading to microsite limitation. However, in the grassland, no treatments improved recruitment, indicating that the grassland was not suitable for seedling recruitment because of intense light and soil drought. In the conifer forest, more recruited seedlings, and lower herbivory on seedling leaves, as well as thicker stem basal diameters, indicated that the P. tabulaeformis plantation was safer for Liaodong oak seedling establishment. Unexpectedly, shrub cover did not affect the recruitment of Liaodong oak in the three stands.     

Regeneration dynamics of dominant tree species along an altitudinal gradient in moist temperate valley slopes of the Garhwal Himalaya

The present study was undertaken in moist temperate forest of Mandal-Chopta area in the Garhwal region of Uttarakhand, India. The aim of the present study was to understand the regeneration dynamics of the dominant tree species along an altitudinal gradient in naturally regenerating, restricted access forest. The overall regeneration status was fairly high in the study area. Most of the native canopy and undercanopy dominants had frequent reproduction and expanding populations, which suggests the stability of forest structure/composition and further expansion of dominant species. The overall regeneration of trees in the forest had a greater contribution of middle and understorey species. Because of infrequent reproduction and declining populations of some of the dominant native species viz., Abies pindrow, Alnus nepalensis and Betula alnoides, structural/compositional changes in the future are expected in respective forests dominated by them. Abies pindrow and Taxus baccata need immediate attention by forest managers for their survival in the area. Seedlings were found to be more prone to competition from herb and shrubs than saplings.     

Recruitment limitation in forests: Lessons from an unprecedented mountain pine beetle epidemic

Since the mid-1990s the forests of central British Columbia have undergone an unprecedented Mountain Pine Beetle (Dendroctonus ponderosae Hopkins) (MPB) epidemic that has resulted in extensive mortality of canopy lodgepole pine (Pinus contorta var. latifolia Engelm.). This study investigated how seed-source availability, seedbed substrate, overstory structure, and time since MPB attack interact to affect post-MPB seedling recruitment of the dominant tree species of these forests. In addition to post-MPB recruitment, these forests may be regenerated by trees established in the understory prior to MPB disturbance. Accordingly, we examined abundance and patterns of all regeneration less than 130 cm tall. We found post-MPB recruitment was sparse. Subalpine fir (Abies laciocarpa (Hook.) Nutt.) comprised the majority of the post-MPB recruitment. It increased with local parent tree basal area and increased strongly with proximity to a major seed source. This resulted in a patchy distribution for subalpine fir post-MPB regeneration. Lodgepole pine post-MPB recruitment was limited by overstory shading. Recruitment of pine decreased as the total overstory basal area increased. Interior spruce (Picea glauca × engelmannii) post-MPB recruitment was similarly limited by total overstory basal area. Seedbed substrates were uniform and dominated by moss. Substrate type distribution did not change as time since MPB disturbance increased. The overall low post-MPB recruitment observed was likely due to a lack of disturbance to the moss-dominated forest floor. Moss is known to be a poor substrate in northern forests. The distribution of all regeneration less than 130 cm tall showed the same trends as the post-MPB regeneration. We believe the post-MPB seedling recruitment dynamics of these forests was not substantially changed from conditions prior to MPB disturbance. There was no pulse of regeneration up to 10 years post-MPB disturbance. Unless this changes, future stand structure will be dominated by the seedling bank established prior to the MPB epidemic. Subalpine fir dominated the seedling bank (68%) and post-MPB recruitment (94%). This suggests that MPB-disturbed forests are undergoing a substantial shift in landscape-level species composition.     

Successional process of Picea crassifolia forest after logging disturbance in semiarid mountains: A case study in the Qilian Mountains,northwestern China

Selective logging is the most widely employed method of commercial timber production in Asia, and its impact on forest structure, composition, and regeneration dynamics is considerable. However, the successional processes in forest communities after logging in semiarid mountains are poorly understood. To provide more information on these processes, we used data from tree rings, direct and indirect age determinations, and field measurements of stand structure to reconstruct the historical disturbance regime, stand development patterns, and successional processes in a natural Picea crassifolia forest community in the Qilian Mountains of northwestern China. The results showed that the density of P. crassifolia forest increased significantly after logging. The densities of second growth forests 30 and 70 years after logging disturbance had increased to 2874% and 294% of primary forest's density, respectively. Logging disturbance did not alter tree species composition of logged stands. However, the diversity of understory species changed significantly among the successional phases. Logging disturbance decreased the spatial heterogeneity of second growth forest. The spatial distributions of recruitment were affected by the location of the remaining trees. There was less recruitment near the remaining trees than near forest that had been cut. In addition, logging disturbance also induced a growth release for the trees on the sites sampled. Our results imply that the succession and regeneration of P. crassifolia forest may be improved if the remaining trees could be retained relative uniform distribution pattern, thinning or selective logging could be performed to height density, exotic shrubs could be removed or the shrubs cover could be reduced during the earlier successional stages.     

