Stand dynamics and tree quality response to precommercial thinning in a northern hardwood forest of the Acadian forest region: 23 years of intermediate results

In the late 1980s, large forest companies began precommercial thinning (PCT) operations in young northern hardwood cutovers in New Brunswick, Canada. To provide supporting growth and yield information, an industrial experiment was established at residual stand densities of 1300, 1600, 1900, and 2200?stems?ha^?1. Stand responses were examined for measurements recorded at 0 (1987), 5 (1992), 10 (1997), 16 (2003), and 23 (2010) years after establishment. Average diameter at breast height, quadratic mean diameter, stand basal area, and stand total volume growth increased as stem density decreased from PCT. There were significant linear differences for many of these variables between treatments and time periods (year). No significant differences were detected in tree height between treatments. In 2010, the four PCT thinning treatments did not exhibit any differences in potential sawlogs at 2.4?m (8?ft) and 3.6?m (12?ft) lengths. Significant differences were observed for 4.9?m (16?ft) sawlogs that were produced at the least dense spacing (1300?stems?ha^?1). Results from this study and recommendations from the European literature suggest that value-added timber products may be produced from more intense PCT treatments than are currently being practiced on sites dominated by yellow birch (Betula alleghaniensis Britt.).     

Influence of precommercial thinning grade on branch diameter and crown ratio in Pinus sylvestris in southern Sweden

Precommercial thinning is regarded as one of the most important measures for influencing timber quality in stands of Scots pine (Pinus sylvestris L.). The present study considers the influence of both thinning intensity and tree height at the time of thinning on diameter of the thickest branch and crown ratio. A total of 115 plots (either naturally regenerated, planted or sown, and either thinned down to 528–8000 stems?ha^?1 or untreated) included in 20 sites in southern Sweden were analysed. The average tree height after thinning varied from 1.2 to 8.3?m. An increasing number of remaining stems resulted in a reduction in branch diameter, although the reduction appeared to be only minor if the number of stems after thinning was more than 3000 stems?ha^?1. It was found that late thinning reduced the diameter of the thickest branch. The crown ratio decreased with stand height, number of stems after thinning and average height at thinning. The results were consistent for all trees and for the 500 thickest trees per hectare.     

Pre-commercial thinning,birch admixture and sprout management in planted Norway spruce stands in South Sweden

Early management of the regenerated seedlings shapes the future stand properties. To address these issues, pre-commercial thinning (PCT) and control treatments were applied to planted Norway spruce (Picea abies L. Karst) and naturally regenerated birch (Betula pendula Roth., Betula pubescens Ehrh.) stands in forest experiments in southern Sweden (lat. 56–57?N) containing 1.1–5.5?m tall saplings. The treatments were retention of 1000 or 2000?stems?ha^?1 of Norway spruce, with no birch or birch at 1000?stems?ha^?1. Treatments were replicated with and without annual removal of birch sprouts from stumps. The periodic annual increment (PAI) over five years was calculated for total stand volume and individual trees. The mean PAI of dominant trees was significantly higher both following all PCT treatments than controls, and following low rather than high-density PCT. Birch retention did not affect growth of the dominant trees but PAI was lower in plots with uncontrolled sprouting. The PAI of birch was significantly higher in low-density Norway spruce plots than in control plots and the high-density plots. The treatment response was significant even in stands with initial heights of only 1–2?m.     

Stem Quality and Height Development in a Betula-Dominated Stand Seven Years After Precommercial Thinning at Different Stump Heights

Topping may provide an attractive alternative to traditional precommercial thinning, offering several advantages. The main stem quality of birch (Betula pubescens Ehrh. and Betula pendula Roth, species not separated) was examined seven growing seasons after such topping in a dense, even-aged and naturally regenerated stand in northern Sweden. The height development and survival of main and secondary stems of birch and secondary stems of willow (Salix spp.) were also studied. At a main stem height of 1.8 m, the following treatments, leaving 3100 main stems ha^?1, were applied to secondary stems: traditional precommercial thinning (cutting just above ground), top-cutting at either 40% (71 cm above ground) or 70% (120 cm above ground) of main stem mean height, and no precommercial thinning. For main stems, topping resulted in fewer forks, straighter stems, higher live crown height and smaller diameter of the thickest branch compared with traditional precommercial thinning. In these respects, topping at the higher level above ground gave the best quality. The main stems were no longer at risk of becoming overtopped by the secondary stems following any of these treatments (except for no precommercial thinning).     

