Vegetation development over four years on two green roofs in the UK

Carefully designed green roofs have the potential to be used as mitigation for habitats lost at ground level. The development of plant assemblages on two green roofs designed to emulate diverse brownfield habitats (brown roofs), by using recycled demolition aggregate as part of a low-fertility growth substrate, were studied over the first four years of their development. The cover-abundance of flowering plants and habitat structural components (e.g. bare ground, moss) were measured on the Domin-Krajina scale within all identified microhabitats. Drought disturbance was one of the main controlling factors on assemblage development. Annual plants were abundant and successful in the first growth season, and thereafter only re-appeared in any numbers following drought disturbances in subsequent years. Moss and Sedum acre L. increased through the study period until these plants dominated coverage. The cover-abundance of perennial wildflower species was strongly influenced by drought disturbance. The influence of drought disturbance varied between different brown roof microhabitats, with plant assemblages in coarser and less fertile microhabitats more resistant to these disturbances. Observed responses to drought were consistent with the following two hypotheses: (i) Areas of coarse substrate can act as disturbance refugia for plants during drought by helping preserve pockets of water under large clasts and within absorbent materials such as brick. (ii) The plant assemblages living in areas of more fertile substrates, which grow more luxuriantly when water availability is high, are more vulnerable to drought disturbance. Green roofs should be designed to include a range of substrate types to create several microhabitats that will collectively support more species than any single microhabitat.     

Substrate temperature fluctuation and dry-weight partitioning of Lantana grown in four green roof growing media

Summary

Four potential green roof substrates were evaluated in a field study for their thermal characteristics and their impact on Lantana camara dry-weight accumulation. The substrates were a) a sandy loam soil (S), b) the sandy loam soil amended with urea formaldehyde resin foam (S:F) at a proportion of 60:40 v/v, c) the sandy loam soil mixed with peat and perlite (S:P:Per) at a proportion of 50:30:20 v/v, respectively and, d) peat amended with urea-formaldehyde resin foam (P:F) at a proportion of 60:40 v/v. Measurements included the monitoring of the substrate temperature and the determination of the dry-weight accumulation rhythm of shoots, leaves and roots. In addition, the lateral spreading of the roots was recorded and the shoot/root ratio determined. Temperature fluctuation within the substrates was found to depend on the type of the substrate, plant growth, and season. Temperature fluctuation was high in S, moderate in S:F and S:P:Per and low in P:F. Shoot and root dry-weight accumulation was reduced in P:F, while root growth was promoted in S substrate. Differences in dry-weight accumulation and the lateral spreading of the roots were affected more by the water holding capacity of the substrate than by the substrate temperature.     

Review on the roles and effects of growing media on plant performance in green roofs in world climates

Growing media (substrate) is a fundamental part of a green roof, providing water, nutrients and support to plants. However, little research has reviewed how it affects plant performances in different climatic regions. This study aims to analyse published research on green roof growing medium across world’s climate zones. Findings are structured according to Köppen–Geiger climate classification, aiming to investigate the prevalence of research conducted in different climate zones. Results from full-scale studies and laboratory or greenhouse experiments were reviewed. The later were included as they provide systematic knowledge on the effect of individual factors on system performances although cannot provide climate specific information. Studies discussed effects of major substrate components and depths on plant survival and establishment using standard test procedures. Results showed that most research in the subject were in temperate (group C climate classification), continental (group D) and dry climates (group B), respectively. Considerable number of investigations was conducted in controlled laboratory or greenhouse environments. Based on the results, future green roof research and guidelines should consider climate specifications of the region in designing growing medium, depths and attribute of green roof substrates in order to ensure enhanced plant performance. Especially, for more fragile but less investigated dry climate, considerations should be made to tackle heat fluctuations and drought stress by enhancing water holding capacity and thermal isolation of the substrate. To move forward, sustainable building solutions as a part of future urban forms, climate-adaptive green roof systems should be included into future research, practice and guidelines.     