Disturbance history and dynamics of an old-growth mixed species mountain forest in the Slovenian Alps

A long history of human presence in the Alps has made studies of natural forest structure, composition, and disturbance processes difficult. In the Slovenian Alps, we identified a mixed European beech (Fagus sylvatica L.)-silver fir (Abies alba Mill.)-Norway spruce (Picea abies (L.) Karst.) stand with old-growth characteristics that escaped past cutting. We reconstructed the dynamics of the stand using a combination of stand and age structure analyses, dendroecological evidence of past canopy disturbance, and historical evidence of past human activities. The disturbance reconstruction revealed a stand-scale disturbance in the 1850s. Based on the presence of pit-and-mound topography throughout much of the stand and the absence of logging activities found in historical records, we assume this event was caused by strong winds. The current structure and composition of the stand are largely a result of this event. Post-disturbance forest development was dominated by the release of shade tolerant regeneration, especially F. sylvatica. The windthrow also created appropriate conditions (i.e. increased light and exposed mineral soil) for recruitment of less shade tolerant species, including larch (Larix decidua Mill.) and maple (Acer pseudoplatanus L.). The results suggest that periodic, intermediate severity wind disturbances, similar to the event documented here, may have had an important influence on pre-settlement forest dynamics in mixed mountain forests of the Alps, and add to the limited available information regarding the historic range of natural variability of disturbance processes. This information may be useful for forest management that attempts to emulate natural processes.     

Saproxylic beetle assemblages in the Mediterranean region: Impact of forest management on richness and structure

Forests cover almost 30% of the Mediterranean region today, yet forest management activities have influenced structure and composition of both natural and planted forests. To date no study has been conducted to evaluate the impact of forest management on saproxylic beetle assemblages, although it is known that the Mediterranean is a biodiversity hotspot with a long-lasting human pressure on natural habitats. We provide an overview of saproxylic beetle assemblages of three forest types (mature Pinus halepensis forests, mature Pinus brutia forests, young Quercus calliprinos forests) in the East Mediterranean region using a one-year sample from 12 forest plots located in the north of Israel.     

昆明西山国家森林公园主要林型地表可燃物的特征

通过外业调查和实验室测定,利用绝对含水率和载量2个关键指标,研究了持续干旱背景下昆明西山国家森林公园的华山松、云南油杉、滇青冈、地盘松、麻栎和旱冬瓜等6种主要林型的可燃物特征。结果表明:地表可燃物、粗死木质残体的含水率最高的都为云南油杉林,分别为55.36%,40.95%,最低都为滇青冈林,分别为10.26%,10.58%,差异显著,最高与最低分别相差约5.4倍、3.9倍;载量都达到较高水平,地表可燃物载量最大为华山松林(4.53±1.11)kg/m2,最低为旱冬瓜林(2.08±0.19)kg/m2,相差约2.2倍,粗死木质残体载量最高为地盘松林为(0.94±0.16)kg/m2,最低为旱冬瓜为(0.07±0.03)kg/m2,相差约13.4倍。     

Forest structure and woody plant species composition along a fire chronosequence in mixed pine–oak forest in the Sierra Madre Oriental, Northeast Mexico

Although wildfires are occurring frequently in the pine–oak forests in the Sierra Madre Oriental (northeastern Mexico), data on post-fire succession and forest structure are still rare. Our objectives were to (1) assess the changes in woody plant species composition after fire and to (2) to relate successional patterns to environmental variability. Based on their fire history 23 plots were selected in the Parque Ecológico Chipinque (PECH). Changes in forest structure across the chronosequence of burned stands were deduced from density, height and diameter measurements of trees and shrubs (>5 cm in diameter) in all plots of 1000 m². Differences in woody plant species composition among the plots were evaluated using Shannon evenness measure and the Whittaker's measure and by Hierarchical cluster analysis and Detrended Correspondence Analysis. Hierarchical cluster analysis showed a high similarity among all recently burned plots, independed of the aspect. Multivariate analysis showed that local environmental factors, including time since fire, continue to structure species composition. Oak species (mainly Quercus rysophylla) resprouted successfully after fire and dominated young post-fire stands. Pine species (Pinus pseudotrobus and Pinus teocote) only appeared 18 years after fire and were the dominating species in mature stands (62 years after fire). In contrast, woody plant species composition in older stands tended also to be influenced by factors such as aspect and by the potential solar radiation (PSR) during the growing season. The results demonstrate that in the PECH, natural regeneration is sufficient and woody plant species composition will be similar to pre-fire conditions after 60 years of succession. We conclude that the park managers should consider incorporating natural disturbance regimes into their management practices.     