Simulation of the long-term effects of different strategies for precommercial thinning in Pinus sylvestris

The influence of the intensity and timing of precommercial thinning (PCT) on stand development and financial return was studied in Scots pine stands. Functions describing the early development of the stand structure after PCT were developed. The functions were based on 195 plots within 41 PCT experiments in Sweden. The dimension distribution of the established stand was estimated and used as input to the decision support system Heureka, to simulate the stand development until final felling. The studied treatments included PCT to 1000, 2000 and 3000?stems?ha^?1 at mean heights of 2, 4 and 6 m. Separate simulations were carried out for different site fertilities. Mean annual volume increment increased with increasing number of stems after PCT whereas the timing of PCT had only a small effect. The land expectation value (LEV) generally decreased with increasing mean height at PCT, primarily because of the increased cost of PCT. LEV decreased with increasing number of stems after PCT at low fertility sites whereas only minor differences were found for PCT to 1000 and 2000?stems?ha^?1 at medium and high fertility sites. The general pattern persisted when different scenarios of future timber quality were simulated.     

Stand density and fertilization effects on aboveground allocation patterns and stem form of Pinus sylvestris in young stands

The increasing demand for forest biomass for energy generation could be partially met by growing denser stands and use of fertilizer. Before this is done at large scale, more knowledge of the effects of stand density and fertilization on aboveground allocation patterns and stem form is needed. Therefore, effects of pre-commercial thinning (PCT) to 3000 stems ha^?1, an unthinned dense control (C), and PCT combined with two levels of fertilization (100 kg ha^?1 of nitrogen applied either during the establishment of the field experiment (F1) or annually (F2)) were examined in 23- to 26-year-old Scots pine (Pinus sylvestris L.) stands six years after the establishment of the field experiment. In total, 114 sample trees were harvested using destructive biomass sampling. The growth allocation and stem form of trees with diameter at breast height (DBH; 1.3 m height) >5.0 cm were not affected by either the PCT or fertilization. Small trees (DBH < 5 cm) in denser, unthinned control plots had more slender stems (lower DBH/height ratios) and allocated less growth to branches and foliage than trees in PCT plots. Fertilization had little effect on the stem form and growth allocation of the smallest trees. Therefore, effects of stem density and fertilization on stem form and growth allocation to foliage were only found for small suppressed trees, and the treatments had very little influence on dominant and codominant trees.     

Analysis of Spacing for Spotted Gum Plantations for Maximizing Merchantable Logs' Volume in Southeast Queensland,Australia

Spotted gum (Corymbia citriodora subspecies Variegata) has the potential to be the major hardwood species for large-scale plantations in Southeast Queensland, Australia, but production research is limited due to the lack of age of research plots. Optimal spacing is a major subject of concern. Based on time series data from a spotted gum experiment site, growth performance was analyzed for five spacing levels: 11.3 m?×?11.3 m (78 stems ha^?1), 7.4 m?×?7.4 m (182 stems ha^?1), 5.4 m?×?5.4 m (343 stems ha^?1), 3.6 m?×?3.6 m (771 stems ha^?1), and 2.9 m?×?2.9 m (1189 stems ha^?1). The major objective was assumed to be to maximize total merchantable log volume. A growth model was produced, and the mean diameter at breast height (DBH) and total merchantable log volume for each spacing level at a range of harvesting ages were estimated. From the analysis, the spacing level of 5.4 m?×?5.4 m was found to be optimal for maximizing merchantable log volume to a 10-cm small-end diameter. Further analysis of mean DBH, height, and volume of the largest 200 and 250 trees from this spacing level indicates that merchantable log volume could be maximized by retaining the 250 largest trees ha^?1. The total financial revenue from the best spacing level in 25 and 30 yr are predicted to be Australian dollars (A$) 13,637 and A$17,779 ha^?1, respectively. If full rotation data could be obtained, more reliable models could be produced, and a more accurate financial estimate could be made.     