Root development of Thuja plicata in peat-substitute rooting media

In this study we investigated the potential usefulness of different substrates produced from gorse (Ulex europaeus L.) for propagating cuttings of Thuja plicata ‘Atrovirens’. The gorse was composted by different methods, to produce four types of compost. These composts were then used as such (100% compost) or in mixes with coarse sand and pine bark compost (25% compost, 25% coarse sand and 50% pine bark compost, v/v/v) to prepare eight different substrates. The substrates were compared in cutting propagation trials carried out in a greenhouse, with a standard rooting medium consisting of 25% Sphagnum peat moss, 25% coarse sand and 50% pine bark compost (v/v/v). Cuttings were taken from a garden hedge in March 2003, and comprised shoots of the previous year's growth. The bases of the cuttings were treated with a 0.4% solution of IBA rooting hormone, then inserted into cells filled with the corresponding substrate and placed on a propagation bench. Rooting percentage and visual rooting score were determined in July. The quality of roots was evaluated, as total root length and number of root tips, in July, September and November 2003, by use of a Delta-T Scan system. The rooting success of the cuttings ranged from 75% to 90%, except when the substrate comprised the pure composted gorse, obtained after the shorter period in an enclosed composting system with addition of fertilizer and reuse of leachates. Most of the cuttings in this substrate died and only 4% of the cuttings rooted successfully. Physicochemical characteristics of this rooting medium included very high electrical conductivity, low C/N ratio and high water retention. Addition of coarse sand and pine bark compost to the composted gorse significantly increased the rooting success. The root quality variables differed depending on the substrate considered and date of harvest. The results showed that substrates made from gorse compost are of potential use as a peat substitute for propagating cuttings of T. plicata ‘Atrovirens’. The physicochemical properties of the composted gorse should be determined before its use to establish whether it can be used pure as a substrate for rooting or whether it should be mixed with other substances.     

Bioreceptivity of living walls: Interactions between building materials and substrates,and effect on plant growth

Bringing nature into urban areas is largely recognized as one of the multiple benefits of green walls. Among the existing types of living walls, those made from durable building materials have a longer lifespan and require less maintenance than many typical green wall systems. In that context, there is a need to study the bioreceptivity of materials, i.e., their ability to host living organisms. This paper presents a novel method to study the bioreceptivity of materials, by enhancing the biological and physicochemical exchanges in different mixtures of crushed materials and substrates, and by measuring their effect on plant growth. For this experiment, we crushed and sieved four different building materials (3 types of concretes and 1 brick) to increase their contact surface. We then mixed them with substrate with or without compost additions. The mixtures were used as growing media for the cultivation of two wild mural plant species in a glasshouse. Plant growth and the physicochemical evolution of the mixtures were monitored for one year. The study shows the physical and chemical differences between the building materials, their interactions with substrate composition, and their effect on plant growth. Two main results appeared: (i) high pH (>8) was the major constraint against plant growth (ii) compost additions significantly compensated for adverse characteristics of the materials. Assuming that higher plant growth is associated with better bioreceptivity, this experiment introduces a quantitative method that could contribute to the evaluation of the agronomic properties of building materials. The results allow us to make recommendations for Living wall and Technosol design.     

An initial experimental assessment of the influence of substrate depth on floral assemblage for extensive green roofs

Extensive green roofs have the potential to be used as mitigation tools to compensate for urban habitat loss, but there is little information about how closely these systems emulate ground-based habitats. This study investigated the effect of limited substrate depth on plant assemblages in the initial phase of growth in extensive green roof substrates. Five replicate mesocosms (1 m²) for each of three design treatments: (A) 10 cm aggregate depth with green roof drainage and solid floor, (B) 15 cm aggregate depth with green roof drainage and solid floor, and (C) 15 cm aggregate depth on top of bare earth; were positioned at ground level. Each mesocosm had an identical growth substrate and was seeded with the same seed mix. Plant assemblages were analysed using point-quadrat methods. Significant differences in species composition were observed between treatments that seemed to be related to water availability. Even the deep (15 cm) solid floor green roof treatment showed many significant differences in floral assemblage compared to the identical treatment (C) where plants had access to water in the soil profile. Therefore, it is not possible to exactly recreate most ground-based urban habitats on roofs by simply copying the soil characteristics and floral composition found on the ground. Like for mitigation for habitat loss using extensive green roofs requires the careful manipulation of design elements in order to counteract the limited water availability on green roofs.     