云南丽江拉市海汇水区不同森林枯落物的持水性能

对云南丽江拉市海汇水区面山上6种不同森林群落的枯落物储量和持水性能进行了测定,结果表明:不同森林群落的枯落物储量和持水性差别较大,其枯落物储量从最大的黄背栎林(22.45 t·hm-2)到最小的云南松林(6.54 t·hm-2),均是半分解与分解层的储量大于未分解层的储量;6种森林枯落物的最大持水量,除滇杨林外均是半分解与分解层的大于未分解层的,其最大总持水量排序为黄背栎林(60.77 t·hm-2)丽江云杉林(36.42 t·hm-2)云南松+黄背栎+杜鹃混交林(33.18 t·hm-2)川滇高山栎林(29.23 t·hm-2)滇杨林(18.82 t·hm-2)云南松林(13.72 t·hm-2)。各层枯落物的吸水速率均随浸水时间的延长而逐渐降低,在2 4 h后明显减缓,最终趋于零;且半分解与分解层的吸水速率均大于未分解层。6种森林枯落物的拦蓄水量也表现出半分解与分解层大于未分解层的规律,从大到小依次为黄背栎林(66.94 t·hm-2)丽江云杉林(41.24 t·hm-2)云南松+黄背栎+杜鹃混交林(36.80 t·hm-2)川滇高山栎林(32.99 t·hm-2)滇杨林(21.18 t·hm-2)云南松林(16.59 t·hm-2),降雨拦蓄量深分别为6.70、4.12、3.68、3.30、2.12、1.66 mm。     

Patterns of litterfall and return of nutrients across anthropogenic disturbance gradients in three subalpine forests of west Himalaya, India

The study was conducted to improve our understanding of the effects of forest disturbance on litterfall and patterns of nutrient return in three subalpine forest ecosystems (i.e. Betula utilis-dominated, Abies pindrow-dominated, and Acer mixed broadleaf) of Indian west Himalaya. Total litterfall (t ha⁻¹ yr⁻¹) ranged between 2.6–3.6 and 2.1–2.6 for pristine and degraded stands, respectively. Whereas total litterfall decrease from pristine to degraded stand was about 25–30% in B. utilis and Acer mixed-broadleaf forests, the level of disturbance did not affect total litterfall in A. pindrow (coniferous) forest. Nutrient (N, P, and K) concentrations in litter components of the forests studied also varied across forest types and disturbance intensities. For pristine stands, among all the forests, return of total nutrients via litterfall was higher. The study revealed that patterns of litterfall and nutrient return in the forests studied were sensitive to intensity of disturbance, although sensitivity varied among forest types and nutrient contents. Increased intensity of disturbance greatly affected the total annual amount of nutrient return in broadleaf forests. Maximum impact was recorded in B. utilis forest with a significant decline in nutrient return from pristine to degraded stands (i.e. 64% for N, 38% for P, and 67% for K). Corresponding values for decline in Acer mixed forest were 17, 13, and 33% for N, P, and K, respectively, whereas in A. Pindrow forest N return was 15% higher and P return was 33% lower. This study indicates that the litterfall and litter nutrient concentrations in these forests are sensitive to the intensity of disturbance, which affects the amount of nutrient return. This will have a strong bearing on forest nutrient cycling.     

Spatial patterning and population structure of a common woodland herb, Trillium erectum, in primary and post-logging secondary forests

We investigated the population age structure and spatial patterning of a common understory perennial, Trillium erectum var. album, in primary and post-logging secondary (industrially logged in the early 1900s) cove forests of the southern Appalachian Mountains. Within each of 8 T. erectum populations (4 in primary forest and 4 in secondary forest), we mapped the locations of all T. erectum plants in a 10 m × 10 m plot located within the center of each population. We recorded the height and life stage of each plant and excavated a randomly selected subset for age determination. Trillium erectum populations in primary forests displayed a somewhat stronger and more consistent spatial structure than those found in secondary forests. However, aside from being on-average younger due to a higher concentration of juveniles, age structures of secondary forest populations were similar to those of primary forests and both forest types had similar maximum plant ages. Our results, when considered with the life history of the species, suggest that logging did not eliminate T. erectum from secondary forests. Rather, residual plants and patches of plants that survived logging provided the propagules needed to expand and reestablish populations following logging. This relatively rapid recovery stands in stark contrast to the slow recovery observed by others in post-agricultural secondary forest. When considered in the context of other studies of post-disturbance recovery in secondary forests, our results suggest that the full history of a site must be carefully considered when examining the impacts of disturbance on forest perennial herbs.     