Effects of thinning paper birch on conifer productivity and understory plant diversity

Naturally regenerated paper birch (Betula papyrifera Marsh.) is commonly removed from juvenile interior Douglas-fir (Pseudotsuga menziesii var. glauca [Beissn.] Franco) plantations in southern interior British Columbia, Canada, to increase conifer productivity and create a free-growing stand; however, this practice is expensive and contentious because of possible negative ecological impacts. One solution is to retain an optimal density of birch where growth gains of understory Douglas-fir are balanced against losses to Armillaria ostoyae (Romagn.) Herink and understory plant species diversity. We sought to find this optimal density by comparing four evenly applied birch density reduction treatments (0, 400, 1111, and 4444 retained birch stems ha^?1) and an unthinned control (>7300 retained birch stems ha^?1). The mortality rate of Douglas-fir due to Armillaria root disease increased non-significantly with thinning intensity. Mean diameter increment of surviving Douglas-fir improved the most where birch was completely removed, with little variation among intermediate thinning treatments. Height growth was unaffected by the thinning treatments. Diversity of cryptogams was significantly greater in the control than where all birch was removed. We suggest that the treatment with 4444 retained birch stems ha^?1 provides the best balance for improving Douglas-fir growth while minimizing risk of increased Armillaria root disease and reduced understory plant diversity in young mixed stands.     

Effects of pruning and stand density on the productionof new epicormic shoots in young stands of pedunculate oak (Quercus robur L.)

Context

Avoidance or control of epicormic shoots is among the major silvicultural challenges for the production of high-quality oak timber. In northern Europe, contemporary oak silviculture aims to produce valuable timber on a relatively short rotation, applying early, heavy thinning combined with artificial pruning.

Aims

The aim of this study was to analyse the effects of pruning and stand density on the production of new epicormic shoots on young trees of pedunculate oak (Quercus robur L.).

Methods

The study was based on two field experiments in even-aged stands of pedunculate oak subjected to different thinning practices and early selection of potential future crop trees. From ages 13 to 15 years, stem density was reduced to 300 trees ha^?1, 1,000 ha^?1 or stands remained unthinned. Pruning was conducted on selected trees at ages 22–24 years. At that age, the stem density in unthinned control plots ranged from 2,500 to 3,100 ha^?1. All treatments were replicated twice within each experiment.

Results

Pruning led to an overall increase in the total production of new epicormic shoots. More epicormic shoots were produced in the lower part of the stem (0–3 m in height) than in the upper part (3–6 m). The number of new epicormic shoots increased with increasing stand density.

Conclusion

Early, heavy thinning combined with high pruning at regular intervals may help shorten the rotation length for pedunculate oak without further reduction in wood quality than that which is caused by wider annual growth rings.     

Canopy density in stands of Picea abies and Pinus sylvestris after different thinning methods

Altogether 82 plots (261 estimations) of Picea abies (L.) Karst, and 193 plots (360 estimations) of Pinus sylvestris (L.) stands were estimated by a vertical tube. The “crown free projection”, CFP, of stands thinned in three methods with different thinning grades was measured: unthinned, heavily and very heavily thinned, heavily thinned delayed first thinning, extra heavily thinned and thinned from the top. Basal area (m²ha^?1) density (stems ha^?1) and diameter sum (m ha^?1) were plotted against CFP. Basal area was the best practical measure of stand in this study. Generally Scots pine stands have higher CFP and the curves are steeper than in Norway spruce stands. Depending on the grade of thinning, heavily and very heavily thinned spruce stands, delayed first thinning included, have CFP values of 10–15% and stands thinned from the top, 20–40%, compared with 30–80% and 30–60% respectively in pine stands. Extra heavily thinned stands have the highest CFP, 20–80% in spruce and 50–90% in pine stands. The CFP levels after thinning are too high in pine stands for avoidance of sucker and sprout production of aspen and birch. In dense Norway spruce stands thinned from the top or heavily and very heavily thinned, the CFP values are low enough (≤30%) to diminish the production of suckers.     