植物工厂中不同供液方式对辣椒育苗的影响

以植物工厂辣椒育苗为对象,研究漂浮式与潮汐式供液方式对育苗的影响。试验结果表明,漂浮式处理营养液和基质pH值显著高于潮汐式处理,EC值显著低于潮汐式处理;与潮汐式处理相比,漂浮式处理中幼苗株高、茎粗分别增加18.4%,23.6%;漂浮式处理幼苗根尖数、总根长、直径在0~0.5 mm的根长分别增加了37.0%,45.5%,50.1%;幼苗地上部鲜质量、全株鲜质量、地上部含水量、植株含水量分别增加45.6%,32.1%,2.5%,1.8%。漂浮式育苗显著促进辣椒幼苗侧根生长,形成良好的根系系统,促进根系对水分和可溶性盐类的吸收,加快地上部形态建成,提升种苗整体质量。     

Effects of artificial amendments in potting media on Orthosiphon aristatus growth and development

Artificial polyacrylamide gel (PAG) and urea-formaldehyde resin foam (UFRF) amendments are used for putative enhancement of soil physical properties, including increasing their water holding capacity (WHC). Effects were investigated of these two amendments alone and combined on growth and development of Orthosiphon aristatus (Cats’ Whiskers) grown in either composted pine bark or washed river sand, including under transient water deficit stress. UFRF and PAG were incorporated into these potting media substrates at recommended rates of 30% (v/v) and 0.1% (w/w), respectively. UFRF incorporation reduced bulk density and increased air-filled porosity of composted pine bark from 0.24 g cm⁻³ and 43.3% to 0.18 g cm⁻³ and 50.2%, respectively. UFRF also reduced bulk density of sand from 1.43 g cm⁻³ to 1.17 g cm⁻³ and increased its air-filled porosity from 18.5% to 25.3%. PAG slightly decreased bulk density of composted pine bark to 0.23 g cm⁻³ and also reduced sand bulk density to 1.32 g cm⁻³. Water content of composted pine bark and sand was increased by PAG addition from 47.6% and 27.7% to 51.0% and 34.2%, respectively. However, UFRF and/or PAG did not increase plant available water (PAW) in either composted pine bark or sand. PAW was 23.8% and 14.4%, 23.6% and 15.8%, 22.8% and 14.8%, and 25.2% and 17.8% for composted pine bark and sand controls, these two substrates amended with UFRF, these substrates amended with PAG, and these substrates with UFRF plus PAG, respectively. Neither shoot length nor number was increased by adding UFRF and/or PAG. Similarly, neither shoot fresh nor dry weight was increased by UFRF and/or PAG amendments. UFRF and, moreso, UFRF plus PAG slightly delayed the onset of wilting by 4–15 h in water deficit stressed O. aristatus compared to control and PAG alone in composted pine bark (experiment 1), but PAG did not. However, in experiment 2, UFRF and/or PAG did not delay wilting in either composted pine bark or sand. Thus, when incorporated at suppliers recommended rates, neither UFRF nor PAG conferred appreciable benefits for O. aristatus plant growth in either composted pine bark or sand potting media.     

Can microbial inoculants boost soil food webs and vegetation development on newly constructed extensive green roofs?

Green roofs are a key to providing nature-based solutions in cities. However, most green roofs installed in the Northern hemisphere are shallow, stonecrop planted systems (“extensive” green roofs), which have been shown to support limited biodiversity and could be more effective at providing ecosystem services. One issue with this type of extensive green roof is that rootzones are almost sterile on construction, relying on natural colonisation to provide a soil food web. This is a slow process, meaning plant growth can also be slow. Our aim was to determine if a soil food web could be introduced when the green roof is built. We applied microbial inoculants (mycorrhizal fungi and bacteria (Bacillus spp.)) to a new green roof and monitored plant growth and the soil food web (bacteria, mycorrhizal fungi and microarthropods). Different inoculants altered the composition of microarthropod communities, potentially impacting later succession. In particular, bacterial inoculants increased microarthropod populations. This is one of the first studies to demonstrate that the addition of microbial inoculants impacts not only plant growth, but also faunal components of the soil food web, which could have implications for long-term resilience. Bacteria were effective at aiding mycorrhizal colonisation of plants roots, but this colonisation had no impact on the growth of our selected stonecrops, Sedum album, Petrosedum reflexum and Phedimus spurius. We suggest that if a beneficial mycorrhiza could be found to promote the growth of these specific species on green roofs, bacteria could be effective “helper” species to aid colonisation. This study enables green roof researchers and the industry to justify further exploration of the impact of microbial inoculants on green roofs.     