Resilience of stand structure and tree species diversity in subtropical forest degraded by clear logging

Subtropical forests in the Ryukyu Islands have been degraded by silvicultural practices, and thus their structural attributes are being shifted to other states dominated by a few tree species. This study clarified the mechanisms of the change, and examined the effect of clear logging on the resilience of a subtropical forest. Sprouting regeneration and typhoon disturbance were introduced into an individual-based model, SEIB-DGVM, for describing stand development and succession. The regeneration dynamics from young secondary to old-growth stands were reproduced fairly well with the model. Sprouting recruitment produced high stem density at the beginning of stand development, which caused a self-thinning trajectory following the −3/2 power law. In the late development stage after 70 years, tree species diversity fluctuated because of the regenerative response of sprouting species and the facilitatory effect of typhoon disturbance on the coexistence of subordinate species. The death of canopy trees because of typhoon disturbances reduced the dominance of Castanopsis sieboldii, and depressed its dominance in the understory. Consequently, the understory species could establish by virtue of fallen canopy trees, and tree species diversity increased at the stand level. Clear logging experiments in the model revealed that species diversity deteriorated, especially in the stand dominated by sprouting species. Resilience of subtropical forests was determined by initial species composition before clear logging. Our simulation results suggest that repeated logging drives subtropical forests with high species diversity to a stand monopolized by C. sieboldii.     

Forest soil and litter as filtering media for suspended sediment

A series of two filtration experiments were conducted to evaluate the filtration function of forest soil experimentally. The first experiment evaluated the differences between the filtration capabilities of the A₀ horizon and A horizon, and the effect of overstory species on the filtration function of the A₀ horizon. Undisturbed A₀, A and A₀+A horizons were collected for the filter mediums with cylindrical samplers. Leaves ofQuercus serrata, Quercus myrsinaeforia, Sasa senanensis Pinus densiflora, Chamaecyparis obtusa, andCryptomeria japonica were also packed in the samplers. Various suspended sediment concentration of water were sprayed at constant intensity on the surface of samples. Filtering coefficients were not affected by SS concentration in all samples, and the order of filtering coefficient was: A₀ horizons>A₀+A horizons>A horizons in undisturbed forest soil samples, andQuercus serrata>Sasa senanensis>Quercus myrsinaeforia>Pinus densiflora”Chamaecyparis obtusa>Cryptomeria, japonica in leaf samples. These results led to the conclusion that SS from managed forests can best be prevented by buffer zones where a thick A₀ horizon is maintained. The second experiment evaluated the effect of turbid water supply rate on the filtration capacity. Undisturbed A horizons and four leaf types,Quercus serrata, Pinus densiflora, Chamaecyparis obtusa, andCryptomeria japonica were used as filter mediums. Filtering coefficients were inversely proportional to supply rate of turbid water in all samples.     

云南宝台山国家森林公园森林资源现状分析

为掌握云南宝台山国家森林公园的森林资源特点,并为制定森林公园的森林资源保护与利用规划提供依据,对该公园2019年森林资源监测数据进行了分析。结果表明:该森林公园林地面积占公园总面积的99.4%,森林覆盖率高达90.50%。林地以有林地为主,面积占比为90.9%。森林起源以天然为主,面积占比为77.8%;人工起源为辅。优势树种以栎类为主,面积占比为45.9%;其次为云南松林,面积占24.1%;再次为核桃林,面积占17.7%。栎类的蓄积量超过总蓄积量的一半,达51.4%;其次为云南松;再次为栲类。森林的龄组以中龄林为主,面积占比为55.2%;幼龄林的面积比例过小。     

Tree recruitment in gaps of various size,clearcuts and undisturbed mixed forest of interior British Columbia,Canada

Tree seedling recruitment was monitored after various types of logging in mixed conifer and deciduous forests of northern British Columbia, Canada. Predicting tree seedling recruitment after disturbance is fundamental to understanding forest dynamics and succession and is vital for forest management purposes. Seedling recruitment success in multi-species northern latitude forests varied as a function of mature tree canopy cover, gap size and position in a gap. Recruitment was abundant within canopy gaps across a wide range of gap sizes (20–5000 m²), but recruit numbers dropped off rapidly under the closed forest canopy and in the open conditions of clearcuts. Inside canopy gaps, recruitment was similar by gap position in small gaps (<300 m²) but, in these northern latitude forests, exhibited a trend of increasing density from the sunny north to shady south end of larger gaps. This was true for all tree species regardless of their shade tolerance ranking. There was no evidence of gap partitioning by any of the tree species during the regeneration phase suggesting that adaptation to the subtleties of gap size during early recruitment are not well developed in these tree species. Favorable locations for emergence and early establishment of germinants were less favorable for growth and survival of established seedlings, i.e. the regeneration niches in these forests were discordant. Tree abundance and species diversity appears to be controlled more by differentiation among growth and survival niches than by the regeneration niches. From the perspective of forest management, abundant natural regeneration of all the dominant tree species of these mixed-species forests can be obtained after partial cutting.