Influence of Shelterwood Density on Survival and Height Increment of Picea abies Advance Growth

Abstract

The increases in the arithmetic mean diameter at breast height (dbh) of the 100, 200, 300 and 400 largest stems ha^?1 by dbh associated with six different thinning regimens for periods up to 35 years were compared, by analyses of variance, to the corresponding stem fractions in unthinned stands using permanent sample plot data. In addition, the expected mean dbh of these stand fractions at future points in time was modelled, based on site and stand data obtained at the time of the first thinning. Thirty-five years after the first thinning, the thinning from below regimens resulted in actual increases (a measure affected by both the thinning intensity and the thinning method), compared with the unthinned stands, in the mean dbh for the 100, 200, 300 and 400 largest stems ha^?1 of 2.6–9.0 cm. The corresponding genuine increases (a measure principally affected solely by the thinning intensity) were 4–11 cm. The functions predicting the future dbh of these stand fractions yielded good fits.     

尾叶桉萌芽林留条数量对生长和产量的影响

对 5 5年生尾叶桉实生林砍伐后 ,保留不同萌条数量对其萌芽林生长和产量的影响开展了研究。结果表明 ,定株使桩的存活率明显下降 ,特别是保留单株 ,但保留一定的株数却有利于萌芽林的生长 ;萌芽林的高、径和单株材积生长随保留株数的减少而显著增加 ;3 5年生单位面积蓄积量和出材率 (顶木 )以保留 3株·桩- 1为最好 ,其次为 4株·桩- 1和 2株·桩 - 1,蓄积量分别为 4 5 4 9、4 4 2 1和4 2 5 1m3·hm- 2 ,分别是对照的 131 7%、12 8 0 %和 12 3.0 % ,出材率 (顶木 )分别是对照的 174 .3%、14 8.6 %和 15 5 .4 %。因此 ,在中等地力 ,种植密度为 3m× 2m的尾叶桉萌芽林 ,以纸浆材和顶木为主要经营目的的林分 ,建议其萌芽林的保留株数为 2　 4株·桩 - 1。     

Precommercial thinning of beech (Fagus sylvatica L.): Early effects of stump height on growth and natural pruning of potential crop trees

Abstract

This paper reports the early effects of stump height on the growth and natural pruning of potential crop trees after precommercial thinning of a young stand of naturally regenerated beech (Fagus sylvatica L.) in Denmark. The experiment comprises five treatments based on combinations of three grades of precommercial thinning and three stump heights. Treatments include the unthinned control, thinning only of whips and wolf trees (using low stumps), and thinning for potential crop trees using low, medium or high stumps. All treatments were replicated three times. Stump heights averaged 10, 90 or 230 cm, respectively. Pretreatment stem number (live trees) varied from 17,500 to 41,000 ha^?1. In the potential crop tree treatment, post-treatment stem number ranged from 4750 to 9500 ha^?1. Following two growth seasons, the quantity of stump regrowth increased with increasing stump height, the rate of stump regrowth increased with increasing stump height, the diameter growth of potential crop trees increased with decreasing stump height, the increase in stand height did not depend on stump height or post-treatment stem number, and the natural pruning of potential crop trees increased with increasing stump height. It remains to be seen whether these trends hold in the long run, and whether additional economic return from the increase in wood quality with increasing stump height compensates for the reduction in diameter growth.     

Practical large-scale forest stand inventory using a small-footprint airborne scanning laser