The Reinvigoration of Apple Trees by the Inarching of Vigorous Rootstocks

Summary

The purpose of this research was to determine if established, orchard-grown apple trees that were con®ned in root-restricting membranes received adequate water for growth when irrigated. Previous data had shown positive effects of root restriction on reducing shoot extension growth in apple. Soil matric potential (Ψ_sm), leaf stomatal conductance (g) and leaf water potentials (Ψ_l) were measured over daily cycles during the period of maximum tree water use (July and August). Measurements were also made of the Ψ_sm and fruit size throughout the growing season. Daily and seasonal Ψ_sm showed that the soil within the root restricting membranes (+R) when irrigated (+I) remained closed to field capacity (<100 HPa), which was not the case for unirrigated soil within the membranes (-I +R). Ψ_l measured before dawn, showed that similar levels of drought stress were evident between irrigated and restricted (+I +R) and unrestricted (+I -R and -I -R) trees. The Ψ_sm, and Ψ_l for trees with roots within restricting membranes were significantly more negative, in the absence of irrigation (-I +R). Measurement of g showed that root restricted trees were transpiring at similar rates in the presence of irrigation (+I +R) as unrestricted trees (+I -R) with or without irrigation. Stomatal closure could not explain the increased Ψ_l observed for the restricted irrigated (+I +R) treatment compared with unrestricted (-R) trees. A reduced stomatal aperture was the most likely explanation for the reduction in growth previously observed with the restricted unirrigated trees. Fruit size was also affected by root restriction and the effect became greater as the roots became more restricted with tree age. Similarly, there was also a negative effect, in one year, of root restriction on fruit size at harvest, even in the presence of irrigation. Data show that reductions in soil water availability, Ψ_l and g, for the root-restricted trees (+I +R), were unlikely to be the causes for the previously observed reductions in shoot growth (tree size). These results imply that other factors were in operation, among which root-synthesized chemical regulators of shoot growth are the most likely candidates.     

铁皮石斛无土栽培基质的研制与优化

为筛选铁皮石斛组培苗的无土栽培基质,在温室内采用细锯末、苔藓、蛭石、松树皮配制成10种不同的栽培基质,栽培60 d后统计成活率;120 d后测定铁皮石斛苗在各基质中的生长指标。结果表明：铁皮石斛组培苗在细锯末∶松树皮∶苔藓=2∶4∶4的基质中成活率最高,移栽60 d时成活率达93.30%,植株高度达2.85 cm;移栽120 d后试管苗在细锯末∶松树皮∶苔藓=2∶4∶4的基质中平均株高12.49 cm、发根数量为8条、最长根系为3.49 cm,平均茎粗为0.93 cm、平均萌芽数为3个,且叶色浓绿、生长健壮,总体情况显著优于其他基质配比,为当地铁皮石斛组培苗栽培松树皮、锯末等废弃物的利用提供参考。     

Performance of herbaceous Evolvulus pilosus on urban green roof in relation to substrate and irrigation

Rooftop gardening (or green roof establishment) is an urban greening afforestation method that has many environmental, economic, and urban landscaping benefits. For rooftop vegetation, stressful environments that include heat, strong winds, sunshine, and drought prevent many plants from growing well, especially in shallow soil. To establish low cost urban rooftop gardens with a low weight substrate in the summer, we evaluated the green coverage and growth of a suitable groundcover ornamental plant, blue daze (Evolvulus pilosus). Blue daze was used as rooftop vegetation in a shallow substrate (10 cm thick) on the flat rooftop of a four-story building in the middle of Kobe city, Japan, which has a temperate and humid climate. On average, the diurnal maximum temperatures and the total amounts of precipitation during each planting experiment between July and September were 30.8 °C and 247 mm in 2008 and 33.2 °C and 458 mm in 2009, respectively. Three different types of planting substrates (AS, amended soil, TM, turf mat as a dried-up blocks of sod, and FBA, furnace bottom ash) were tested to determine how they influenced plant cover, flowering, and growth. We evaluated the physical and chemical properties of the substrates that were used to establish the rooftop greeneries. In addition, the economy, ornamental and ecological values, reuse of material, and low rooftop weight burdens of the substrates were evaluated. The greatest blue daze ground cover occurred on the TM substrate (relative to the AS and FBA substrates) approximately three months after transplanting. The blue daze actively bloomed on all substrates throughout midsummer. However, alternate day irrigation largely reduced their shoot growth and flowering relative to daily irrigation. The AS and TM substrates had suitable physicochemical properties for use as planting substrates and had ideal solid, liquid, and vapor phase distributions and high water and nutrient holding capacities. FBA was a poor substrate with very low water and nutrient retention capacities, which resulted in depressed plant growth, especially under low watering conditions. To recycle FBA efficiently as a shallow low-cost planting substrate for rooftop gardening, the addition of coated fertilizers and organic matter to FBA should be considered to improve its water and nutrient holding capacities.     