Mean tree height, dominant height, mean diameter, stem number, basal area and timber volume of 116 georeferenced field sample plots were estimated from various canopy height and canopy density metrics derived by means of a small-footprint laser scanner over young and mature forest stands using regression analysis. The sample plots were distributed systematically throughout a 6500 ha study area, and the size of each plot was 232.9 m². Regressions for coniferous forest explained 60–97% of the variability in ground reference values of the six studied characteristics. A proposed practical two-phase procedure for prediction of corresponding characteristics of entire forest stands was tested. Fifty-seven test plots within the study area with a size of approximately 3740 m² each were divided into 232.9 m² regular grid cells. The six examined characteristics were predicted for each grid cell from the corresponding laser data using the estimated regression equations. Average values for each test plot were computed and compared with ground-based estimates measured over the entire plot. The bias and standard deviations of the differences between predicted and ground reference values (in parentheses) of mean height, dominant height, mean diameter, stem number, basal area and volume were ?0.58 to ?0.85 m (0.64–1.01 m), ?0.60 to ?0.99 m (0.67–0.84 m), 0.15–0.74 cm (1.33–2.42 cm), 34–108 ha^?1 (97–466 ha^?1), 0.43–2.51 m² ha^?1 (1.83–3.94 m² ha^?1) and 5.9–16.1 m³ ha^?1 (15.1–35.1 m³ ha^?1), respectively.     

Simulated yield and quality distribution of sawn timber from final felling in a Norway spruce [Picea abies (L.) Karst.] stand with varying thinning regimes: A case study

Abstract

An integrated simulation tool, formed by integrating the InnoSIM sawing simulation system with the RetroSTEM simulator, was used to convert available wood raw materials from final felling into sawn timber, allowing for calculation of the three-dimensional wood properties of individual stems (stem geometry, heartwood formation, knottiness) as well as the volume, quality and value of sawn timber in a Norway spruce stand with different thinning regimes (unthinned, normal and intensively thinned). Based on the input data of sawing patterns, the simulations indicate that there are relatively small differences (<8%) in the volume yield (m³ha^?1) of sawn goods resulting from sawlogs available from final felling with different thinning practices. However, intensive thinning yielded the largest stem diameters and the greatest volumes (m³ha^?1) of large-sized centre goods (thickness: 50, 63, 75 mm) of rather poor quality. Normal thinning yielded the largest volume of A-grade side boards and centre goods (m³ha^?1), as well as the best total value ([euro]ha^?1) of sawn timber. Differences observed in sawn timber quality distribution can contribute to even more significant variation in value yields, if pricing mechanisms of timber products change to favour higher grade timber products.     

Foliar Responses of Understorey Abies lasiocarpa to Different Degrees of Release Cutting in a Betula papyrifera and Conifer Mixed Species Stand

Foliar responses of subalpine fir [Abies lasiocarpa (Hook.) Nutt.] to thinning were studied in a 35-yr-old mixed stand of paper birch (Betula papyrifera Marsh.) and conifers. The stand regenerated naturally after a wildfire with a canopy dominated by paper birch (average height 9.8 m) and an understorey dominated by subalpine fir (average height 1.6 m). The stand was thinned to four densities of birch: 0, 600 and 1200 stems ha-¹ and control (unthinned at 2300-6400 stems ha-¹) in the autumn of 1995. The understorey conifers, mainly subalpine fir, were thinned to 1200 stems ha-¹. The study used a completely randomized split-plot design. Three sample trees were systematically selected from each treatment replicate and each tree stratum (upper, intermediate and lower understorey). One-year-old and older age class needles were collected from one south-facing branch within the fifth whorl from the tree top. Thinning of paper birch significantly (p <0.001) increased leaf area and dry weight per 100 needles for intermediate and short trees except in the 0 birch treatment. Understorey subalpine fir trees in 600 stems ha¹ birch (T3) had the largest leaf area and leaf dry weight per 100 1-yr-old needles. Specific leaf area (SLA) decreased from unthinned (T1) to 0 birch (T4). Lower understorey trees had the largest SLA. One-year-old needles had significantly higher N, P and K concentrations in all the thinning treatments. These responses are consistent with the shade tolerance of subalpine fir. The results suggest that when managing a paper birch-conifers mixed-wood forest it may be of benefit to understorey conifers to leave a birch canopy as a nursing crop.     

Economic feasibility of and vegetation responses to biomass harvest in managed loblolly pine plantations

We evaluated two biomass harvest methods, (1) Whole Tree Thinning (WTT; third-row thinning), and (2) Whole Tree Thinning with Fuelchips (WTTF; third-row thinning plus remove all accessible hardwood stems >2.5 cm diameter at breast height and understory shrubs in thinned area of stand) in a 21-year-old loblolly pine (Pinus taeda L.) plantation in northwest Florida for their harvesting yields and productivities, costs, and effects on groundcover. Both WTT and WTTF produced similar quantities of roundwood (~70 Mg ha^?1) and chips (8.43–13.12 Mg ha^?1) without significantly added operational time (15.77–28.12 Mg h^?1). On-board costs of chip production (US$6.93 to 10.60 Mg^?1) and total cost of roundwood and chip production (~22.5 US$ Mg^?1) of the two harvest methods also did not differ significantly. Following either WTT or WTTF, overall percent groundcover recovered within 6 months. While shrub and grass cover were similar, forb cover in WTT increased significantly following six months of harvests. Importantly, the study suggests that biomass removal is an attractive option that could be integrated with traditional silvicultural thinning methods to manage vegetation and alleviate hazardous fuel and wildfire conditions, leading to improved forest health.     

Natural regeneration in patch clear-cutting in Picea abies stands in Southern Finland

Abstract

The success of natural regeneration on patch clear-cuts in Norway spruce stands in Southern Finland was examined in terms of stocking and tree height. The experiment was established in mature spruce stands in the submesic Myrtillus (MT) site type. In each of the eight study sites, three plots were treated with clear-cutting and planting, clear-cutting with partial tree retention and planting, and patch clear-cutting without site preparation, respectively. In three of the study sites, one plot with patch clear-cutting and site preparation was established. Each plot was 1 ha, out of which three patches of 40×40 m were clear-cut in the patch treatments. 10–11 years after cutting, the patches had on average 1316 crop trees ha^–1, of which 91% were spruces. Some 27% of the stands were up to the target stocking level (≥1600 ha^?1), and 36% were at least satisfactory (≥1300 ha^?1). Site preparation did not yield greater stocking levels on patches, but that result is ambiguous due to a difference in initial stocking. The average spruce tree height in the patches (0.76 m) was much smaller than in the case of clear-cutting and planting (2.42 m). In conclusion, the patches had been restocked tolerably well with spruce and birch for practical purposes during the 10- to 11-year period, but the regeneration process had been very slow compared to clear-cutting and planting.     

Optimal management of Korean pine plantations in multifunctional forestry

Korean pine is one of the most important plantation species in northeast China.Besides timber,it produces edible nuts and plantations sequester carbon dioxide from the atmosphere.This study optimized the management of Korean pine plantations for timber production,seed production,carbon sequestration and for the joint production of multiple benefits.As the first step,models were developed for stand dynamics and seed production.These models were used in a simulation–optimization system to find optimal timing and type of thinning treatments and optimal rotation lengths.It was found that three thinnings during the rotation period were optimal.When the amount or profitability of timber production is maximized,suitable rotation lengths are 65–70 years and wood production is 5.5–6.0 m~3 ha~(-1) a~(-1).The optimal thinning regime is thinning from above.In seed production,optimal rotation lengths are over 100 years.When carbon sequestration in living biomass is maximized,stands should not be clear-cut until trees start to die due to senescence.In the joint production of multiple benefits,the optimal rotation length is 86 years if all benefits(wood,economic profits,seed,carbon sequestration) are equally important.In this management schedule,mean annual wood production is 5.5 m~2 ha~(-1) and mean annual seed yield 141 kg ha~(-1).It was concluded that it is better to produce timber and seeds in the same stands rather than assign stands to either timber production or seed production.     

Response of young densely stocked stands of Pinus radiata to thinning and refertilization

The effects of thinning and refertilization on growth of Pinus radiata in three experiments planted at 11 960 stems ha^–1 were examined. Responses to thinning and refertilization were obtained at 3 or 5 years after planting despite heavy applications of fertilizer at establishment. During the 3 years following retreatment, trees which had been thinned to 2990 stems ha^–1 had twice the sectional area increments of unthinned trees, while up to a 3-fold increase in increment was obtained on trees which had been thinned and refertilized. Responses to nitrogen were obtained in all experiments. Response to phosphate was dependent upon soil type and prior treatment. Foliar analyses do not appear to be reliable for predicting responses to fertilization applied in combination with thinning.