Using superpixel- or pixel-based segmentation for efficient green roof digital image classification and rapid estimation of plant species cover

Green roofs are “nature-based solutions” that provide numerous ecosystem services in the context of urban green infrastructures. Plant species diversity and the associated vegetation communities, in strong interactions with green roof substrate, play a central role in the green infrastructure functioning. In order to better understand the influence of vegetation in relation with the co-benefits provided by green roofs as well as to select suitable species for these usually harsh environments, it is essential to be able to achieve accurate and long-term monitoring of plant communities. In this short communication, two free plugins recently developed for the open-source image analysis software Fiji (a distribution of the freely available ImageJ platform, initially dedicated to biological image analysis) were investigated for their capacity to rapidly and efficiently perform supervised machine-learning for the classification of green roof vegetation photographs, with the aim of estimating individual plant species abundance. Two simple methods are thus described using the Trainable Weka Pixel Segmentation (Arganda-Carreras et al., 2017) or the Trainable Superpixel Segmentation (Salinas Colina et al., 2018), which allowed for rapid, efficient and reproducible classification and estimation of multispecies colonized green roof regardless the color or shape similarities among species or ground cover materials. Finally, recommendations are made for the use of the Trainable Superpixel Segmentation which is particularly convenient for quick and efficient green roof image analysis.     

砖栽应用于番茄农业生产的可行性研究

采用砖块栽培和基质栽培,通过设定不同的砖孔密度（0、6、12 孔和18 孔）,研究其对番茄
生长的影响。结果表明：18 孔、12 孔、6 孔和无孔砖栽番茄果实产量分别比基质栽培番茄果实产量增加
了27.63%、30.84%、23.38%、24.65%,且生产投入成本小。砖栽番茄植株的株高、茎粗、叶片数、叶干
质量、茎干质量、根系干质量、果实产量、营养液利用效率相对于基质栽培番茄都有所增加,砖栽应用于
现代化农业生产具有一定的可行性。     

Bio-efficiency of compost extracts on the wet rot incidence,morphological and physiological growth of okra (Abelmoschus esculentus [(L.) Moench])

An experiment was carried out to investigate the efficacy of rice straw and empty fruit bunch (EFB) of oil palm compost extracts either fortified or unfortified with Trichoderma harzianum on morpho-physiological growth and occurrence of Choanephora wet rot of okra. Treatments tested were water (control) (T1), rice straw (RST) compost extract (T2), Trichoderma-enriched RST compost extract (T3), empty fruit bunch of oil palm compost extract (T4), Trichoderma-enriched EFB compost extract (T5), aqueous suspension of Trichoderma (T6), and a fungicide Dithane M-45^® (2 g l⁻¹ of water) (T7). The experimental results revealed significant variations amongst the treatments in respect of morphological characters, e.g. shoot length, tap root length, number of leaves per plant, and leaf area. The shoot and tap root length, number of leaves per plant, leaf area were significantly (P ≤ 0.05) higher in Trichoderma-enriched RST compost extracts (T3) followed by Dithane M-45^® (T7), Trichoderma-enriched EFB extracts (T5), RST (T2), EFB (T4) and aqueous suspension of T. harzianum (T6) in both Choanephora inoculated and uninoculated (control) plots. Similarly, net photosynthetic rate and chlorophyll content were higher in plants receiving Trichoderma-enriched RST compost extracts (T3) with 76.2% reduction in Choanephora wet rot incidence when compared with rest of the treatments. These experimental results revealed that morpho-physiological characters of okra could be modified by the application of Trichoderma-enriched compost extracts. This suggests that use of Trichoderma-enriched compost extracts would be more beneficial in environmentally friendly okra cultivation and may be used as an alternative to inorganic fertilizers/fungicides to enhance plant growth and reduce disease incidence subsequently, resulting in higher yield.     

本特草球盘区土壤改良的研究

以树皮堆肥４０％＋发酵鸡粪１５％＋蟹皮粉１０＋沸石粉３５％配制的土壤改良材料脂明显地促进本特草的生长，合理地调整土壤的三相分布，改善土壤透水何水性，提高土壤对氨态氮的吸收容量。超缓效氮肥可促进本特草后期生长；与改良材料配合比例无互作效应。     

Vegetation establishment on ‘Green Walls’: Integrating shotcrete walls from road construction into the landscape

The establishment of a sustainable vegetation layer on shotcrete walls subjected to road environment represents enormous challenges. High inclination of almost 90°, the exposed position of the walls near streets as well as lack of soil and water supply pose major limiting conditions. The objective of this study was to evaluate the performance of an innovative greening technology for vertical structures in terms of vegetation development on varying plant substrates and geotextiles.The field experiment included testing three plant substrates on basis of nearby rocky excavation material (‘Innsbrucker Quarzphyllit’, ‘Bündnerschiefer’ and ‘Zentralgneis’) combined with compost. Additionally, five geotextiles (geogrid (3x4 mm), geogrid (9x10 mm), coir net, coir mat, geo mat) were applied for assessment. All test combinations were evaluated regarding vegetation cover, species composition, and biomass production from 2015 to 2016. Analyses of chemical properties were conducted for all plant substrates.Results showed highest vegetation cover ratio on ‘Bündnerschiefer’ and ‘Innsbrucker Quarzphyllit’, which can be explained by the favorable mineral composition (nutrient storage capacity) and chemical properties of compost (lower values of electrical conductivity and C/N ratio).In conclusion, the use of ‘Green Walls’ filled with ‘Bündnerschiefer’ or ‘Innsbrucker Quarzphyllit’ plant substrate in combination with netlike geotextiles like geogrid or coir net proved best. They are promising in terms of establishing an optimal vegetation cover on vertical structures and are well suited for integrating shotcrete walls into the landscape. The use of local excavation material for greening purposes thus can be confirmed. Though, the use of high-quality compost is crucial.     

控释肥对菇渣基质电导率及容器苗生长的影响

通过木槿容器苗栽培试验,研究了控释肥、控释肥+少量普通复合肥(BB肥)、普通复合肥、营养液对菇渣基质电导率及木槿苗叶片数、枝条长度、最大叶面积、叶绿素含量、植株鲜重等生长指标的影响。结果表明:普通肥料处理前期菇渣基质电导率过高,后期电导率偏低,木槿容器苗生长不理想;控释肥处理前期菇渣基质EC值稍低,但后期木槿各项生长指标与营养液处理相当;控释肥加入少量普通复合肥制成的BB肥,养分释放与木槿容器苗的养分需求基本一致,木槿生长总体优于其他施肥处理。控释肥施肥方法简单,效果理想,是基质容器育苗值得推广的一种施肥方法。     

不同粉碎度的椰糠基质对黄瓜穴盘苗生长发育及其质量的影响

设置5、10、15目3个不同粉碎度的椰糠基质,按照椰糠∶蛭石=3 V∶1 V进行复配,以草炭∶蛭石=3 V∶1 V为对照,通过研究基质的理化性质（容重、总孔隙度、持水孔隙度、pH、电导率）及其对黄瓜幼苗生长发育的影响,筛选出适宜黄瓜幼苗生长的椰糠基质粉碎度。结果表明,在3个不同的粉碎度中,10目粉碎度的处理最有利于黄瓜穴盘苗的生长。植株达到了较为理想的生长状态,壮苗指数、根冠比、干质量,以及幼苗中叶绿素含量和净光合速率等较其他两个处理均有所提高。     

Specification of Radiant Flux and Radiant Flux Density in Irradiation of Plants with Artificial Light

Aseptic culture of green shoot tips or etiolated bud-cuttings in Galston-Loo agar with α-naphthylacetic acid induced callus formation and root initiation. Continued aseptic culture in liquid medium without NAA led to root and shoot growth, and development of a plant able to survive transfer and continue growth in soil.